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Early human migrations are the earliest migrations and expansions of archaic and modern humans across continents. They are believed to have begun approximately 2 million years ago with the early expansions out of Africa by ''
Homo erectus ''Homo erectus'' (; meaning "upright man") is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Several human species, such as '' H. heidelbergensis'' and '' H. antecessor' ...
''. This initial migration was followed by other
archaic humans A number of varieties of ''Homo'' are grouped into the broad category of archaic humans in the period that precedes and is contemporary to the emergence of the earliest early modern humans (''Homo sapiens'') around 300 ka. Omo-Kibish I (Omo I) f ...
including ''
H. heidelbergensis ''Homo heidelbergensis'' (also ''H. sapiens heidelbergensis''), sometimes called Heidelbergs, is an extinct species or subspecies of archaic human which existed during the Middle Pleistocene. It was subsumed as a subspecies of '' H. erectus'' in ...
'', which lived around 500,000 years ago and was the likely ancestor of
Denisovan The Denisovans or Denisova hominins ) are an extinct species or subspecies of archaic human that ranged across Asia during the Lower and Middle Paleolithic. Denisovans are known from few physical remains and consequently, most of what is known ...
s and
Neanderthal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ago. While th ...
s as well as modern humans. Early hominids had likely crossed
land bridge In biogeography, a land bridge is an isthmus or wider land connection between otherwise separate areas, over which animals and plants are able to cross and Colonisation (biology), colonize new lands. A land bridge can be created by marine regre ...
s that have now sunk. Within Africa, ''
Homo sapiens Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
'' dispersed around the time of its
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within ...
, roughly 300,000 years ago. The
recent African origin In paleoanthropology, the recent African origin of modern humans, also called the "Out of Africa" theory (OOA), recent single-origin hypothesis (RSOH), replacement hypothesis, or recent African origin model (RAO), is the dominant model of the ...
paradigm suggests that the anatomically modern humans outside of Africa descend from a population of ''Homo sapiens'' migrating from
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
roughly 70–50,000 years ago and spreading along the southern coast of Asia and to Oceania by about 50,000 years ago. Modern humans spread across Europe about 40,000 years ago. Early Eurasian ''Homo sapiens'' fossils have been found in Palestine and Greece, dated to 194,000–177,000 and 210,000 years old respectively. These fossils seem to represent failed dispersal attempts by early ''Homo sapiens'', who were likely replaced by local Neanderthal populations. The migrating modern human populations are known to have
interbred In biology, a hybrid is the offspring resulting from combining the qualities of two organisms of different breeds, varieties, species or genera through sexual reproduction. Hybrids are not always intermediates between their parents (such as in ...
with earlier local populations, so that contemporary human populations are descended in small part (below 10% contribution) from regional varieties of archaic humans. After the
Last Glacial Maximum The Last Glacial Maximum (LGM), also referred to as the Late Glacial Maximum, was the most recent time during the Last Glacial Period that ice sheets were at their greatest extent. Ice sheets covered much of Northern North America, Northern Eur ...
, North Eurasian populations migrated to the Americas about 20,000 years ago. Northern Eurasia was peopled after 12,000 years ago, at the beginning of the
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...
period. Arctic Canada and Greenland were reached by the
Paleo-Eskimo The Paleo-Eskimo (also pre-Thule or pre-Inuit) were the peoples who inhabited the Arctic region from Chukotka Autonomous Okrug, Chukotka (e.g., Chertov Ovrag) in present-day Russia across North America to Greenland prior to the arrival of the mode ...
expansion around 4,000 years ago. Finally,
Polynesia Polynesia () "many" and νῆσος () "island"), to, Polinisia; mi, Porinihia; haw, Polenekia; fj, Polinisia; sm, Polenisia; rar, Porinetia; ty, Pōrīnetia; tvl, Polenisia; tkl, Polenihia (, ) is a subregion of Oceania, made up of ...
was populated within the past 2,000 years in the last wave of the
Austronesian expansion The Austronesian peoples, sometimes referred to as Austronesian-speaking peoples, are a large group of peoples in Taiwan, Maritime Southeast Asia, Micronesia, coastal New Guinea, Island Melanesia, Polynesia, and Madagascar that speak Austron ...
.


Early humans (before ''Homo sapiens'')

The earliest humans developed out of
australopithecine Australopithecina or Hominina is a subtribe in the tribe Hominini. The members of the subtribe are generally ''Australopithecus'' (cladistically including the genus, genera ''Homo'', ''Paranthropus'', and ''Kenyanthropus''), and it typically in ...
ancestors about 3 million years ago, most likely in
Eastern Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
, most likely in the area of the
Kenyan Rift Valley The Great Rift Valley is part of an intra-continental ridge system that runs through Kenya from north to south. It is part of the Gregory Rift, the eastern branch of the East African Rift, which starts in Tanzania to the south and continues no ...
, where the oldest known stone tools were found. Stone tools recently discovered at the
Shangchen Shangchen () is a Lower Palaeolithic archaeological site in Lantian County, Shaanxi, China, some 25 km south of Weinan. It was discovered in 1964, and excavated during 2004 and 2017. Stone tools found at the site were dated based on magne ...
site in China and dated to 2.12 million years ago are claimed to be the earliest known evidence of hominins outside Africa, surpassing
Dmanisi Dmanisi ( ka, დმანისი, tr, , az, Başkeçid) is a town and archaeological site in the Kvemo Kartli region of Georgia approximately 93 km southwest of the nation’s capital Tbilisi in the river valley of Mashavera. The hominin ...
in Georgia by 300,000 years.


''Homo erectus''

Between 2 and less than a million years ago, ''
Homo ''Homo'' () is the genus that emerged in the (otherwise extinct) genus ''Australopithecus'' that encompasses the extant species ''Homo sapiens'' ( modern humans), plus several extinct species classified as either ancestral to or closely relate ...
'' spread throughout East Africa and to
Southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number of ...
(''
Homo ergaster ''Homo ergaster'' is an extinct species or subspecies of archaic humans who lived in Africa in the Early Pleistocene. Whether ''H. ergaster'' constitutes a species of its own or should be subsumed into ''Homo erectus, H. erectus'' is an ongoin ...
''), but not yet to West Africa. Around 1.8 million years ago, ''
Homo erectus ''Homo erectus'' (; meaning "upright man") is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Several human species, such as '' H. heidelbergensis'' and '' H. antecessor' ...
'' migrated out of Africa via the
Levantine corridor The Levantine corridor is the relatively narrow strip between the Mediterranean Sea to the northwest and deserts to the southeast which connects Africa to Eurasia. This corridor is a land route of migrations of animals between Eurasia and Africa ...
and
Horn of Africa The Horn of Africa (HoA), also known as the Somali Peninsula, is a large peninsula and geopolitical region in East Africa.Robert Stock, ''Africa South of the Sahara, Second Edition: A Geographical Interpretation'', (The Guilford Press; 2004), ...
to
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
. This migration has been proposed as being related to the operation of the Saharan pump, around 1.9 million years ago. ''Homo erectus'' dispersed throughout most of the
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
, reaching as far as
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
. Its distribution is traced by the
Oldowan The Oldowan (or Mode I) was a widespread stone tool archaeological industry (style) in prehistory. These early tools were simple, usually made with one or a few flakes chipped off with another stone. Oldowan tools were used during the Lower ...
lithic industry, by 1.3 million years ago extending as far north as the 40th parallel (
Xiaochangliang Xiaochangliang () is the site of some of the earliest paleolithic remains in East Asia, located in the Nihewan (泥河灣) Basin in Yangyuan County, Hebei, China, most famous for the stone tools discovered there. Stone tools The tool forms discov ...
). Key sites for this early migration out of Africa are
Riwat Riwat (Rawat, Murree) is a Paleolithic site in Punjab, northern Pakistan. Another site, called Riwat Site 55, shows a later occupation dated to around 45,000 years ago. Site The site was discovered in 1983. The artifacts consist of flakes and c ...
in Pakistan (~2 Ma?),
Ubeidiya Ubeidiya commonly refers to: * Ubeidiya prehistoric site * Ubeidiya, West Bank * Ubeidiya, Tiberias See also * Abd (Arabic) ʿAbd ( ar, عبد) is an Arabic word meaning one who is subordinated as a slave or a servant, and it means also to w ...
in the Levant (1.5 Ma) and
Dmanisi Dmanisi ( ka, დმანისი, tr, , az, Başkeçid) is a town and archaeological site in the Kvemo Kartli region of Georgia approximately 93 km southwest of the nation’s capital Tbilisi in the river valley of Mashavera. The hominin ...
in the Caucasus (1.81 ± 0.03 Ma, p=0.05).
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
shows evidence of Erectus from 2.12 Mya in Gongwangling in Lantian county.Zhu, Zhaoyu, Robin Dennell, Weiwen Huang, Yi Wu, Shifan Qiu, Shixia Yang, Zhiguo Rao, et al. 2018. “Hominin Occupation of the Chinese Loess Plateau since about 2.1 Million Years Ago.” Nature 559 (7715): 608–612. . Two ''Homo erectus'' incisors have been found near Yuanmou, south China, and are dated to 1.7 Mya, and a cranium from Lantian has been dated to 1.63 Ma. Artefacts from Majuangou III and Shangshazui in the
Nihewan basin Xiaochangliang () is the site of some of the earliest paleolithic remains in East Asia, located in the Nihewan (泥河灣) Basin in Yangyuan County, Hebei, China, most famous for the stone tools discovered there. Stone tools The tool forms disco ...
, north China, have been dated to 1.6–1.7 Ma. The archaeological site of
Xihoudu Xihoudu () is an archeological site located in the Shanxi Province of China. The site dates to the Paleolithic Age. In total 32 stone implements were found at the site. Discovery In 1929, Chinese prehistoric archaeologist and paleontologist Prof ...
() in
Shanxi Shanxi (; ; formerly romanised as Shansi) is a landlocked province of the People's Republic of China and is part of the North China region. The capital and largest city of the province is Taiyuan, while its next most populated prefecture-lev ...
Province is the earliest recorded use of fire by ''Homo erectus'', which is dated 1.27 million years ago.
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
(
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...
) was reached about 1.7 million years ago (
Meganthropus ''Meganthropus'' is an extinct genus of non-hominin hominid ape, known from the Pleistocene of Indonesia. It is known from a series of large jaw and Hominid skull, skull fragments found at the Sangiran site near Surakarta in Central Java (island) ...
). Western
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
was first populated around 1.2 million years ago ( Atapuerca).
Robert G. Bednarik Robert G. Bednarik (born 6 April 1944 in Vienna) is an Australian prehistorian and cognitive archeologist. Robert Bednarik moved from Austria to Australia in 1966. Bednarik, who considers himself an autodidact, is an expert in the fields of rock ...
has suggested that ''Homo erectus'' may have built rafts and sailed oceans, a theory that has raised some controversy.


After ''H. erectus''

One million years after its dispersal, ''H. erectus'' was diverging into new species. ''H. erectus'' is a
chronospecies A chronospecies is a species derived from a anagenesis, sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a populatio ...
and was never extinct, so its "late survival" is a matter of taxonomic convention. Late forms of ''H. erectus'' are thought to have survived until after about 0.5 million ago to 143,000 years ago at the latest, with derived forms classified as ''
H. antecessor ''Homo antecessor'' (Latin "pioneer man") is an extinct species of archaic human recorded in the Spanish Sierra de Atapuerca, a productive archaeological site, from 1.2 to 0.8 million years ago during the Early Pleistocene. Populations of this s ...
'' in Europe around 800,000 years ago and ''
H. heidelbergensis ''Homo heidelbergensis'' (also ''H. sapiens heidelbergensis''), sometimes called Heidelbergs, is an extinct species or subspecies of archaic human which existed during the Middle Pleistocene. It was subsumed as a subspecies of '' H. erectus'' in ...
'' in Africa around 600,000 years ago. ''H. heidelbergensis'' in its turn spread across East Africa (''
H. rhodesiensis ''Homo rhodesiensis'' is the species name proposed by Arthur Smith Woodward (1921) to classify Kabwe 1 (the "Kabwe skull" or "Broken Hill skull", also "Rhodesian Man"), a Middle Stone Age fossil recovered from a cave at Broken Hill, or Kabwe, ...
'') and to Eurasia, where it gave rise to
Neanderthals Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an Extinction, extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ag ...
and
Denisovans The Denisovans or Denisova hominins ) are an extinct species or subspecies of archaic human that ranged across Asia during the Lower and Middle Paleolithic. Denisovans are known from few physical remains and consequently, most of what is known ...
. ''H. heidelbergensis'', Neanderthals and Denisovans expanded north beyond the 50th parallel (
Eartham Pit, Boxgrove Eartham Pit is an internationally important archaeological site north-east of Boxgrove in West Sussex with findings that date to the Lower Palaeolithic. The oldest human remains in Britain have been discovered on the site, fossils of ''Homo heidel ...
500kya,
Swanscombe Heritage Park Swanscombe Skull Site or Swanscombe Heritage Park is a geological Site of Special Scientific Interest in Swanscombe in north-west Kent, England. It contains two Geological Conservation Review sites and a National Nature Reserve. The park lies ...
400kya,
Denisova Cave Denisova Cave (russian: Денисова пещера, lit= the cave of Denis, translit= Denísova peshchéra; alt, Аю-Таш, lit= Bear Rock, translit= Ayu Tash) is a cave in the Bashelaksky Range of the Altai mountains, Siberia, Russia. The ...
50 kya). It has been suggested that late Neanderthals may even have reached the boundary of the
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
, by c. 32,000 years ago, when they were being displaced from their earlier habitats by ''H. sapiens'', based on 2011 excavations at the site of Byzovaya in the
Urals The Ural Mountains ( ; rus, Ура́льские го́ры, r=Uralskiye gory, p=ʊˈralʲskʲɪjə ˈɡorɨ; ba, Урал тауҙары) or simply the Urals, are a mountain range that runs approximately from north to south through European ...
(
Komi Republic The Komi Republic (russian: Республика Коми; kv, Коми Республика), sometimes simply referred to as Komi, is a republic of Russia located in Eastern Europe. Its capital is the city of Syktyvkar. The population of the ...
, ). Other archaic human species are assumed to have spread throughout Africa by this time, although the fossil record is sparse. Their presence is assumed based on traces of admixture with modern humans found in the genome of African populations. ''
Homo naledi '' Homo naledi'' is an extinct species of archaic human discovered in 2013 in the Rising Star Cave, Cradle of Humankind, South Africa dating to the Middle Pleistocene 335,000–236,000 years ago. The initial discovery comprises 1,550 specimen ...
'', discovered in South Africa in 2013 and tentatively dated to about 300,000 years ago, may represent fossil evidence of such an archaic human species. Neanderthals spread across the Near East and Europe, while Denisovans appear to have spread across Central and East Asia and to Southeast Asia and Oceania. There is evidence that Denisovans interbred with Neanderthals in Central Asia where their habitats overlapped. Neanderthal evidence has also been found quite late at 33,000 years ago at the 65th latitude of the Byzovaya site in the
Ural Mountains The Ural Mountains ( ; rus, Ура́льские го́ры, r=Uralskiye gory, p=ʊˈralʲskʲɪjə ˈɡorɨ; ba, Урал тауҙары) or simply the Urals, are a mountain range that runs approximately from north to south through western ...
. This is far outside of any otherwise known habitat, during a high ice cover period, and perhaps reflects a refugia of near extinction.


''Homo sapiens''


Dispersal throughout Africa

''
Homo sapiens Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
'' are believed to have emerged in Africa about 300,000 years ago, based in part on thermoluminescence dating of artefacts and remains from
Jebel Irhoud Jebel Irhoud or Adrar n Ighoud ( zgh, ⴰⴷⵔⴰⵔ ⵏ ⵉⵖⵓⴷ, Adrar n Iɣud; ar, جبل إيغود, žbəl iġud), is an archaeological site located just north of the locality known as Tlet Ighoud, approximately south-east of the cit ...
, Morocco, published in 2017. The
Florisbad Skull The Florisbad Skull is an important human fossil of the early Middle Stone Age, representing either late ''Homo heidelbergensis'' or early ''Homo sapiens''. It was discovered in 1932 by T. F. Dreyer at the Florisbad archaeological and paleontolo ...
from Florisbad, South Africa, dated to about 259,000 years ago, has also been classified as early ''Homo sapiens''. Previously, the
Omo remains The Omo remains are a collection of homininThis article quotes historic texts that use the terms 'hominid' and 'hominin' with meanings that may be different from their modern usages. This is because several revisions in classifying the great apes h ...
, excavated between 1967 and 1974 in
Omo National Park Omo National Park is a national park in Ethiopia founded in 1980. Located in the Southern Nations, Nationalities, and Peoples' Region on the west bank of the Omo River, the park covers approximately 4,068 square kilometers, about 870 kilometers so ...
,
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
, and dated to 200,000 years ago, were long held to be the oldest known fossils of ''Homo sapiens''. In September 2019, scientists reported the computerized determination, based on 260
CT scan A computed tomography scan (CT scan; formerly called computed axial tomography scan or CAT scan) is a medical imaging technique used to obtain detailed internal images of the body. The personnel that perform CT scans are called radiographers ...
s, of a virtual skull shape of the last common human ancestor to anatomically modern humans, representative of the earliest modern humans, and suggested that modern humans arose between 260,000 and 350,000 years ago through a merging of populations in
East East or Orient is one of the four cardinal directions or points of the compass. It is the opposite direction from west and is the direction from which the Sun rises on the Earth. Etymology As in other languages, the word is formed from the fa ...
and
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
. In July 2019, anthropologists reported the discovery of 210,000 year old remains of a ''H. sapiens'' and 170,000 year old remains of a ''H. neanderthalensis'' in
Apidima Cave The Apidima Cave (, ''Spilaio Apidima'') is a complex of five caves four small caves located on the western shore of Mani Peninsula in Southern Greece. A systematic investigation of the cave has yielded Neanderthal and ''Homo sapiens'' fossils f ...
in southern
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
, more than 150,000 years older than previous ''H. sapiens'' finds in Europe. Early modern humans expanded to Western Eurasia and Central, Western and Southern Africa from the time of their emergence. While early expansions to Eurasia appear not to have persisted, expansions to
Southern Southern may refer to: Businesses * China Southern Airlines, airline based in Guangzhou, China * Southern Airways, defunct US airline * Southern Air, air cargo transportation company based in Norwalk, Connecticut, US * Southern Airways Express, M ...
and
Central Africa Central Africa is a subregion of the African continent comprising various countries according to different definitions. Angola, Burundi, the Central African Republic, Chad, the Democratic Republic of the Congo, the Republic of the Congo, ...
resulted in the deepest temporal divergence in living human populations. Early modern human expansion in sub-Saharan Africa appears to have contributed to the end of late
Acheulean Acheulean (; also Acheulian and Mode II), from the French ''acheuléen'' after the type site of Saint-Acheul, is an archaeological industry of stone tool manufacture characterized by the distinctive oval and pear-shaped "hand axes" associated ...
(
Fauresmith Fauresmith is located 130 km south west of Bloemfontein, South Africa. The town, named after Rev Phillip Faure and Sir Harry Smith, is the second oldest town in the Free State. Fauresmith is the only town in South Africa, and one of only t ...
) industries at about 130,000 years ago, although very late coexistence of archaic and early modern humans, until as late as 12,000 years ago, has been argued for West Africa in particular. The ancestors of the modern
Khoi-San Khoisan , or (), according to the contemporary Khoekhoegowab orthography, is a catch-all term for those indigenous peoples of Southern Africa who do not speak one of the Bantu languages, combining the (formerly "Khoikhoi") and the or ( in t ...
expanded to Southern Africa before 150,000 years ago, possibly as early as before 260,000 years ago, so that by the beginning of the
MIS 5 Marine Isotope Stage 5 or MIS 5 is a marine isotope stage in the geologic temperature record, between 130,000 and 80,000 years ago. Sub-stage MIS 5e, called the Eemian or Ipswichian, covers the last major interglacial period before the Holocene, w ...
"
megadrought A megadrought (or mega-drought) is a prolonged drought lasting two decades or longer. Past megadroughts have been associated with persistent multiyear La Niña conditions (cooler than normal water temperatures in the tropical eastern Pacific Oc ...
", 130,000 years ago, there were two ancestral population clusters in Africa, bearers of mt-DNA haplogroup L0 in southern Africa, ancestral to the Khoi-San, and bearers of haplogroup L1-6 in central/eastern Africa, ancestral to everyone else. There was a significant back-migration of bearers of L0 towards eastern Africa between 120 and 75 kya. Expansion to Central Africa by the ancestors of the
Central African forager The African Pygmies (or Congo Pygmies, variously also Central African foragers, "African rainforest hunter-gatherers" (RHG) or "Forest People of Central Africa") are a group of ethnicities Indigenous peoples of Africa, native to Central Africa ...
populations (African Pygmies) most likely took place before 130,000 years ago, and certainly before 60,000 years ago. The situation in
West Africa West Africa or Western Africa is the westernmost region of Africa. The United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Mali, Maurit ...
is difficult to interpret due to a sparsity of fossil evidence. ''Homo sapiens'' seems to have reached the western Sahelian zone by 130,000 years ago, while tropical West African sites associated with ''H. sapiens'' are known only from after 130,000 years ago. Unlike elsewhere in Africa, archaic
Middle Stone Age The Middle Stone Age (or MSA) was a period of African prehistory between the Early Stone Age and the Late Stone Age. It is generally considered to have begun around 280,000 years ago and ended around 50–25,000 years ago. The beginnings of pa ...
sites appear to persist until very late, down to the Holocene boundary (12,000 years ago), pointing to the possibility of late survival of
archaic human A number of varieties of ''Homo'' are grouped into the broad category of archaic humans in the period that precedes and is contemporary to the emergence of the earliest early modern humans (''Homo sapiens'') around 300 ka. Omo-Kibish I (Omo I) f ...
s, and late
hybridization Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...
with ''H. sapiens'' in West Africa.


Early northern Africa dispersal

Populations of ''Homo sapiens'' migrated to the Levant and to Europe between 130,000 and 115,000 years ago, and possibly in earlier waves as early as 185,000 years ago. A fragment of a jawbone with eight teeth found at Misliya Cave has been dated to around 185,000 years ago. Layers dating from between 250,000 and 140,000 years ago in the same cave contained tools of the Levallois type which could put the date of the first migration even earlier if the tools can be associated with the modern human jawbone finds. These early migrations do not appear to have led to lasting colonisation and receded by about 80,000 years ago. There is a possibility that this first wave of expansion may have reached China (or even North America) as early as 125,000 years ago, but would have died out without leaving a trace in the genome of contemporary humans. There is some evidence that modern humans left Africa at least 125,000 years ago using two different routes: through the
Nile Valley The Nile, , Bohairic , lg, Kiira , Nobiin: Áman Dawū is a major north-flowing river in northeastern Africa. It flows into the Mediterranean Sea. The Nile is the longest river in Africa and has historically been considered the longest rive ...
heading to the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabian Peninsula, Arabia (including the Arabian Peninsula and Bahrain), Anatolia, Asia Minor (Asian part of Turkey except Hatay Pro ...
, at least into modern Israel (
Qafzeh Mount Precipice ( he, הר הקפיצה, "''Har HaKfitsa''"; ar, جبل القفزة, "''Jebel al-Qafzeh''", "Mount of the Leap"), also known as Mount of Precipitation, Mount of the Leap of the Lord and Mount Kedumim is located just outside the ...
: 120,000–100,000 years ago); and a second route through the present-day
Bab-el-Mandeb The Bab-el-Mandeb (Arabic: , , ) is a strait between Yemen on the Arabian Peninsula, and Djibouti and Eritrea in the Horn of Africa. It connects the Red Sea to the Gulf of Aden. Name The strait derives its name from the dangers attendin ...
Strait on the Red Sea (at that time, with a much lower sea level and narrower extension), crossing to the
Arabian Peninsula The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plate ...
and settling in places like the present-day United Arab Emirates (125,000 years ago) and Oman (106,000 years ago), and possibly reaching the Indian Subcontinent (
Jwalapuram Jwalapuram (meaning "City of fire" in Telugu) is an archaeological site in the Kurnool district of Andhra Pradesh, southern India, which shows hominid habitation before and after the Toba event (73 kya) according to the Toba catastrophe theory ...
: 75,000 years ago.) Although no human remains have yet been found in these three places, the apparent similarities between the stone tools found at
Jebel Faya Jebel Faya ( ar, جَبَل ٱلْفَايَة, Jabal Al-Fāyah; FAY-NE1) is an archaeological site and limestone hill or escarpment near Al Madam in the Emirate of Sharjah, the UAE, located about east of the city of Sharjah, and between the sho ...
, those from Jwalapuram and some from Africa suggest that their creators were all modern humans. These findings might give some support to the claim that modern humans from Africa arrived at southern China about 100,000 years ago (
Zhiren Cave Zhiren Cave () is a karstic cave in the Mulan Mountains that overlooks the Hejiang River in Chongzuo, Guangxi, China. Zhiren Cave is an early Late Pleistocene site that has yielded the fossil remains of possibly anatomically modern humans with som ...
,
Zhirendong Zhiren Cave () is a karstic cave in the Mulan Mountains that overlooks the Hejiang River in Chongzuo, Guangxi, China. Zhiren Cave is an early Late Pleistocene site that has yielded the fossil remains of possibly anatomically modern humans with som ...
,
Chongzuo Chongzuo (; za, Cungzcoj) is a prefecture-level city in the south of Guangxi Zhuang Autonomous Region near the Sino-Vietnamese border. It is home to one of China's largest Zhuang populations. Geography and climate Chongzuo is located in sout ...
City: 100,000 years ago; and the
Liujiang hominid The Liujiang men () are among the earliest modern humans (''Homo sapiens'') found in East Asia. Their remains were discovered in the Tongtianyan Cave (通天岩) in Liujiang, Guangxi, China. The remains were excavated in 1958. The remains cons ...
(
Liujiang County Liujiang District (; Standard Zhuang: ) is under the administration of Liuzhou, Guangxi Zhuang Autonomous Region, China, located on the southwest bank of the Liu River. It covers a land area of and had a population of 562,351 . The southernmost co ...
): controversially dated at 139,000–111,000 years ago ). Dating results of the Lunadong ( Bubing Basin,
Guangxi Guangxi (; ; Chinese postal romanization, alternately romanized as Kwanghsi; ; za, Gvangjsih, italics=yes), officially the Guangxi Zhuang Autonomous Region (GZAR), is an Autonomous regions of China, autonomous region of the People's Republic ...
,
southern China South China () is a geographical and cultural region that covers the southernmost part of China. Its precise meaning varies with context. A notable feature of South China in comparison to the rest of China is that most of its citizens are not n ...
) teeth, which include a right upper second molar and a left lower second molar, indicate that the molars may be as old as 126,000 years. Since these previous exits from Africa did not leave traces in the results of genetic analyses based on the Y chromosome and on MtDNA (which represent only a small part of the human genetic material), it seems that those modern humans did not survive in large numbers and were assimilated by our major antecessors. An explanation for their extinction (or small genetic imprint) may be the
Toba eruption The Youngest Toba eruption was a supervolcano eruption that occurred around 74,000 years ago at the site of present-day Lake Toba in Sumatra, Indonesia. It is one of the Earth's largest known explosive eruptions. The Toba catastrophe theory ho ...
(74,000 years ago), though some argue it scarcely affected human population.


Coastal migration

The so-called " recent dispersal" of modern humans took place about 70–50,000 years ago. It is this migration wave that led to the lasting spread of modern humans throughout the world. A small group from a population in East Africa, bearing mitochondrial haplogroup L3 and numbering possibly fewer than 1,000 individuals, crossed the
Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; T ...
strait at
Bab-el-Mandeb The Bab-el-Mandeb (Arabic: , , ) is a strait between Yemen on the Arabian Peninsula, and Djibouti and Eritrea in the Horn of Africa. It connects the Red Sea to the Gulf of Aden. Name The strait derives its name from the dangers attendin ...
, to what is now
Yemen Yemen (; ar, ٱلْيَمَن, al-Yaman), officially the Republic of Yemen,, ) is a country in Western Asia. It is situated on the southern end of the Arabian Peninsula, and borders Saudi Arabia to the Saudi Arabia–Yemen border, north and ...
, after around 75,000 years ago. A recent review has also shown support for the northern route through Sinai/Israel/Syria (Levant). Their descendants spread along the coastal route around
Arabia The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plate. ...
and
Persia Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
to the
Indian subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
before 55,000 years ago. Other research supports a migration out of Africa between about 65,000 and 50,000 years ago. The coastal migration between roughly 70,000 and 50,000 years ago is associated with mitochondrial haplogroups M and N, both derivative of L3. Along the way ''H. sapiens'' interbred with Neanderthals and Denisovans, with Denisovan DNA making 0.2% of mainland Asian and Native American DNA.


Nearby Oceania

Migrations continued along the Asian coast to Southeast Asia and Oceania, colonising
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
by around 65,000–50,000 years ago. By reaching Australia, ''H. sapiens'' for the first time expanded its habitat beyond that of ''H. erectus''. Denisovan ancestry is shared by
Melanesians Melanesians are the predominant and indigenous inhabitants of Melanesia, in a wide area from Indonesia's New Guinea to as far East as the islands of Vanuatu and Fiji. Most speak either one of the many languages of the Austronesian language fa ...
,
Aboriginal Australians Aboriginal Australians are the various Indigenous peoples of the Australian mainland and many of its islands, such as Tasmania, Fraser Island, Hinchinbrook Island, the Tiwi Islands, and Groote Eylandt, but excluding the Torres Strait Islands ...
, and smaller scattered groups of people in Southeast Asia, such as the
Mamanwa The Lumad are a group of Austronesian indigenous people in the southern Philippines. It is a Cebuano term meaning "native" or "indigenous". The term is short for Katawhang Lumad (Literally: "indigenous people"), the autonym officially adopte ...
, a
Negrito The term Negrito () refers to several diverse ethnic groups who inhabit isolated parts of Southeast Asia and the Andaman Islands. Populations often described as Negrito include: the Andamanese peoples (including the Great Andamanese, the Onge, ...
people in the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
, suggesting the interbreeding took place in Eastern Asia where the Denisovans lived. Denisovans may have crossed the
Wallace Line The Wallace Line or Wallace's Line is a faunal boundary line drawn in 1859 by the British naturalist Alfred Russel Wallace and named by English biologist Thomas Henry Huxley that separates the biogeographical realms of Asia and Wallacea, a tran ...
, with
Wallacea Wallacea is a biogeographical designation for a group of mainly Indonesian islands separated by deep-water straits from the Asian and Australian continental shelves. Wallacea includes Sulawesi, the largest island in the group, as well as Lo ...
serving as their last refugium. ''Homo erectus'' had crossed the Lombok gap reaching as far as Flores, but never made it to Australia. During this time sea level was much lower and most of
Maritime Southeast Asia Maritime Southeast Asia comprises the countries of Brunei, Indonesia, Malaysia, the Philippines, Singapore, and East Timor. Maritime Southeast Asia is sometimes also referred to as Island Southeast Asia, Insular Southeast Asia or Oceanic Sout ...
formed one land mass known as
Sunda Sunda may refer to: Europe * Sunda, Faroe Islands India * Sunda (asura), an asura brother of Upasunda * Sunda (clan), a clan (gotra) of Jats in Haryana and Rajasthan, India Southeast Asia * Sundanese (disambiguation) ** Sundanese people ...
. Migration continued Southeast on the coastal route to the
strait A strait is an oceanic landform connecting two seas or two other large areas of water. The surface water generally flows at the same elevation on both sides and through the strait in either direction. Most commonly, it is a narrow ocean channe ...
s between Sunda and
Sahul __NOTOC__ Sahul (), also called Sahul-land, Meganesia, Papualand and Greater Australia, was a paleocontinent that encompassed the modern-day landmasses of mainland Australia, Tasmania, New Guinea, and the Aru Islands. Sahul was in the south-we ...
, the continental land mass of present-day Australia and
New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...
. The gaps on the
Weber Line Max Carl Wilhelm Weber van Bosse or Max Wilhelm Carl Weber (5 December 1852, in Bonn – 7 February 1937, in Eerbeek) was a German-Dutch zoologist and biogeographer. Weber studied at the University of Bonn, then at the Humboldt University in B ...
are up to 90 km wide, so the migration to Australia and New Guinea would have required seafaring skills. Migration also continued along the coast eventually turning northeast to
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
and finally reaching
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
before turning inland. This is evidenced by the pattern of mitochondrial haplogroups descended from haplogroup M, and in
Y-chromosome The Y chromosome is one of two sex chromosomes (allosomes) in therian mammals, including humans, and many other animals. The other is the X chromosome. Y is normally the sex-determining chromosome in many species, since it is the presence or abse ...
haplogroup C. Sequencing of one Aboriginal genome from an old hair sample in
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
revealed that the individual was descended from people who migrated into East Asia between 62,000 and 75,000 years ago. This supports the theory of a single migration into Australia and New Guinea before the arrival of Modern Asians (between 25,000 and 38,000 years ago) and their later migration into North America. This migration is believed to have happened around 50,000 years ago, before Australia and New Guinea were separated by rising sea levels approximately 8,000 years ago. This is supported by a date of 50,000–60,000 years ago for the oldest evidence of settlement in Australia, around 40,000 years ago for the oldest human remains, the earliest humans artifacts which are at least 65,000 years old and the extinction of the
Australian megafauna The term Australian megafauna refers to the megafauna in Australia during the Pleistocene Epoch. Most of these species became extinct during the latter half of the Pleistocene, and the roles of human and climatic factors in their extinction are ...
by humans between 46,000 and 15,000 years ago argued by Tim Flannery, which is similar to what happened in the Americas. The continued use of Stone Age tools in Australia has been much debated.


Dispersal throughout Eurasia

The population brought to
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;;;;; ...
by
coastal migration In the context of the recent African origin of modern humans, the Southern Dispersal scenario (also the coastal migration or great coastal migration hypothesis) refers to the early migration along the southern coast of Asia, from the Arabian Pen ...
appears to have remained there for some time, during roughly 60,000 to 50,000 years ago, before spreading further throughout Eurasia. This dispersal of early humans, at the beginning of the
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
, gave rise to the major population groups of the
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
and the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...
. Towards the West, Upper Paleolithic populations associated with mitochondrial haplogroup R and its derivatives, spread throughout Asia and Europe, with a back-migration of M1 to North Africa and the Horn of Africa several millennia ago. Presence in Europe is certain after 40,000 years ago, possibly as early as 43,000 years ago, rapidly replacing the Neanderthal population. Contemporary Europeans have Neanderthal ancestry, but it seems likely that substantial interbreeding with Neanderthals ceased before 47,000 years ago, i.e. took place before modern humans entered Europe. There is evidence from
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
that modern humans have passed through at least one
genetic bottleneck A population bottleneck or genetic bottleneck is a sharp reduction in the size of a population due to environmental events such as famines, earthquakes, floods, fires, disease, and droughts; or human activities such as specicide, widespread violen ...
, in which genome diversity was drastically reduced.
Henry Harpending Henry Cosad Harpending (January 13, 1944 – April 3, 2016) was an American anthropologist and writer. He was a distinguished professor at the University of Utah, and formerly taught at Penn State and the University of New Mexico. He was a membe ...
has proposed that humans spread from a geographically restricted area about 100,000 years ago, the passage through the geographic bottleneck and then with a dramatic growth amongst geographically dispersed populations about 50,000 years ago, beginning first in Africa and thence spreading elsewhere. Climatological and geological evidence suggests evidence for the bottleneck. The explosion of
Toba Toba may refer to: Languages * Toba Sur language, spoken in South America * Batak Toba, spoken in Indonesia People * Toba people, indigenous peoples of the Gran Chaco in South America * Toba Batak people, a sub-ethnic group of Batak people from ...
, the largest volcanic eruption of the
Quaternary The Quaternary ( ) is the current and most recent of the three periods of the Cenozoic Era in the geologic time scale of the International Commission on Stratigraphy (ICS). It follows the Neogene Period and spans from 2.58 million years ...
, may have created a 1,000 year cold period, potentially reducing human populations to a few tropical refugia. It has been estimated that as few as 15,000 humans survived. In such circumstances genetic drift and
founder effects In population genetics, the founder effect is the loss of genetic variation that occurs when a new population is established by a very small number of individuals from a larger population. It was first fully outlined by Ernst Mayr in 1942, usi ...
may have been maximised. The greater diversity amongst African genomes may reflect the extent of African refugia during the Toba incident. However, a recent review highlights that the single-source hypothesis of non-African populations is less consistent with ancient DNA analysis than multiple sources with genetic mixing across Eurasia.


Europe

The recent expansion of
anatomically modern humans Early modern human (EMH) or anatomically modern human (AMH) are terms used to distinguish ''Homo sapiens'' (the only extant Hominina species) that are anatomically consistent with the range of phenotypes seen in contemporary humans from extin ...
reached Europe around 40,000 years ago from Central Asia and the Middle East, as a result of cultural adaption to big game hunting of sub-glacial steppe fauna.
Neanderthals Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an Extinction, extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ag ...
were present both in the Middle East and in Europe, and the arriving populations of anatomically modern humans (also known as "
Cro-Magnon Early European modern humans (EEMH), or Cro-Magnons, were the first early modern humans (''Homo sapiens'') to settle in Europe, migrating from Western Asia, continuously occupying the continent possibly from as early as 56,800 years ago. They ...
" or
European early modern humans Early European modern humans (EEMH), or Cro-Magnons, were the first early modern humans (''Homo sapiens'') to settle in Europe, migrating from Western Asia, continuously occupying the continent possibly from as early as 56,800 years ago. They ...
) interbred with Neanderthal populations to a limited degree. Populations of modern humans and Neanderthal overlapped in various regions such as the Iberian peninsula and the Middle East. Interbreeding may have contributed Neanderthal genes to palaeolithic and ultimately modern Eurasians and Oceanians. An important difference between Europe and other parts of the inhabited world was the northern latitude. Archaeological evidence suggests humans, whether Neanderthal or Cro-Magnon, reached sites in Arctic Russia by 40,000 years ago. Cro-Magnon are considered the first anatomically modern humans in Europe. They entered
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
by the
Zagros Mountains The Zagros Mountains ( ar, جبال زاغروس, translit=Jibal Zaghrus; fa, کوه‌های زاگرس, Kuh hā-ye Zāgros; ku, چیاکانی زاگرۆس, translit=Çiyakani Zagros; Turkish: ''Zagros Dağları''; Luri: ''Kuh hā-ye Zāgro ...
(near present-day
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
and eastern
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
) around 50,000 years ago, with one group rapidly settling coastal areas around the
Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by th ...
and another migrating north to the steppes of
Central Asia Central Asia, also known as Middle Asia, is a subregion, region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes t ...
. Modern human remains dating to 43,000–45,000 years ago have been discovered in Italy and Britain, as well as in the European Russian Arctic from 40,000 years ago. Humans colonised the environment west of the Urals, hunting reindeer especially, but were faced with adaptive challenges; winter temperatures averaged from with fuel and shelter scarce. They travelled on foot and relied on hunting highly mobile herds for food. These challenges were overcome through technological innovations: tailored clothing from the pelts of fur-bearing animals; construction of shelters with hearths using bones as fuel; and digging "ice cellars" into the permafrost to store meat and bones. A
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
sequence of two Cro-Magnons from the
Paglicci Cave Paglicci Cave is an archaeological site situated in Paglicci, near Rignano Garganico, Apulia, southern Italy. The cave, discovered in the 1950s, is the most important cave of Gargano. The cave is an attraction of the Gargano National Park. Desc ...
in Italy, dated to 23,000 and 24,000 years old (Paglicci 52 and 12), identified the
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...
as Haplogroup N, typical of the latter group. The expansion of modern human population is thought to have begun 45,000 years ago, and it may have taken 15,000–20,000 years for Europe to be colonized. During this time, the Neanderthals were slowly being displaced. Because it took so long for Europe to be occupied, it appears that humans and Neanderthals may have been constantly competing for territory. The Neanderthals had larger brains, and were larger overall, with a more robust or heavily built frame, which suggests that they were physically stronger than modern ''Homo sapiens''. Having lived in Europe for 200,000 years, they would have been better adapted to the cold weather. The anatomically modern humans known as the
Cro-Magnons Early European modern humans (EEMH), or Cro-Magnons, were the first early modern humans (''Homo sapiens'') to settle in Europe, migrating from Western Asia, continuously occupying the continent possibly from as early as 56,800 years ago. They ...
, with widespread trade networks, superior technology and bodies likely better suited to running, would eventually completely displace the Neanderthals, whose last refuge was in the
Iberian peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
. After about 25,000 years ago the fossil record of the Neanderthals ends, indicating extinction. The last known population lived around a cave system on the remote south-facing coast of
Gibraltar ) , anthem = " God Save the King" , song = " Gibraltar Anthem" , image_map = Gibraltar location in Europe.svg , map_alt = Location of Gibraltar in Europe , map_caption = United Kingdom shown in pale green , mapsize = , image_map2 = Gib ...
from 30,000 to 24,000 years ago. From the extent of linkage disequilibrium, it was estimated that the last Neanderthal gene flow into early ancestors of Europeans occurred 47,000–65,000 years BP. In conjunction with archaeological and fossil evidence, interbreeding is thought to have occurred somewhere in Western Eurasia, possibly the Middle East. Studies show a higher Neanderthal admixture in East Asians than in Europeans. North African groups share a similar excess of derived alleles with Neanderthals as non-African populations, whereas Sub-Saharan African groups are the only modern human populations with no substantial Neanderthal admixture. The Neanderthal-linked haplotype B006 of the dystrophin gene has also been found among nomadic pastoralist groups in the Sahel and Horn of Africa, who are associated with northern populations. Consequently, the presence of this B006 haplotype on the northern and northeastern perimeter of Sub-Saharan Africa is attributed to gene flow from a non-African point of origin.


East, Southeast and North Asia

"
Tianyuan man Tianyuan man ( zh, t=田園洞人, s=田园洞人, p=Tiányuándòng Rén) are the remains of one of the earliest modern humans to inhabit East Asia. In 2007, researchers found 34 bone fragments belonging to a single individual at the Tianyuan Ca ...
", an individual who lived in China c. 40,000 years ago, showed substantial Neanderthal admixture. A 2017 study of the ancient DNA of Tianyuan Man found that the individual is related to modern Asian and Native American populations. A 2013 study found Neanderthal introgression of 18 genes within the chromosome 3p21.31 region (HYAL region) of East Asians. The introgressive haplotypes were positively selected in only East Asian populations, rising steadily from 45,000 years ago until a sudden increase of growth rate around 5,000 to 3,500 years ago. They occur at very high frequencies among East Asian populations in contrast to other Eurasian populations (e.g. European and South Asian populations). The findings also suggest that this Neanderthal introgression occurred within the ancestral population shared by East Asians and Native Americans.. A 2016 study presented an analysis of the population genetics of the
Ainu Ainu or Aynu may refer to: *Ainu people, an East Asian ethnic group of Japan and the Russian Far East *Ainu languages, a family of languages **Ainu language of Hokkaido **Kuril Ainu language, extinct language of the Kuril Islands **Sakhalin Ainu la ...
people of northern Japan as key to the reconstruction of the early peopling of East Asia. The Ainu were found to represent a more basal branch than the modern farming populations of East Asia, suggesting an ancient (pre-Neolithic) connection with northeast Siberians. A 2013 study associated several
phenotypical In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology or physical form and structure, its developmental processes, its biochemical and physiological proper ...
traits associated with Mongoloids with a single mutation of the
EDAR Ectodysplasin A receptor (EDAR) is a protein that in humans is encoded by the EDAR gene. EDAR is a cell surface receptor for ectodysplasin A which plays an important role in the development of ectodermal tissues such as the skin. It is structura ...
gene, dated to c. 35,000 years ago. Mitochondrial haplogroups A, B and G originated about 50,000 years ago, and bearers subsequently colonized
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
,
Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic o ...
and
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
, by about 35,000 years ago. Parts of these populations migrated to North America during the
Last Glacial Maximum The Last Glacial Maximum (LGM), also referred to as the Late Glacial Maximum, was the most recent time during the Last Glacial Period that ice sheets were at their greatest extent. Ice sheets covered much of Northern North America, Northern Eur ...
. A review paper by Melinda A. Yang (in 2022) summarized and concluded that a distinctive "Basal-East Asian population" referred to as East- and Southeast Asian lineage''' (ESEA); which is ancestral to modern East Asians,
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
ns,
Polynesians Polynesians form an ethnolinguistic group of closely related people who are native to Polynesia (islands in the Polynesian Triangle), an expansive region of Oceania in the Pacific Ocean. They trace their early prehistoric origins to Island Sou ...
, and
Siberians The Siberians, or Siberiaks, (russian: сибиряки, sibiryaki, ) are the majority inhabitants of Siberia, as well as the (sub)ethnic or ethnographic group of the Russians. As demonym The demonym ''Siberian'' can be restricted to either t ...
, originated in
Mainland Southeast Asia Mainland Southeast Asia, also known as the Indochinese Peninsula or Indochina, is the continental portion of Southeast Asia. It lies east of the Indian subcontinent and south of Mainland China and is bordered by the Indian Ocean to the west an ...
at ~50,000BC, and expanded through multiple migration waves southwards and northwards respectively. This ESEA lineage gave rise to various sublineages, and is also ancestral to the Hoabinhian hunter-gatherers of Southeast Asia and the ~40,000 year old Tianyuan lineage found in
Northern China Northern China () and Southern China () are two approximate regions within China. The exact boundary between these two regions is not precisely defined and only serve to depict where there appears to be regional differences between the climate ...
, but already differentiated and distinct from European-related and Australasian-related lineages, found in other regions of prehistoric Eurasia. The ESEA lineage trifurcated from an earlier East-Eurasian or "eastern non-African" (ENA) lineage somewhere in
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;;;;; ...
, which also contributed to the formation of indigenous South Asian hunter-gatherers (AASI) as well as to Australasians.


Last Glacial Maximum


Eurasia

Around 20,000 years ago, approximately 5,000 years after the Neanderthal extinction, the
Last Glacial Maximum The Last Glacial Maximum (LGM), also referred to as the Late Glacial Maximum, was the most recent time during the Last Glacial Period that ice sheets were at their greatest extent. Ice sheets covered much of Northern North America, Northern Eur ...
forced northern hemisphere inhabitants to migrate to several shelters ( refugia) until the end of this period. The resulting populations are presumed to have resided in such refuges during the LGM to ultimately reoccupy Europe, where archaic historical populations are considered their descendants. The composition of European populations was later altered by further migrations, notably the
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several parts ...
expansion from the Middle East, and still later the
Chalcolithic The Copper Age, also called the Chalcolithic (; from grc-gre, χαλκός ''khalkós'', "copper" and  ''líthos'', "stone") or (A)eneolithic (from Latin '' aeneus'' "of copper"), is an archaeological period characterized by regular ...
population movements associated with
Indo-European expansion The Indo-European migrations were hypothesized migrations of Proto-Indo-European language (PIE) speakers, and subsequent migrations of people speaking derived Indo-European languages, which took place approx. 4000 to 1000 BCE, potentially expla ...
. A Paleolithic site on the Yana River, Siberia, at 71°N, lies well above the Arctic Circle and dates to 27,000 radiocarbon years before present, during glacial times. This site shows that people adapted to this harsh, high-latitude, Late Pleistocene environment much earlier than previously thought.


Americas

Paleo-Indians originated from
Central Asia Central Asia, also known as Middle Asia, is a subregion, region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes t ...
, crossing the Beringia land bridge between eastern Siberia and present-day Alaska. Humans lived throughout the Americas by the end of the last glacial period, or more specifically what is known as the
late glacial maximum The Last Glacial Maximum (LGM), also referred to as the Late Glacial Maximum, was the most recent time during the Last Glacial Period that ice sheets were at their greatest extent. Ice sheets covered much of Northern North America, Northern Eur ...
. Details of Paleo-Indian migration to and throughout the American continent, including the dates and the routes traveled, are subject to ongoing research and discussion. Conventional estimates have it that humans reached North America at some point between 15,000 and 20,000 years ago. The traditional theory is that these early migrants moved when sea levels were significantly lowered due to the
Quaternary glaciation The Quaternary glaciation, also known as the Pleistocene glaciation, is an alternating series of glacial and interglacial periods during the Quaternary period that began 2.58 Ma (million years ago) and is ongoing. Although geologists describe ...
, following herds of now-extinct
pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
megafauna In terrestrial zoology, the megafauna (from Greek μέγας ''megas'' "large" and New Latin ''fauna'' "animal life") comprises the large or giant animals of an area, habitat, or geological period, extinct and/or extant. The most common threshold ...
along ''ice-free corridors'' that stretched between the
Laurentide The Laurentide Ice Sheet was a massive sheet of ice that covered millions of square miles, including most of Canada and a large portion of the Northern United States, multiple times during the Quaternary glacial epochs, from 2.58 million years a ...
and Cordilleran ice sheets. Another route proposed is that, either on foot or using primitive boats, they migrated down the Pacific coast to
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
as far as
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
. Any archaeological evidence of coastal occupation during the last Ice Age would now have been covered by the
sea level rise Globally, sea levels are rising due to human-caused climate change. Between 1901 and 2018, the globally averaged sea level rose by , or 1–2 mm per year on average.IPCC, 2019Summary for Policymakers InIPCC Special Report on the Ocean and Cry ...
, up to a hundred metres since then. The recent finding of indigenous
Australasia Australasia is a region that comprises Australia, New Zealand and some neighbouring islands in the Pacific Ocean. The term is used in a number of different contexts, including geopolitically, physiogeographically, philologically, and ecologica ...
n genetic markers in Amazonia supports that a coastal route and subsequent isolation did occur with some migrants.


Holocene migrations

The
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...
is taken to begin 12,000 years ago, after the end of the
Last Glacial Maximum The Last Glacial Maximum (LGM), also referred to as the Late Glacial Maximum, was the most recent time during the Last Glacial Period that ice sheets were at their greatest extent. Ice sheets covered much of Northern North America, Northern Eur ...
. During the
Holocene climatic optimum The Holocene Climate Optimum (HCO) was a warm period that occurred in the interval roughly 9,000 to 5,000 years ago BP, with a thermal maximum around 8000 years BP. It has also been known by many other names, such as Altithermal, Climatic Optimu ...
, beginning about 9,000 years ago, human populations which had been geographically confined to refugia began to migrate. By this time, most parts of the globe had been settled by ''H. sapiens''; however, large areas that had been covered by
glaciers A glacier (; ) is a persistent body of dense ice that is constantly moving under its own weight. A glacier forms where the accumulation of snow exceeds its ablation over many years, often centuries. It acquires distinguishing features, such as ...
were now re-populated. This period sees the transition from the
Mesolithic The Mesolithic (Greek: μέσος, ''mesos'' 'middle' + λίθος, ''lithos'' 'stone') or Middle Stone Age is the Old World archaeological period between the Upper Paleolithic and the Neolithic. The term Epipaleolithic is often used synonymous ...
to the
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several parts ...
stage throughout the
temperate zone In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
. The Neolithic subsequently gives way to the
Bronze Age The Bronze Age is a historic period, lasting approximately from 3300 BC to 1200 BC, characterized by the use of bronze, the presence of writing in some areas, and other early features of urban civilization. The Bronze Age is the second pri ...
in
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
cultures and the gradual emergence of the
historical record Recorded history or written history describes the historical events that have been recorded in a written form or other documented communication which are subsequently evaluated by historians using the historical method. For broader world hist ...
in the
Near East The ''Near East''; he, המזרח הקרוב; arc, ܕܢܚܐ ܩܪܒ; fa, خاور نزدیک, Xāvar-e nazdik; tr, Yakın Doğu is a geographical term which roughly encompasses a transcontinental region in Western Asia, that was once the hist ...
and
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
beginning around 4,000 years ago. Large-scale migrations of the Mesolithic to Neolithic era are thought to have given rise to the pre-modern distribution of the world's major
language families A language family is a group of languages related through descent from a common ''ancestral language'' or ''parental language'', called the proto-language of that family. The term "family" reflects the tree model of language origination in hi ...
such as the Niger-Congo,
Nilo-Saharan The Nilo-Saharan languages are a proposed family of African languages spoken by some 50–60 million people, mainly in the upper parts of the Chari and Nile rivers, including historic Nubia, north of where the two tributaries of the Nile meet. T ...
,
Afro-Asiatic The Afroasiatic languages (or Afro-Asiatic), also known as Hamito-Semitic, or Semito-Hamitic, and sometimes also as Afrasian, Erythraean or Lisramic, are a language family of about 300 languages that are spoken predominantly in the geographic su ...
,
Uralic The Uralic languages (; sometimes called Uralian languages ) form a language family of 38 languages spoken by approximately 25million people, predominantly in Northern Eurasia. The Uralic languages with the most native speakers are Hungarian (w ...
,
Sino-Tibetan Sino-Tibetan, also cited as Trans-Himalayan in a few sources, is a family of more than 400 languages, second only to Indo-European in number of native speakers. The vast majority of these are the 1.3 billion native speakers of Chinese languages. ...
or
Indo-European The Indo-European languages are a language family native to the overwhelming majority of Europe, the Iranian plateau, and the northern Indian subcontinent. Some European languages of this family, English, French, Portuguese, Russian, Dutch ...
phyla. The speculative Nostratic theory postulates the derivation of the major language families of Eurasia (excluding Sino-Tibetan) from a single proto-language spoken at the beginning of the Holocene period.


Eurasia

Evidence published in 2014 from genome analysis of ancient human remains suggests that the modern native populations of Europe largely descend from three distinct lineages: "
Western Hunter-Gatherers In archaeogenetics, the term Western Hunter-Gatherer (WHG), West European Hunter-Gatherer or Western European Hunter-Gatherer names a distinct ancestral component of modern Europeans, representing descent from a population of Mesolithic hunter-gat ...
", derivative of the Cro-Magnon population of Europe,
Early European Farmers Early European Farmers (EEF), First European Farmers (FEF), Neolithic European Farmers, Ancient Aegean Farmers, or Anatolian Neolithic Farmers (ANF) are names used to describe a distinct group of early Neolithic farmers who brought agriculture to E ...
introduced to Europe from the Near East during the
Neolithic Revolution The Neolithic Revolution, or the (First) Agricultural Revolution, was the wide-scale transition of many human cultures during the Neolithic period from a lifestyle of hunting and gathering to one of agriculture and settlement, making an incre ...
and
Ancient North Eurasians In archaeogenetics, the term Ancient North Eurasian (generally abbreviated as ANE) is the name given to an ancestral component that represents a lineage ancestral to the people of the Mal'ta–Buret' culture and populations closely related to the ...
which expanded to Europe in the context of the
Indo-European expansion The Indo-European migrations were hypothesized migrations of Proto-Indo-European language (PIE) speakers, and subsequent migrations of people speaking derived Indo-European languages, which took place approx. 4000 to 1000 BCE, potentially expla ...
. The Ancient North Eurasian component was introduced to Western Europe by people related to the
Yamnaya culture The Yamnaya culture or the Yamna culture (russian: Ямная культура, ua, Ямна культура literal translation, lit. 'culture of pits'), also known as the Pit Grave culture or Ochre Grave culture, was a late Copper Age to ea ...
. Additional ANE ancestry is found in European populations through Paleolithic interactions with
Eastern Hunter-Gatherer In archaeogenetics, the term Eastern Hunter-Gatherer (EHG), sometimes East European Hunter-Gatherer, or Eastern European Hunter-Gatherer is the name given to a distinct ancestral component that represents descent from Mesolithic hunter-gatherers of ...
s.


Sub-Saharan Africa

West-Eurasian back-migrations started in the early
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...
or already earlier in the
Paleolithic The Paleolithic or Palaeolithic (), also called the Old Stone Age (from Greek: παλαιός ''palaios'', "old" and λίθος ''lithos'', "stone"), is a period in human prehistory that is distinguished by the original development of stone too ...
period (30-15kya),followed by pre-Neolithic and
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several parts ...
migration events from the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabian Peninsula, Arabia (including the Arabian Peninsula and Bahrain), Anatolia, Asia Minor (Asian part of Turkey except Hatay Pro ...
, mostly affecting Northern Africa, the Horn of Africa, and wider regions of the Sahel zone and East Africa. The
Nilotic peoples The Nilotic peoples are people Indigenous peoples of Africa, indigenous to the Nile Valley who speak Nilotic languages. They inhabit South Sudan, Sudan, Ethiopia, Uganda, Kenya, the Democratic Republic of the Congo, Rwanda, Burundi and Tanzania. ...
are thought to be derived from an earlier undifferentiated
Eastern Sudanic In most classifications, the Eastern Sudanic languages are a group of nine families of languages that may constitute a branch of the Nilo-Saharan language family. Eastern Sudanic languages are spoken from southern Egypt to northern Tanzania. Nub ...
unity by the 3rd millennium BCE. The development of the Proto-Nilotes as a group may have been connected with their domestication of
livestock Livestock are the domesticated animals raised in an agricultural setting to provide labor and produce diversified products for consumption such as meat, eggs, milk, fur, leather, and wool. The term is sometimes used to refer solely to animals ...
. The Eastern Sudanic unity must have been considerably earlier still, perhaps around the 5th millennium BCE (while the proposed
Nilo-Saharan The Nilo-Saharan languages are a proposed family of African languages spoken by some 50–60 million people, mainly in the upper parts of the Chari and Nile rivers, including historic Nubia, north of where the two tributaries of the Nile meet. T ...
unity would date to the
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
about 15kya). The original locus of the early Nilotic speakers was presumably east of the Nile in what is now
South Sudan South Sudan (; din, Paguot Thudän), officially the Republic of South Sudan ( din, Paankɔc Cuëny Thudän), is a landlocked country in East Africa. It is bordered by Ethiopia, Sudan, Central African Republic, Democratic Republic of the C ...
. The Proto-Nilotes of the 3rd millennium BCE were
pastoralists Pastoralism is a form of animal husbandry where domesticated animals (known as "livestock") are released onto large vegetated outdoor lands (pastures) for grazing, historically by nomadic people who moved around with their herds. The animal ...
, while their neighbors, the Proto-
Central Sudanic Central Sudanic is a family of about sixty languages that have been included in the proposed Nilo-Saharan language family. Central Sudanic languages are spoken in the Central African Republic, Chad, South Sudan, Uganda, Congo (DRC), Nigeria and ...
peoples, were mostly agriculturalists. The Niger-Congo phylum is thought to have emerged around 6,000 years ago in West or Central Africa. Its expansion may have been associated with the expansion of Sahel agriculture in the African Neolithic period, following the desiccation of the Sahara in c. 3900 BCE. The
Bantu expansion The Bantu expansion is a hypothesis about the history of the major series of migrations of the original Proto-Bantu-speaking group, which spread from an original nucleus around Central Africa across much of sub-Saharan Africa. In the process, t ...
has spread the
Bantu languages The Bantu languages (English: , Proto-Bantu: *bantʊ̀) are a large family of languages spoken by the Bantu people of Central, Southern, Eastern africa and Southeast Africa. They form the largest branch of the Southern Bantoid languages. The t ...
to Central, Eastern and Southern Africa, partly replacing the
indigenous Indigenous may refer to: *Indigenous peoples *Indigenous (ecology), presence in a region as the result of only natural processes, with no human intervention *Indigenous (band), an American blues-rock band *Indigenous (horse), a Hong Kong racehorse ...
populations of these regions, including the
African Pygmies The African Pygmies (or Congo Pygmies, variously also Central African foragers, "African rainforest hunter-gatherers" (RHG) or "Forest People of Central Africa") are a group of ethnicities native to Central Africa, mostly the Congo Basin, trad ...
,
Hadza people The Hadza, or Hadzabe (''Wahadzabe'' in Swahili), are a Tanzanian indigenous ethnic group mostly based in southwest Karatu District of Arusha Region. They live around Lake Eyasi in the central Rift Valley and in the neighboring Serengeti Platea ...
and
San people The San peoples (also Saan), or Bushmen, are members of various Khoe, Tuu, or Kxʼa-speaking indigenous hunter-gatherer cultures that are the first cultures of Southern Africa, and whose territories span Botswana, Namibia, Angola, Zambia, ...
. Beginning about 3,000 years ago, it reached South Africa about 1,700 years ago. Some evidence (including a 2016 study by Busby et al.) suggests admixture from ancient and recent migrations from
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
into parts of Sub-Saharan Africa. Another study (Ramsay et al. 2018) also shows evidence that ancient Eurasians migrated into Africa and that Eurasian admixture in modern Sub-Saharan Africans ranges from 0% to 50%, varying by region and generally higher in the Horn of Africa and parts of the
Sahel The Sahel (; ar, ساحل ' , "coast, shore") is a region in North Africa. It is defined as the ecoclimatic and biogeographic realm of transition between the Sahara to the north and the Sudanian savanna to the south. Having a hot semi-arid c ...
zone, and found to a lesser degree in certain parts of Western Africa, and
Southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number of ...
(excluding recent immigrants).


Indo-Pacific

The first seaborne human migrations were by the
Austronesian peoples The Austronesian peoples, sometimes referred to as Austronesian-speaking peoples, are a large group of peoples in Taiwan, Maritime Southeast Asia, Micronesia, coastal New Guinea, Island Melanesia, Polynesia, and Madagascar that speak Austro ...
originating from
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
known as the "
Austronesian expansion The Austronesian peoples, sometimes referred to as Austronesian-speaking peoples, are a large group of peoples in Taiwan, Maritime Southeast Asia, Micronesia, coastal New Guinea, Island Melanesia, Polynesia, and Madagascar that speak Austron ...
". Using advanced sailing technologies like
catamaran A Formula 16 beachable catamaran Powered catamaran passenger ferry at Salem, Massachusetts, United States A catamaran () (informally, a "cat") is a multi-hulled watercraft featuring two parallel hulls of equal size. It is a geometry-stab ...
s,
outrigger boat Outrigger boats are various watercraft featuring one or more lateral support floats known as outriggers, which are fastened to one or both sides of the main hull. They can range from small dugout canoes to large plank-built vessels. Outrigger ...
s, and
crab claw sail The crab claw sail is a fore-and-aft triangular sail with spars along upper and lower edges. The crab claw sail was first developed by the Austronesian peoples some time around 1500 BC. It is used in many traditional Austronesian cultures in Isl ...
s, they built the first sea-going ships and rapidly colonized
Island Southeast Asia Maritime Southeast Asia comprises the countries of Brunei, Indonesia, Malaysia, the Philippines, Singapore, and East Timor. Maritime Southeast Asia is sometimes also referred to as Island Southeast Asia, Insular Southeast Asia or Oceanic Sout ...
at around 3000 to 1500 BCE. From the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
and
Eastern Indonesia This is a list of some of the regions of Indonesia. Many regions are defined in law or regulations by the central government. At different times of Indonesia's history, the nation has been designated as having regions that do not necessarily corr ...
they colonized
Micronesia Micronesia (, ) is a subregion of Oceania, consisting of about 2,000 small islands in the western Pacific Ocean. It has a close shared cultural history with three other island regions: the Philippines to the west, Polynesia to the east, and ...
by 2200 to 1000 BCE. A branch of the Austronesians reached
Island Melanesia Island Melanesia is a subregion of Melanesia in Oceania. It is located east of New Guinea island, from the Bismarck Archipelago to New Caledonia.Steadman, 2006. ''Extinction & biogeography of tropical Pacific birds'' See also Archaeology a ...
between 1600 and 1000 BCE, establishing the
Lapita culture The Lapita culture is the name given to a Neolithic Austronesian people and their material culture, who settled Island Melanesia via a seaborne migration at around 1600 to 500 BCE. They are believed to have originated from the northern Philipp ...
(named after the archaeological site in Lapita,
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
, where their characteristic pottery was first discovered). They are the direct ancestors of the modern
Polynesians Polynesians form an ethnolinguistic group of closely related people who are native to Polynesia (islands in the Polynesian Triangle), an expansive region of Oceania in the Pacific Ocean. They trace their early prehistoric origins to Island Sou ...
. They ventured into
Remote Oceania Remote Oceania is the part of Oceania settled within the last 3,000 to 3,500 years, comprising south-eastern Island Melanesia and islands in the open Pacific east of the Solomon Islands: Fiji, Micronesia, New Caledonia, New Zealand, Polynesia, t ...
reaching
Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (french: link=no, République de Vanuatu; bi, Ripablik blong Vanuatu), is an island country located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of no ...
,
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
, and
Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists ...
by 1200 BCE, and
Samoa Samoa, officially the Independent State of Samoa; sm, Sāmoa, and until 1997 known as Western Samoa, is a Polynesian island country consisting of two main islands (Savai'i and Upolu); two smaller, inhabited islands (Manono Island, Manono an ...
and
Tonga Tonga (, ; ), officially the Kingdom of Tonga ( to, Puleʻanga Fakatuʻi ʻo Tonga), is a Polynesian country and archipelago. The country has 171 islands – of which 45 are inhabited. Its total surface area is about , scattered over in ...
by around 900 to 800 BCE. This was the furthest extent of the Lapita culture expansion. During a period of around 1,500 years, they gradually lost the technology for pottery (likely due to the lack of clay deposits in the islands), replacing it with carved wooden and bamboo containers. Back-migrations from the Lapita culture also merged back Island Southeast Asia in 1500 BCE, and into Micronesia at around 200 BCE. It was not until 700 CE when they started voyaging further into the Pacific Ocean, when they colonized the
Cook Islands ) , image_map = Cook Islands on the globe (small islands magnified) (Polynesia centered).svg , capital = Avarua , coordinates = , largest_city = Avarua , official_languages = , lan ...
, the
Society Islands The Society Islands (french: Îles de la Société, officially ''Archipel de la Société;'' ty, Tōtaiete mā) are an archipelago located in the South Pacific Ocean. Politically, they are part of French Polynesia, an overseas country of the F ...
, and the
Marquesas The Marquesas Islands (; french: Îles Marquises or ' or '; Marquesan: ' ( North Marquesan) and ' ( South Marquesan), both meaning "the land of men") are a group of volcanic islands in French Polynesia, an overseas collectivity of France in t ...
. From there, they further colonized
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
by 900 CE,
Rapa Nui Easter Island ( rap, Rapa Nui; es, Isla de Pascua) is an island and special territory of Chile in the southeastern Pacific Ocean, at the southeasternmost point of the Polynesian Triangle in Oceania. The island is most famous for its nearly ...
by 1000 CE, and
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
by 1200 CE. In the
Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by th ...
, Austronesians from
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and eas ...
also colonized
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
and the
Comoros Islands The Comoro Islands or Comoros (Shikomori ''Komori''; ar, جزر القمر , ''Juzur al-qamar''; french: Les Comores) form an archipelago of volcanic islands situated off the southeastern coast of Africa, to the east of Mozambique and northwe ...
by around 500 CE. Austronesians remain the dominant ethnolinguistic group of the islands of the Indo-Pacific, and were the first to establish a maritime trade network reaching as far west as
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
and the
Arabian peninsula The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plate ...
. They assimilated earlier
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
to early
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...
human overland migrations through
Sundaland Sundaland (also called Sundaica or the Sundaic region) is a biogeographical region of South-eastern Asia corresponding to a larger landmass that was exposed throughout the last 2.6 million years during periods when sea levels were lower. It ...
like the
Papuans The indigenous peoples of West Papua in Indonesia and Papua New Guinea, commonly called Papuans, are Melanesians. There is genetic evidence for two major historical lineages in New Guinea and neighboring islands: a first wave from the Malay Arch ...
and the
Negrito The term Negrito () refers to several diverse ethnic groups who inhabit isolated parts of Southeast Asia and the Andaman Islands. Populations often described as Negrito include: the Andamanese peoples (including the Great Andamanese, the Onge, ...
s in Island Southeast Asia. The Austronesian expansion was the last and the most far-reaching
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several parts ...
human migration event.


Caribbean

The
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
was one of the last places in the Americas that were settled by humans. The oldest remains are known from the Greater Antilles (Cuba and Hispaniola) dating between 4000 and 3500 BCE, and comparisons between tool-technologies suggest that these peoples moved across the Yucatán Channel from Central America. All evidence suggests that later migrants from 2000 BCE and onwards originated from South America, via the Orinoco region. The descendants of these migrants include the ancestors of the
Taíno The Taíno were a historic Indigenous peoples of the Caribbean, indigenous people of the Caribbean whose culture has been continued today by Taíno descendant communities and Taíno revivalist communities. At the time of European contact in the ...
and
Kalinago The Kalinago, also known as the Island Caribs or simply Caribs, are an indigenous people of the Lesser Antilles in the Caribbean. They may have been related to the Mainland Caribs (Kalina) of South America, but they spoke an unrelated language ...
(Island Carib) peoples.


Arctic

The earliest inhabitants of North America's central and eastern Arctic are referred to as the
Arctic small tool tradition The Arctic Small Tool tradition (ASTt) was a broad cultural entity that developed along the Alaska Peninsula, around Bristol Bay, and on the eastern shores of the Bering Strait around 2500 BC. ASTt groups were the first human occupants of Arctic ...
(AST) and existed c. 2500 BCE. AST consisted of several
Paleo-Eskimo The Paleo-Eskimo (also pre-Thule or pre-Inuit) were the peoples who inhabited the Arctic region from Chukotka Autonomous Okrug, Chukotka (e.g., Chertov Ovrag) in present-day Russia across North America to Greenland prior to the arrival of the mode ...
cultures, including the Independence cultures and
Pre-Dorset The Pre-Dorset is a loosely defined term for a Paleo-Eskimo culture or group of cultures that existed in the Eastern Canadian Arctic from c. 3200 to 850 cal BC, and preceded the Dorset culture. Due to its vast geographical expanse and to history ...
culture.Gibbon, pp. 28–31 The
Inuit Inuit (; iu, ᐃᓄᐃᑦ 'the people', singular: Inuk, , dual: Inuuk, ) are a group of culturally similar indigenous peoples inhabiting the Arctic and subarctic regions of Greenland, Labrador, Quebec, Nunavut, the Northwest Territories ...
are the descendants of the
Thule culture The Thule (, , ) or proto-Inuit were the ancestors of all modern Inuit. They developed in coastal Alaska by the year 1000 and expanded eastward across northern Canada, reaching Greenland by the 13th century. In the process, they replaced people o ...
, which emerged from western Alaska around CE 1000 and gradually displaced the Dorset culture.


See also

*
List of first human settlements This is a list of dates associated with the prehistoric peopling of the world (first known presence of ''Homo sapiens''). The list is divided into four categories, Middle Paleolithic (before 50,000 years ago), Upper Paleolithic (50,000 to 12,500 ...
*
Middle Paleolithic The Middle Paleolithic (or Middle Palaeolithic) is the second subdivision of the Paleolithic or Old Stone Age as it is understood in Europe, Africa and Asia. The term Middle Stone Age is used as an equivalent or a synonym for the Middle Paleoli ...
*
Quaternary extinction The Quaternary period (from 2.588 ± 0.005 million years ago to the present) has seen the extinctions of numerous predominantly megafaunal species, which have resulted in a collapse in faunal density and diversity and the extinction of key ecol ...
*
Timeline of human evolution The timeline of human evolution outlines the major events in the evolutionary lineage of the modern human species, ''Homo sapiens'', throughout the history of life, beginning some 4 billion years ago down to recent evolution within ''H. sapiens ...
*
Timeline of maritime migration and exploration This timeline is an incomplete list of significant events of human migration and exploration by sea. This timeline does not include migration and exploration over land, including migration across land that has subsequently submerged beneath the s ...


Notes


References


Further reading

* * Diamond, Jared, '' Guns, germs and steel. A short history of everybody for the last 13,000 years'', 1997. * * ; *


External links


Journey of Mankind – Genetic Map
Bradshaw Foundation The Gwion Gwion rock paintings, Gwion figures, Kiro Kiro or Kujon (previously known as the Bradshaw rock paintings, Bradshaw rock art, Bradshaw figures and the Bradshaws) are one of the two major regional traditions of rock art found in the nort ...

Prehistoric Human Migration From Africa to World Video May 2015
{{Human Evolution Prehistoric migrations
Migration Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...
Paleolithic Early migration Demographic history Global events