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The Pliocene ( ; also Pleiocene) is the
epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided by ...
in the
geologic time scale The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochrono ...
that extends from 5.333 million to 2.58See the 2014 version of the ICS geologic time scale
million years ago. It is the second and most recent epoch of the
Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...
Period in the
Cenozoic Era The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
. The Pliocene follows the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
Epoch and is followed by the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
Epoch. Prior to the 2009 revision of the geologic time scale, which placed the four most recent major glaciations entirely within the Pleistocene, the Pliocene also included the
Gelasian The Gelasian is an age (geology), age in the international geologic timescale or a stage (stratigraphy), stage in chronostratigraphy, being the earliest or lowest subdivision of the Quaternary Period/System and Pleistocene Epoch/Series. It spans ...
Stage, which lasted from 2.588 to 1.806 million years ago, and is now included in the Pleistocene. As with other older geologic periods, the
geological strata In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as ei ...
that define the start and end are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The boundaries defining the Pliocene are not set at an easily identified worldwide event but rather at regional boundaries between the warmer Miocene and the relatively cooler Pliocene. The upper boundary was set at the start of the Pleistocene glaciations.


Etymology

Charles Lyell Sir Charles Lyell, 1st Baronet, (14 November 1797 – 22 February 1875) was a Scottish geologist who demonstrated the power of known natural causes in explaining the earth's history. He is best known as the author of ''Principles of Geolo ...
(later Sir Charles) gave the Pliocene its name in ''Principles of Geology'' (volume 3, 1833). The word ''pliocene'' comes from the Greek words (, "more") and (, "new" or "recent") and means roughly "continuation of the recent", referring to the essentially modern marine
mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
fauna.


Subdivisions

In the official timescale of the ICS, the Pliocene is subdivided into two
stages Stage or stages may refer to: Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly British theatre newspaper * S ...
. From youngest to oldest they are: *
Piacenzian The Piacenzian is in the international geologic time scale the upper stage (stratigraphy), stage or latest age (geology), age of the Pliocene. It spans the time between 3.6 ± 0.005 year#SI prefix multipliers, Ma and 2.588 ± 0.005 Ma (million yea ...
(3.600–2.58 Ma) *
Zanclean The Zanclean is the lowest stage or earliest age on the geologic time scale of the Pliocene. It spans the time between 5.332 ± 0.005 Ma (million years ago) and 3.6 ± 0.005 Ma. It is preceded by the Messinian Age of the Miocene Epoch, and fol ...
(5.333–3.600 Ma) The Piacenzian is sometimes referred to as the Late Pliocene, whereas the Zanclean is referred to as the Early Pliocene. In the system of *
North American Land Mammal Ages The North American land mammal ages (NALMA) establishes a geologic timescale for North American fauna beginning during the Late Cretaceous and continuing through to the present. These periods are referred to as ages or intervals (or stages when ref ...
(NALMA) include
Hemphillian The Hemphillian North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 10,300,000 to 4,900,000 years BP. It is usually considered ...
(9–4.75 Ma), and
Blancan The Blancan North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 4,750,000 to 1,806,000 years Before Present, BP, a period of .
(4.75–1.6 Ma). The Blancan extends forward into the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
. *
South American Land Mammal Ages The South American land mammal ages (SALMA) establish a geologic timescale for prehistoric South American fauna beginning 64.5 Ma during the Paleocene and continuing through to the Late Pleistocene (0.011 Ma). These periods are referred to as age ...
(SALMA) include
Montehermosan The Montehermosan age is a period of geologic time (6.8–4.0 Ma) within the Miocene and Pliocene epochs of the Neogene used more specifically with South American Land Mammal Ages. It follows the Huayquerian and precedes the Chapadmalalan The C ...
(6.8–4.0 Ma),
Chapadmalalan The Chapadmalalan age is a period of geologic time (4.0–3.0 Ma) within the Pliocene epoch of the Neogene used more specifically with South American Land Mammal Ages. It follows the Montehermosan and precedes the Uquian age. Fossil content F ...
(4.0–3.0 Ma) and
Uquian The Uquian age is a period of geologic time (3.0–1.5 Ma) within the Pliocene and Pleistocene epochs of the Neogene used more specifically with South American Land Mammal Ages. It follows the Montehermosan and precedes the Ensenadan The Ensenad ...
(3.0–1.2 Ma). In the
Paratethys The Paratethys sea, Paratethys ocean, Paratethys realm or just Paratethys was a large shallow inland sea that stretched from the region north of the Alps over Central Europe to the Aral Sea in Central Asia. Paratethys was peculiar due to its pa ...
area (central
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
and parts of western Asia) the Pliocene contains the Dacian (roughly equal to the Zanclean) and Romanian (roughly equal to the Piacenzian and Gelasian together) stages. As usual in stratigraphy, there are many other regional and local subdivisions in use. In
Britain Britain most often refers to: * The United Kingdom, a sovereign state in Europe comprising the island of Great Britain, the north-eastern part of the island of Ireland and many smaller islands * Great Britain, the largest island in the United King ...
, the Pliocene is divided into the following stages (old to young): Gedgravian, Waltonian, Pre-Ludhamian, Ludhamian, Thurnian, Bramertonian or Antian,
Pre-Pastonian The Pre-Pastonian Stage or Baventian Stage (from Easton Bavents in Suffolk), is the name for an early Pleistocene stage used in the British Isles. It precedes the Pastonian Stage and follows the Bramertonian Stage. This stage ended 1.806 Ma (millio ...
or Baventian, Pastonian and Beestonian. In the
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
the Pliocene is divided into these stages (old to young): Brunssumian C, Reuverian A, Reuverian B, Reuverian C,
Praetiglian The Tiglian, also referred to as the Tegelen, is a temperate complex stage in the glacial history of Northern Europe. It is preceded by the Praetiglian (stage). The stage was introduced by Zagwijn in 1957 based on geological formations in Tegelen i ...
,
Tiglian The Tiglian, also referred to as the Tegelen, is a temperate complex stage in the glacial history of Northern Europe. It is preceded by the Praetiglian (stage). The stage was introduced by Zagwijn in 1957 based on geological formations in Tegelen i ...
A, Tiglian B, Tiglian C1-4b, Tiglian C4c, Tiglian C5, Tiglian C6 and
Eburonian The Eburonian (german: Eburon or ''Eburonium''), or, much less commonly, the Eburonian Stage, is a glacial complex in the Calabrian age of the Pleistocene epoch and lies between the Tegelen and the Waalian interglacial. The transition from the Te ...
. The exact correlations between these local stages and the
International Commission on Stratigraphy The International Commission on Stratigraphy (ICS), sometimes referred to unofficially as the "International Stratigraphic Commission", is a daughter or major subcommittee grade scientific daughter organization that concerns itself with stratigra ...
(ICS) stages is still a matter of detail.


Climate

The beginning of the Pliocene was marked by an increase in global temperatures relative to the cooler
Messinian The Messinian is in the geologic timescale the last age or uppermost stage of the Miocene. It spans the time between 7.246 ± 0.005 Ma and 5.333 ± 0.005 Ma (million years ago). It follows the Tortonian and is followed by the Zanclean, the first ...
related to the 1.2 million year obliquity amplitude modulation cycle. The global average temperature in the mid-Pliocene (3.3–3 mya) was 2–3 °C higher than today, carbon dioxide levels were the same as today, and global sea level was 25 m higher. The northern hemisphere ice sheet was ephemeral before the onset of extensive
glaciation A glacial period (alternatively glacial or glaciation) is an interval of time (thousands of years) within an ice age that is marked by colder temperatures and glacier advances. Interglacials, on the other hand, are periods of warmer climate betw ...
over
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...
that occurred in the late Pliocene around 3 Ma. The formation of an Arctic ice cap is signaled by an abrupt shift in
oxygen Oxygen is the chemical element with the symbol O and atomic number 8. It is a member of the chalcogen group in the periodic table, a highly reactive nonmetal, and an oxidizing agent that readily forms oxides with most elements as wel ...
isotope Isotopes are two or more types of atoms that have the same atomic number (number of protons in their nuclei) and position in the periodic table (and hence belong to the same chemical element), and that differ in nucleon numbers (mass numbers) ...
ratios and
ice-rafted Ice rafting is the transport of various materials by ice. Various objects deposited on ice may eventually become embedded in the ice. When the ice melts after a certain amount of drifting, these objects are deposited onto the bottom of the water ...
cobbles in the
North Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the "Old World" of Africa, Europe and ...
and
North Pacific Ocean North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating direction or geography. Etymology The word ''north'' is ...
beds.Van Andel (1994), p. 226. Mid-latitude glaciation was probably underway before the end of the epoch. The global cooling that occurred during the Pliocene may have spurred on the disappearance of forests and the spread of grasslands and savannas.


Paleogeography

Continents continued to
drift Drift or Drifts may refer to: Geography * Drift or ford (crossing) of a river * Drift, Kentucky, unincorporated community in the United States * In Cornwall, England: ** Drift, Cornwall, village ** Drift Reservoir, associated with the village ...
, moving from positions possibly as far as 250 km from their present locations to positions only 70 km from their current locations.
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
became linked to North America through the
Isthmus of Panama The Isthmus of Panama ( es, Istmo de Panamá), also historically known as the Isthmus of Darien (), is the narrow strip of land that lies between the Caribbean Sea and the Pacific Ocean, linking North and South America. It contains the country ...
during the Pliocene, making possible the
Great American Interchange The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which lan ...
and bringing a nearly complete end to South America's distinctive native ungulate fauna, though other South American lineages like its predatory mammals were already extinct by this point and others like
xenarthra Xenarthra (; from Ancient Greek ξένος, xénos, "foreign, alien" + ἄρθρον, árthron, "joint") is a major clade of placental mammals native to the Americas. There are 31 living species: the anteaters, tree sloths, and armadillos. Ex ...
ns continued to do well afterwards. The formation of the Isthmus had major consequences on global temperatures, since warm equatorial ocean currents were cut off and an Atlantic cooling cycle began, with cold Arctic and Antarctic waters dropping temperatures in the now-isolated Atlantic Ocean.
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
's collision with
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
formed the
Mediterranean Sea The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the ea ...
, cutting off the remnants of the
Tethys Ocean The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...
. The border between the Miocene and the Pliocene is also the time of the
Messinian salinity crisis The Messinian salinity crisis (MSC), also referred to as the Messinian event, and in its latest stage as the Lago Mare event, was a geological event during which the Mediterranean Sea went into a cycle of partial or nearly complete desiccation (dr ...
. The
land bridge In biogeography, a land bridge is an isthmus or wider land connection between otherwise separate areas, over which animals and plants are able to cross and Colonisation (biology), colonize new lands. A land bridge can be created by marine regre ...
between
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
and
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
(
Beringia Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip ...
) was first flooded near the start of the Pliocene, allowing marine organisms to spread between the Arctic and Pacific Oceans. The bridge would continue to be periodically flooded and restored thereafter. Pliocene marine formations are exposed in northeast
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
, southern
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
,
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
, and
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
. During the Pliocene parts of southern Norway and southern Sweden that had been near sea level rose. In Norway this rise elevated the Hardangervidda plateau to 1200 m in the Early Pliocene. In Southern Sweden similar movements elevated the
South Swedish highlands 300px, Aerial view of farms and forest in Ydre Municipality. 250px, The forested landscape of the South Swedish highlands, seen from Eksjö_Municipality.html"_;"title="Skuruhatt_in_Eksjö_Municipality">Skuruhatt_in_Eksjö_Municipality. The_Sout ...
leading to a deflection of the ancient Eridanos river from its original path across south-central Sweden into a course south of Sweden.


Environment and evolution of human ancestors

The Pliocene is bookended by two significant events in the evolution of human ancestors. The first is the appearance of the
hominin The Hominini form a taxonomic tribe of the subfamily Homininae ("hominines"). Hominini includes the extant genera ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos) and in standard usage excludes the genus ''Gorilla'' (gorillas). The t ...
''
Australopithecus anamensis ''Australopithecus anamensis'' is a hominin species that lived approximately between 4.2 and 3.8 million years ago and is the oldest known ''Australopithecus'' species, living during the Plio-Pleistocene era. Nearly one hundred fossil specimens ...
'' in the early Pliocene, around 4.2 million years ago. The second is the appearance of ''
Homo ''Homo'' () is the genus that emerged in the (otherwise extinct) genus ''Australopithecus'' that encompasses the extant species ''Homo sapiens'' ( modern humans), plus several extinct species classified as either ancestral to or closely relate ...
'', the genus that includes
modern humans Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, an ...
and their closest extinct relatives, near the end of the Pliocene at 2.6 million years ago. Key traits that evolved among hominins during the Pliocene include terrestrial bipedality and, by the end of the Pliocene, encephalized brains (brains with a large
neocortex The neocortex, also called the neopallium, isocortex, or the six-layered cortex, is a set of layers of the mammalian cerebral cortex involved in higher-order brain functions such as sensory perception, cognition, generation of motor commands, sp ...
relative to body mass and stone tool manufacture. Improvements in
dating methods Chronological dating, or simply dating, is the process of attributing to an object or event a date in the past, allowing such object or event to be located in a previously established chronology. This usually requires what is commonly known as a "d ...
and in the use of climate proxies have provided scientists with the means to test hypotheses of the evolution of human ancestors. Early hypotheses of the evolution of human traits emphasized the selective pressures produced by particular habitats. For example, many scientists have long favored the
savannah hypothesis The savannah hypothesis (or savanna hypothesis) is a hypothesis that human bipedalism evolved as a direct result of human ancestors' transition from an arboreal lifestyle to one on the savannas. According to the hypothesis, millions of years ago h ...
. This proposes that the evolution of terrestrial bipedality and other traits was an adaptive response to Pliocene climate change that transformed forests into more open
savannah A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the Canopy (forest), canopy does not close. The open canopy allows sufficient light to rea ...
. This was championed by
Grafton Elliot Smith Sir Grafton Elliot Smith (15 August 1871 – 1 January 1937) was an Australian-British anatomist, Egyptologist and a proponent of the hyperdiffusionist view of prehistory. He believed in the idea that cultural innovations occur only once and ...
in his 1924 book, ''The Evolution of Man'', as "the unknown world beyond the trees", and was further elaborated by
Raymond Dart Raymond Arthur Dart (4 February 1893 – 22 November 1988) was an Australian anatomist and anthropologist, best known for his involvement in the 1924 discovery of the first fossil ever found of ''Australopithecus africanus'', an extinct homi ...
as the
killer ape theory The killer ape theory or killer ape hypothesis is the theory that war and interpersonal aggression was the driving force behind human evolution. It was originated by Raymond Dart in the 1950s; it was developed further in ''African Genesis'' by Rob ...
. Other scientists, such as
Sherwood L. Washburn Sherwood Larned Washburn ( – ), nicknamed "Sherry", was an American physical anthropologist, and "a legend in the field." He was pioneer in the field of primatology, opening it to the study of primates in their natural habitats. His research ...
, emphasized an intrinsic model of hominin evolution. According to this model, early evolutionary developments triggered later developments. The model placed little emphasis on the surrounding environment. Anthropologists tended to focus on intrinsic models while geologists and vertebrate paleontologists tended to put greater emphasis on habitats. Alternatives to the savanna hypothesis include the woodland/forest hypothesis, which emphasizes the evolution of hominins in closed habitats, or hypotheses emphasizing the influence of colder habitats at higher latitudes or the influence of seasonal variation. More recent research has emphasized the variability selection hypothesis, which proposes that variability in climate fostered development of hominin traits. Improved climate proxies show that the Pliocene climate of east Africa was highly variable, suggesting that adaptability to varying conditions was more important in driving hominin evolution than the steady pressure of a particular habitat.


Flora

The change to a cooler, drier, more seasonal climate had considerable impacts on Pliocene vegetation, reducing tropical species worldwide.
Deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
forests proliferated,
coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extant ...
forests and
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless moun ...
covered much of the north, and
grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
s spread on all continents (except Antarctica). Tropical forests were limited to a tight band around the equator, and in addition to dry savannahs,
deserts A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About one ...
appeared in Asia and Africa.


Fauna

Both marine and continental faunas were essentially modern, although continental faunas were a bit more primitive than today. The land mass collisions meant great migration and mixing of previously isolated species, such as in the
Great American Interchange The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which lan ...
.
Herbivores A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
got bigger, as did specialized predators. Image:Oliva sayana.jpg, The
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
''
Oliva sayana The lettered olive, ''Oliva sayana'', is a species of large predatory sea snail, a marine gastropod mollusc in the family Olividae, the olive shells, olive snails, or olives. MolluscaBase eds. (2021). MolluscaBase. Oliva sayana Ravenel, 1834. Acc ...
'', from the Pliocene of
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
Image:Cladocora.jpg, The
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and sec ...
''
Cladocora ''Cladocora'' is a genus of corals in the order of stony corals. Species Species in this genus include: * ''Cladocora arbuscula'' (Le Sueur, 1821) * ''Cladocora caespitosa'' (Linnaeus, 1767) * ''Cladocora debilis'' Milne Edwards & Haime, 1849 ...
'' from the Pliocene of
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is geo ...
Image:CyprusPlioceneGastropod.JPG, A gastropod and attached serpulid wormtube from the Pliocene of Cyprus Image:Turritellatricarinata.jpg, The gastropod ''
Turritella ''Turritella'' is a genus of medium-sized sea snails with an operculum, marine gastropod mollusks in the family Turritellidae.Vos, C.; Gofas, S. (2013). Turritella Lamarck, 1799. Accessed through: World Register of Marine Species at http://www. ...
carinata'' from the Pliocene of Cyprus Image:SpondylusPliocene.jpg, The thorny
oyster Oyster is the common name for a number of different families of salt-water bivalve molluscs that live in marine or brackish habitats. In some species, the valves are highly calcified, and many are somewhat irregular in shape. Many, but not al ...
''
Spondylus ''Spondylus'' is a genus of bivalve molluscs, the only genus in the family (biology), family Spondylidae.MolluscaBase (2019). MolluscaBase. Spondylus Linnaeus, 1758. Accessed through: World Register of Marine Species at: http://www.marinespecies ...
'' right and left valve interiors from the Pliocene of Cyprus Image:Diodoraitalica.jpg, The
limpet Limpets are a group of aquatic snails that exhibit a conical shell shape (patelliform) and a strong, muscular foot. Limpets are members of the class Gastropoda, but are polyphyletic, meaning the various groups called "limpets" descended indep ...
''Diodora italica'' from the Pliocene of Cyprus Image:DentaliumPliocene.jpg, The scaphopod '' Dentalium'' from the Pliocene of Cyprus File:Aporrhais from Pliocene.jpg, The gastropod ''
Aporrhais ''Aporrhais'' is a genus of medium-sized sea snails, marine gastropod mollusks in the family Aporrhaidae and the superfamily Stromboidea.Anadara ''Anadara'' is a genus of saltwater bivalves, ark clams, in the family Arcidae. It is also called ''Scapharca''. This genus is known in the fossil record from the Cretaceous period to the Quaternary period (age range: 140.2 to 0.0 million year ...
'' from the Pliocene of Cyprus Image:Amusium cristatum Cyprus.jpg, The pectenid bivalve ''Ammusium cristatum'' from the Pliocene of Cyprus Image:Petaloconchus Cyprus Pliocene.JPG, Vermetid gastropod ''
Petaloconchus intortus ''Petaloconchus intortus'' is an extinct species of sea snail, a marine gastropod mollusk in the family Vermetidae The Vermetidae, the worm snails or worm shells, are a taxonomic family of small to medium-sized sea snails, marine gastropod ...
'' attached to a branch of the
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and sec ...
''Cladocora'' from the Pliocene of Cyprus Image:Chesapecten barnacles Pliocene VA.jpg, ''
Chesapecten ''Chesapecten'' is an extinct genus of scallop known from marine strata from the early Miocene to the early Pleistocene of the Eastern United States. It flourished in the shallow seas along the Mid-Atlantic during this period. Other scallop ...
'',
barnacles A barnacle is a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in erosive ...
and sponge borings (''
Entobia ''Entobia'' is a trace fossil in a hard substrate (typically a shell, rock or hardground made of calcium carbonate) formed by sponges as a branching network of galleries, often with regular enlargements termed chambers. Apertural canals connec ...
'') from the Pliocene of York River, Virginia


Mammals

In North America,
rodents Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are nat ...
, large
mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of th ...
s and
gomphothere Gomphotheres are any members of the diverse, extinct taxonomic family Gomphotheriidae. Gomphotheres were elephant-like proboscideans, but do not belong to the family Elephantidae. They were widespread across Afro-Eurasia and North America during ...
s, and
opossum Opossums () are members of the marsupial order Didelphimorphia () endemic to the Americas. The largest order of marsupials in the Western Hemisphere, it comprises 93 species in 18 genera. Opossums originated in South America and entered North ...
s continued successfully, while hoofed animals (
ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...
s) declined, with
camel A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. C ...
,
deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
and
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million y ...
all seeing populations recede. Three-toed horses (''
Nannippus ''Nannippus'' is an extinct genus of three-toed horse endemic to North America during the Miocene through Pleistoceneabout 13.3—1.8 million years ago (Mya), living around 11.5 million years. This ancient species of three-toed horse grew up to 3 ...
''),
oreodont Merycoidodontoidea, sometimes called "oreodonts" or "ruminating hogs", is an extinct superfamily of prehistoric cud-chewing artiodactyls with short faces and fang-like canine teeth. As their name implies, some of the better known forms were gen ...
s,
protoceratid Protoceratidae is an extinct family of herbivorous North American artiodactyls (even-toed ungulates) that lived during the Eocene through Pliocene at around 46.2—4.9 Mya, existing for about 41 million years. Classification Protoceratidae was ...
s, and
chalicothere Chalicotheres (from Greek '' chalix'', "gravel" and '' therion'', "beast") are an extinct clade of herbivorous, odd-toed ungulate (perissodactyl) mammals that lived in North America, Eurasia, and Africa from the Middle Eocene until the Early Plei ...
s became extinct. Borophagine dogs and ''
Agriotherium ''Agriotherium'' is an extinct genus of bears whose fossils are found in Miocene through Pleistocene-aged strata of North America, Europe, Africa, and Asia. This long-lived genus persisted from at least ~11.6–2.5 Mya. Materials from the late ...
'' became extinct, but other
carnivores A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
including the
weasel Weasels are mammals of the genus ''Mustela'' of the family Mustelidae. The genus ''Mustela'' includes the least weasels, polecats, stoats, ferrets and European mink. Members of this genus are small, active predators, with long and slender bo ...
family diversified, and
dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the wolf. Also called the domestic dog, it is derived from the extinct Pleistocene wolf, and the modern wolf is the dog's nearest living relative. Do ...
s and
short-faced bear The Tremarctinae or short-faced bears is a subfamily of Ursidae that contains one living representative, the spectacled bear (''Tremarctos ornatus'') of South America, and several extinct species from four genera: the Florida spectacled bear ('' ...
s did well.
Ground sloth Ground sloths are a diverse group of extinct sloths in the mammalian superorder Xenarthra. The term is used to refer to all extinct sloths because of the large size of the earliest forms discovered, compared to existing tree sloths. The Caribbe ...
s, huge
glyptodont Glyptodonts are an extinct subfamily of large, heavily armoured armadillos. They arose in South America around 48 million years ago and spread to southern North America after the continents became connected several million years ago. The best-kn ...
s, and
armadillo Armadillos (meaning "little armored ones" in Spanish) are New World placental mammals in the order Cingulata. The Chlamyphoridae and Dasypodidae are the only surviving families in the order, which is part of the superorder Xenarthra, along wi ...
s came north with the formation of the Isthmus of Panama. In
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
rodents did well, while
primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
distribution declined.
Elephant Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae an ...
s,
gomphothere Gomphotheres are any members of the diverse, extinct taxonomic family Gomphotheriidae. Gomphotheres were elephant-like proboscideans, but do not belong to the family Elephantidae. They were widespread across Afro-Eurasia and North America during ...
s and
stegodon ''Stegodon'' ("roofed tooth" from the Ancient Greek words , , 'to cover', + , , 'tooth' because of the distinctive ridges on the animal's molars) is an extinct genus of proboscidean, related to elephants. It was originally assigned to the famil ...
ts were successful in Asia (the largest land mammals of the Pliocene were such proboscideans as ''
Deinotherium ''Deinotherium'' was a large elephant-like proboscidean that appeared in the Middle Miocene and survived until the Early Pleistocene. Although superficially resembling modern elephants, they had notably more flexible necks, limbs adapted to a mo ...
'', ''
Anancus ''Anancus'' is an extinct genus of elephantoid proboscideans ("gomphothere" ''sensu lato'') native to Afro-Eurasia, that lived from the Tortonian stage of the late Miocene until the genus' extinction during the early Pleistocene, roughly from 8.5 ...
'' and ''
Mammut borsoni ''Zygolophodon'' is an extinct genus of African, Asian, and European mammutid that lived from the Miocene to the Late Pliocene. Taxonomy ''Zygolophodon'' belongs in the family Mammutidae, whose best known member is the American mastodon. ''Zy ...
''), and
hyrax Hyraxes (), also called dassies, are small, thickset, herbivorous mammals in the order Hyracoidea. Hyraxes are well-furred, rotund animals with short tails. Typically, they measure between long and weigh between . They are superficially simil ...
es migrated north from Africa.
Horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million y ...
diversity declined, while tapirs and rhinos did fairly well.
Bovines Bovines ( subfamily Bovinae) comprise a diverse group of 10 genera of medium to large-sized ungulates, including cattle, bison, African buffalo, water buffalos, and the four-horned and spiral-horned antelopes. The evolutionary relationship betw ...
and
antelope The term antelope is used to refer to many species of even-toed ruminant that are indigenous to various regions in Africa and Eurasia. Antelope comprise a wastebasket taxon defined as any of numerous Old World grazing and browsing hoofed mammals ...
s were successful; some camel species crossed into Asia from North America.
Hyenas Hyenas, or hyaenas (from Ancient Greek , ), are feliform carnivoran mammals of the family Hyaenidae . With only four extant species (each in its own genus), it is the fifth-smallest family in the Carnivora and one of the smallest in the cla ...
and early
saber-toothed cat Machairodontinae is an extinct subfamily of carnivoran mammals of the family Felidae (true cats). They were found in Asia, Africa, North America, South America, and Europe from the Miocene to the Pleistocene, living from about 16 million until ...
s appeared, joining other predators including dogs, bears and weasels. Africa was dominated by hoofed animals, and primates continued their evolution, with
australopithecine Australopithecina or Hominina is a subtribe in the tribe Hominini. The members of the subtribe are generally ''Australopithecus'' (cladistically including the genus, genera ''Homo'', ''Paranthropus'', and ''Kenyanthropus''), and it typically in ...
s (some of the first
hominin The Hominini form a taxonomic tribe of the subfamily Homininae ("hominines"). Hominini includes the extant genera ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos) and in standard usage excludes the genus ''Gorilla'' (gorillas). The t ...
s) and baboon-like monkeys such as the ''
Dinopithecus ''Dinopithecus'' ("terrible ape") is an extinct genus of very large primate closely related to the baboon that lived during the Pliocene to the Pleistocene epoch of South Africa and Ethiopia. It was named by British paleontologist Robert Broom i ...
'' appearing in the late Pliocene. Rodents were successful, and elephant populations increased. Cows and antelopes continued diversification and overtook
pig The pig (''Sus domesticus''), often called swine, hog, or domestic pig when distinguishing from other members of the genus '' Sus'', is an omnivorous, domesticated, even-toed, hoofed mammal. It is variously considered a subspecies of ''Sus s ...
s in numbers of species. Early
giraffe The giraffe is a large African hoofed mammal belonging to the genus ''Giraffa''. It is the tallest living terrestrial animal and the largest ruminant on Earth. Traditionally, giraffes were thought to be one species, ''Giraffa camelopardalis ...
s appeared. Horses and modern rhinos came onto the scene. Bears, dogs and weasels (originally from North America) joined cats, hyenas and
civet A civet () is a small, lean, mostly nocturnal mammal native to tropical Asia and Africa, especially the tropical forests. The term civet applies to over a dozen different species, mostly from the family Viverridae. Most of the species diversity ...
s as the African predators, forcing hyenas to adapt as specialized scavengers. Most mustelids in Africa declined as a result of increased competition from the new predators, although '' Enhydriodon omoensis'' remained an unusually successful terrestrial predator. South America was invaded by North American species for the first time since the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
, with North American rodents and primates mixing with southern forms.
Litoptern Litopterna (from grc, λῑτή πτέρνα "smooth heel") is an extinct order of fossil hoofed mammals from the Cenozoic era. The order is one of the five great orders of South American ungulates that were endemic to the continent, until th ...
s and the
notoungulate Notoungulata is an extinct order of mammalian ungulates that inhabited South America from the early Paleocene to the Holocene, living from approximately 61 million to 11,000 years ago. Notoungulates were morphologically diverse, with forms resemb ...
s, South American natives, were mostly wiped out, except for the macrauchenids and
toxodonts Toxodontia. Retrieved April 2013. is a suborder of the meridiungulate order Notoungulata. Most of the members of the five included families, including the largest notoungulates, share several dental, auditory and tarsal specializations. The gr ...
, which managed to survive. Small weasel-like carnivorous
mustelid The Mustelidae (; from Latin ''mustela'', weasel) are a family of carnivorous mammals, including weasels, badgers, otters, ferrets, martens, minks and wolverines, among others. Mustelids () are a diverse group and form the largest family in ...
s,
coati Coatis, also known as coatimundis (), are members of the family Procyonidae in the genera ''Nasua'' and ''Nasuella''. They are diurnal mammals native to South America, Central America, Mexico, and the southwestern United States. The name "c ...
s and short-faced bears migrated from the north. Grazing
glyptodont Glyptodonts are an extinct subfamily of large, heavily armoured armadillos. They arose in South America around 48 million years ago and spread to southern North America after the continents became connected several million years ago. The best-kn ...
s, browsing
giant ground sloth ''Megatherium'' ( ; from Greek () 'great' + () 'beast') is an extinct genus of ground sloths endemic to South America that lived from the Early Pliocene through the end of the Pleistocene. It is best known for the elephant-sized type species ' ...
s and smaller caviomorph rodents,
pampathere Pampatheriidae ("Pampas beasts") is an extinct family of large plantigrade armored armadillos related to extant armadillos in the order Cingulata. However, pampatheriids have existed as a separate lineage since at least the middle Eocene Mus ...
s, and
armadillos Armadillos (meaning "little armored ones" in Spanish) are New World placental mammals in the order Cingulata. The Chlamyphoridae and Dasypodidae are the only surviving families in the order, which is part of the superorder Xenarthra, along w ...
did the opposite, migrating to the north and thriving there. The marsupials remained the dominant Australian mammals, with herbivore forms including
wombat Wombats are short-legged, muscular quadrupedal marsupials that are native to Australia. They are about in length with small, stubby tails and weigh between . All three of the extant species are members of the family Vombatidae. They are adap ...
s and
kangaroo Kangaroos are four marsupials from the family Macropodidae (macropods, meaning "large foot"). In common use the term is used to describe the largest species from this family, the red kangaroo, as well as the antilopine kangaroo, eastern gre ...
s, and the huge ''
Diprotodon ''Diprotodon'' (Ancient Greek: "two protruding front teeth") is an extinct genus of marsupial from the Pleistocene of Australia, containing one species, ''D. optatum''. The earliest finds date to 1.77 million to 780,000 years ago, but most speci ...
''. Carnivorous marsupials continued hunting in the Pliocene, including
dasyurid The Dasyuridae are a family of marsupials native to Australia and New Guinea, including 71 extant species divided into 17 genera. Many are small and mouse-like or shrew-like, giving some of them the name marsupial mice or marsupial shrews, but th ...
s, the dog-like
thylacine The thylacine ( , or , also ) (''Thylacinus cynocephalus'') is an extinct carnivorous marsupial that was native to the Australian mainland and the islands of Tasmania and New Guinea. The last known live animal was captured in 1930 in Tasman ...
and cat-like ''
Thylacoleo ''Thylacoleo'' ("pouch lion") is an extinct genus of carnivorous marsupials that lived in Australia from the late Pliocene to the late Pleistocene (2 million to 46 thousand years ago). Some of these marsupial lions were the largest mammalian pred ...
''. The first rodents arrived in Australia. The modern
platypus The platypus (''Ornithorhynchus anatinus''), sometimes referred to as the duck-billed platypus, is a semiaquatic, egg-laying mammal Endemic (ecology), endemic to Eastern states of Australia, eastern Australia, including Tasmania. The platypu ...
, a
monotreme Monotremes () are prototherian mammals of the order Monotremata. They are one of the three groups of living mammals, along with placentals (Eutheria), and marsupials (Metatheria). Monotremes are typified by structural differences in their brain ...
, appeared.


Birds

The predatory South American
phorusrhacids Phorusrhacids, colloquially known as terror birds, are an extinct clade of large carnivorous flightless birds that were one of the largest species of apex predators in South America during the Cenozoic era; their conventionally accepted temporal ...
were rare in this time; among the last was ''
Titanis ''Titanis'' was an extinct genus of giant flightless terror birds that inhabited North America during the early Pliocene to early Pleistocene epochs. The generic name, ''Titanis'', refers to the titans, Ancient Greek gods that preceded the Twelv ...
'', a large phorusrhacid that migrated to North America and rivaled mammals as top predator. Other birds probably evolved at this time, some modern (such as the genera '' Cygnus'', ''
Bubo A bubo (Greek βουβών, ''boubṓn'', 'groin') is adenitis or inflammation of the lymph nodes and is an example of reactive lymphadenopathy. Classification Buboes are a symptom of bubonic plague and occur as painful swellings in the thigh ...
'', ''
Struthio Ostriches are large flightless birds of the genus ''Struthio'' in the order Struthioniformes, part of the infra-class Palaeognathae, a diverse group of flightless birds also known as ratites that includes the emus, rheas, and kiwis. There are ...
'' and ''
Corvus ''Corvus'' is a widely distributed genus of medium-sized to large birds in the family Corvidae. It includes species commonly known as crows, ravens and Rook (bird), rooks. The species commonly encountered in Europe are the carrion crow, the hoo ...
''), some now extinct.


Reptiles and amphibians

Alligator An alligator is a large reptile in the Crocodilia order in the genus ''Alligator'' of the family Alligatoridae. The two extant species are the American alligator (''A. mississippiensis'') and the Chinese alligator (''A. sinensis''). Additiona ...
s and
crocodile Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term crocodile is sometimes used even more loosely to inclu ...
s died out in Europe as the climate cooled.
Venomous snake Venomous snakes are Species (biology), species of the Suborder (biology), suborder Snake, Serpentes that are capable of producing Snake venom, venom, which they use for killing prey, for defense, and to assist with digestion of their prey. The v ...
genera continued to increase as more rodents and birds evolved.
Rattlesnake Rattlesnakes are venomous snakes that form the genera ''Crotalus'' and ''Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting small anim ...
s first appeared in the Pliocene. The modern species ''
Alligator mississippiensis The American alligator (''Alligator mississippiensis''), sometimes referred to colloquially as a gator or common alligator, is a large crocodilian reptile native to the Southeastern United States. It is one of the two extant species in the gen ...
'', having evolved in the Miocene, continued into the Pliocene, except with a more northern range; specimens have been found in very late Miocene deposits of
Tennessee Tennessee ( , ), officially the State of Tennessee, is a landlocked state in the Southeastern region of the United States. Tennessee is the 36th-largest by area and the 15th-most populous of the 50 states. It is bordered by Kentucky to th ...
.
Giant tortoise Giant tortoises are any of several species of various large land tortoises, which include a number of extinct species, as well as two extant species with multiple subspecies formerly common on the islands of the western Indian Ocean and on the ...
s still thrived in North America, with genera like ''
Hesperotestudo ''Hesperotestudo'' ("Western turtle") is an extinct genus of tortoise native to North and Central America from the Oligocene to the Late Pleistocene. Species of ''Hesperotesudo'' varied widely in size, with the largest unnamed species from El Sal ...
''. Madtsoid
snake Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Ma ...
s were still present in Australia. The amphibian order
Allocaudata The Albanerpetontidae are an extinct family of small amphibians, native to the Northern Hemisphere during the Mesozoic and Cenozoic. The only members of the order Allocaudata, they are thought to be allied with living amphibians belonging to Lissa ...
became extinct.


Oceans

Oceans continued to be relatively warm during the Pliocene, though they continued cooling. The Arctic ice cap formed, drying the climate and increasing cool shallow currents in the North Atlantic. Deep cold currents flowed from the Antarctic. The formation of the Isthmus of Panama about 3.5 million years ago cut off the final remnant of what was once essentially a circum-equatorial current that had existed since the Cretaceous and the early
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
. This may have contributed to further cooling of the oceans worldwide. The Pliocene seas were alive with
sea cow The Sirenia (), commonly referred to as sea-cows or sirenians, are an order of fully aquatic, herbivorous mammals that inhabit swamps, rivers, estuaries, marine wetlands, and coastal marine waters. The Sirenia currently comprise two distinct f ...
s,
seal Seal may refer to any of the following: Common uses * Pinniped, a diverse group of semi-aquatic marine mammals, many of which are commonly called seals, particularly: ** Earless seal, or "true seal" ** Fur seal * Seal (emblem), a device to impr ...
s,
sea lion Sea lions are pinnipeds characterized by external ear flaps, long foreflippers, the ability to walk on all fours, short and thick hair, and a big chest and belly. Together with the fur seals, they make up the family Otariidae, eared seals. ...
s and
shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimo ...
s.


Supernovae

In 2002, Narciso Benítez ''et al.'' calculated that roughly 2 million years ago, around the end of the Pliocene Epoch, a group of bright O and B stars called the Scorpius–Centaurus
OB association In astronomy, stellar kinematics is the observational study or measurement of the kinematics or motions of stars through space. Stellar kinematics encompasses the measurement of stellar velocities in the Milky Way and its satellites as well as t ...
passed within 130 light-years of Earth and that one or more
supernova A supernova is a powerful and luminous explosion of a star. It has the plural form supernovae or supernovas, and is abbreviated SN or SNe. This transient astronomical event occurs during the last evolutionary stages of a massive star or when ...
explosions gave rise to a feature known as the
Local Bubble The Local Bubble, or Local Cavity, is a relative cavity in the interstellar medium (ISM) of the Orion Arm in the Milky Way. It contains the closest of celestial neighbours and among others, the Local Interstellar Cloud (which contains the Sola ...
. Such a close explosion could have damaged the Earth's
ozone layer The ozone layer or ozone shield is a region of Earth's stratosphere that absorbs most of the Sun's ultraviolet radiation. It contains a high concentration of ozone (O3) in relation to other parts of the atmosphere, although still small in rela ...
and caused the extinction of some ocean life (at its peak, a supernova of this size could have the same
absolute magnitude Absolute magnitude () is a measure of the luminosity of a celestial object on an inverse Logarithmic scale, logarithmic Magnitude (astronomy), astronomical magnitude scale. An object's absolute magnitude is defined to be equal to the apparent mag ...
as an entire galaxy of 200 billion stars).Comins & Kaufmann (2005), p. 359. Radioactive
iron-60 Naturally occurring iron (26Fe) consists of four stable isotopes: 5.845% of 54Fe (possibly radioactive with a half-life over years), 91.754% of 56Fe, 2.119% of 57Fe and 0.286% of 58Fe. There are 24 known radioactive isotopes, the most stable of w ...
isotopes that have been found in ancient seabed deposits further back this finding, as there are no natural sources for this radioactive isotope on Earth, but they can be produced in supernovae. Furthermore, iron-60 residues point to a huge spike 2.6 million years ago, but an excess scattered over 10 million years can also be found, suggesting that there may have been multiple, relatively close supernovae. In 2019, researchers found more of these interstellar iron-60 isotopes in Antarctica, which have been associated with the
Local Interstellar Cloud The Local Interstellar Cloud (LIC), also known as the Local Fluff, is an interstellar cloud roughly across, through which the Solar System is moving. This feature overlaps a region around the Sun referred to as the solar neighborhood. It is unk ...
.


See also

*
List of fossil sites This list of fossil sites is a worldwide list of localities known well for the presence of fossils. Some entries in this list are notable for a single, unique find, while others are notable for the large number of fossils found there. Many of t ...
''(with link directory)''


Notes


References


Further reading

* * ; 2004: ''A Geologic Time Scale 2004'',
Cambridge University , mottoeng = Literal: From here, light and sacred draughts. Non literal: From this place, we gain enlightenment and precious knowledge. , established = , other_name = The Chancellor, Masters and Schola ...
Press. * *


External links


Mid-Pliocene Global Warming: NASA/GISS Climate Modeling







"Supernova dealt deaths on Earth? Stellar blasts may have killed ancient marine life" ''Science News Online''
retrieved February 2, 2002


Pliocene Microfossils: 100+ images of Pliocene Foraminifera

Human Timeline (Interactive)
Smithsonian,
National Museum of Natural History The National Museum of Natural History is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. In 2021, with 7 ...
(August 2016). {{Authority control Geological epochs Neogene geochronology