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Sparassodonta
Sparassodonta (from Ancient Greek, Greek ['], to tear, rend; and , gen. [', '], tooth) is an extinct order (biology), order of carnivore, carnivorous metatherian mammals native to South America, related to modern marsupials. They were once considered to be true marsupials, but are now thought to be a separate side branch that split before the last common ancestor of all modern marsupials. A number of these mammalian predators closely resemble placental predators that evolved separately on other continents, and are cited frequently as examples of convergent evolution. They were first described by Florentino Ameghino, from fossils found in the Santa Cruz beds of Patagonia. Sparassodonts were present throughout South America's long period of "splendid isolation" during the Cenozoic; during this time, they shared the niches for large warm-blooded predators with the flightless Phorusrhacidae, terror birds. Previously, it was thought that these mammals died out in the face of competit ...
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Proborhyaenidae
Proborhyaenidae is an extinct family of metatherian mammals of the order Sparassodonta, which lived in South America from the Eocene (Mustersan) until the Oligocene (Deseadan). Sometimes it has been included as a subfamily of their relatives, the borhyaenids (as Proborhyaeninae). The largest species, ''Proborhyaena gigantea'', is estimated to be about the size of a spectacled bear, with its skull reaching in length, and body mass estimates up to approximately , making the proborhyaenids some of the largest known metatherians. Proborhyaenid remains have been found in western Bolivia, Uruguay, southern Brazil, and the provinces of Mendoza, Salta, and Chubut, in Argentina. Most proborhyaenids had a robust, hyena-like skull, although one species, ''Callistoe vincei'', had an elongate, narrow skull more reminiscent of a thylacine. The teeth were strongly specialized as carnassials for eating meat, and in '' Arminiheringia'' rotated throughout the animal's life to maintain a continu ...
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Great American Interchange
The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which land and freshwater fauna migrated from North America via Central America to South America and vice versa, as the volcanic Isthmus of Panama rose up from the sea floor and bridged the formerly separated continents. Although earlier dispersals had occurred, probably over water, the migration accelerated dramatically about 2.7 million years ( Ma) ago during the Piacenzian age. It resulted in the joining of the Neotropic (roughly South American) and Nearctic (roughly North American) biogeographic realms definitively to form the Americas. The interchange is visible from observation of both biostratigraphy and nature (neontology). Its most dramatic effect is on the zoogeography of mammals, but it also gave an opportunity for reptiles, amphibia ...
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Borhyaenidae
Borhyaenidae is an extinct metatherian family of low-slung, heavily built predatory mammals in the order Sparassodonta. Borhyaenids are not true marsupials, but members of a sister taxon, Sparassodonta. Like most metatherians, borhyaenids and other sparassodonts are thought to have had a pouch to carry their offspring around. Borhyaenids had strong and powerful jaws, like those of the unrelated placentalians ''Hyaenodon'' and ''Andrewsarchus'', for crushing bones. Borhyaenids grew up to an average of long. ''Borhyaena'' is the type genus of this group. Originally, the borhyaenids were one of the most diverse groups of sparassodonts, including all species not originally included in the Thylacosmilidae. However, in recent years, with the elevation of most sparassodont subfamilies to family rank and the discovery that borhyaenids are more closely related to proborhyaenids and thylacosmilids than other sparassodonts, the family has been reduced to seven species in four genera. Th ...
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Metatheria
Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well as many extinct non-marsupial relatives. There are three extant subclasses of mammals, one being metatherians: #monotremes: egg laying mammals like the platypus and the echidna, #metatheria: marsupials, which includes three American orders ( Didelphimorphia, Paucituberculata and Microbiotheria) and four Australasian orders (Notoryctemorphia, Dasyuromorphia, Peramelemorphia and Diprotodontia), and the # eutherians: placental mammals, consisting of four superorders divided into 21 orders. Metatherians belong to a subgroup of the northern tribosphenic mammal clade or Boreosphenida. They differ from all other mammals in certain morphologies like their dental formula, which includes about five upper and four lower incisors, a canine, three pre ...
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Thylacosmilidae
Thylacosmilidae is an extinct Family (biology), family of metatherian predators, related to the modern marsupials, which lived in South America between the Miocene and Pliocene epochs. Like other South American mammalian predators that lived prior to the Great American Biotic Interchange, these animals belonged to the order Sparassodonta, which occupied the ecological niche of many eutherian mammals of the order Carnivora from other continents. The family's most notable feature are the elongated, laterally flattened fangs, which is a remarkable evolutionary convergence with other saber-toothed mammals like ''Barbourofelis'' and ''Smilodon''. Taxonomic history The family Thylacosmilidae was originally erected by Riggs in 1933, to accommodate ''Thylacosmilus'', found in the Pliocene Brochero Formation of Argentina. Later, the family was demoted to a subfamily, as Thylacosmilinae, within Borhyaenidae, a group of superficially canid-like sparassodonts, under the assumption that '' ...
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Lycopsis
''Lycopsis'' is an extinct genus of South American metatherian, that lived during the Miocene in Argentina and Colombia. History Although not named until 1927, Florentino Ameghino described a species now seen as synonymous with ''Lycopsis torresi'', ''Anatherium oxyrhynchus'', in 1895 based on a mandibular ramus with several teeth.Marshall, L. G. (1977). A new species of Lycopsis (Borhyaenidae: Marsupialia) from the La Venta fauna (Late Miocene) of Colombia, South America. ''Journal of Paleontology'', 633-642. The fossil was recovered from Puesto Estancia La Costa in Santa Cruz, Argentina, dating to the Miocene. The type material of ''Lycopsis'' was collected in July 1895 by "C. Berry" from the middle Miocene strata of the Santa Cruz Formation along the Santa Cruz River in the same area.Cabrera, Á. (1927). Datos para el conocimiento de los dasiuroideos fósiles argentinos. ''Revista del Museo de la Plata'', ''30'', 271-315. The fossils (MLP 11-113) were fragmentary, constitu ...
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Pliocene
The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58See the 2014 version of the ICS geologic time scale
million years ago. It is the second and most recent epoch of the Neogene Period in the . The Pliocene follows the Epoch and is followed by the Epoch. Prior to the 2009 ...
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Carnivore
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other soft tissues) whether through hunting or scavenging. Nomenclature Mammal order The technical term for mammals in the order Carnivora is ''carnivoran'', and they are so-named because most member species in the group have a carnivorous diet, but the similarity of the name of the order and the name of the diet causes confusion. Many but not all carnivorans are meat eaters; a few, such as the large and small cats (felidae) are ''obligate'' carnivores (see below). Other classes of carnivore are highly variable. The Ursids, for example: While the Arctic polar bear eats meat almost exclusively (more than 90% of its diet is meat), almost all other bear species are omnivorous, and one species, the giant panda, is nearly exclusively herbivorous. ...
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Phorusrhacidae
Phorusrhacids, colloquially known as terror birds, are an extinct clade of large carnivorous flightless birds that were one of the largest species of apex predators in South America during the Cenozoic era; their conventionally accepted temporal range covers from 62 to 0.1 million years ( Ma) ago. They ranged in height from . Their closest modern-day relatives are believed to be the seriemas. ''Titanis walleri'', one of the larger species, is known from Texas and Florida in North America. This makes the phorusrhacids the only known large South American predator to migrate north in the Great American Interchange that followed the formation of the Isthmus of Panama land bridge (the main pulse of the interchange began about 2.6 Ma ago; ''Titanis'' at 5 Ma was an early northward migrant). It was once believed that ''T. walleri'' became extinct in North America around the time of the arrival of humans, but subsequent datings of ''Titanis'' fossils provided no evidence for their surv ...
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Cenozoic
The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configuration of continents. It is the latest of three geological eras since complex life evolved, preceded by the Mesozoic and Paleozoic. It started with the Cretaceous–Paleogene extinction event, when many species, including the non-avian dinosaurs, became extinct in an event attributed by most experts to the impact of a large asteroid or other celestial body, the Chicxulub impactor. The Cenozoic is also known as the Age of Mammals because the terrestrial animals that dominated both hemispheres were mammalsthe eutherians (placentals) in the northern hemisphere and the metatherians (marsupials, now mainly restricted to Australia) in the southern hemisphere. The extinction of many groups allowed mammals and birds to greatly diversify so that l ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Journal Of Mammalian Evolution
The ''Journal of Mammalian Evolution'' is the official journal of the Society for the Study of Mammalian Evolution. The journal is peer-reviewed Bi-monthly is a multidisciplinary forum devoted to studies on the comparative morphology, molecular biology, paleobiology, genetics, developmental and reproductive biology, biogeography, systematics, ethology and ecology, and population dynamics of mammals and the ways that these diverse data can be analyzed for the reconstruction of mammalian evolution. According to the ''Journal Citation Reports'', the journal has a 2014 impact factor The impact factor (IF) or journal impact factor (JIF) of an academic journal is a scientometric index calculated by Clarivate that reflects the yearly mean number of citations of articles published in the last two years in a given journal, as i ... of 2.357. References {{Reflist English-language journals ...
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