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Thylacosmilidae
Thylacosmilidae is an extinct Family (biology), family of metatherian predators, related to the modern marsupials, which lived in South America between the Miocene and Pliocene epochs. Like other South American mammalian predators that lived prior to the Great American Biotic Interchange, these animals belonged to the order Sparassodonta, which occupied the ecological niche of many eutherian mammals of the order Carnivora from other continents. The family's most notable feature are the elongated, laterally flattened fangs, which is a remarkable evolutionary convergence with other saber-toothed mammals like ''Barbourofelis'' and ''Smilodon''. Taxonomic history The family Thylacosmilidae was originally erected by Riggs in 1933, to accommodate ''Thylacosmilus'', found in the Pliocene Brochero Formation of Argentina. Later, the family was demoted to a subfamily, as Thylacosmilinae, within Borhyaenidae, a group of superficially canid-like sparassodonts, under the assumption that '' ...
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Sparassodonta
Sparassodonta (from Ancient Greek, Greek ['], to tear, rend; and , gen. [', '], tooth) is an extinct order (biology), order of carnivore, carnivorous metatherian mammals native to South America, related to modern marsupials. They were once considered to be true marsupials, but are now thought to be a separate side branch that split before the last common ancestor of all modern marsupials. A number of these mammalian predators closely resemble placental predators that evolved separately on other continents, and are cited frequently as examples of convergent evolution. They were first described by Florentino Ameghino, from fossils found in the Santa Cruz beds of Patagonia. Sparassodonts were present throughout South America's long period of "splendid isolation" during the Cenozoic; during this time, they shared the niches for large warm-blooded predators with the flightless Phorusrhacidae, terror birds. Previously, it was thought that these mammals died out in the face of competit ...
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Thylacosmilus
''Thylacosmilus'' is an extinct genus of Saber-toothed predator, saber-toothed metatherian mammals that inhabited South America from the Miocene, Late Miocene to Pliocene Epoch (geology), epochs. Though ''Thylacosmilus'' looks similar to the "Machairodontinae, saber-toothed cats", it was not a felid, like the well-known North American ''Smilodon'', but a Sparassodonta, sparassodont, a group closely related to marsupials, and only superficially resembled other saber-toothed mammals due to convergent evolution. A 2005 study found that the bite forces of ''Thylacosmilus'' and ''Smilodon'' were low, which indicates the killing-techniques of saber-toothed animals differed from those of extant species. Remains of ''Thylacosmilus'' have been found primarily in Catamarca Province, Catamarca, Entre Ríos Province, Entre Ríos, and La Pampa Provinces in northern Argentina. Taxonomy In 1926, the Marshall Field Paleontological Expeditions collected mammal fossils from the Ituzaingó Format ...
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Borhyaenidae
Borhyaenidae is an extinct metatherian family of low-slung, heavily built predatory mammals in the order Sparassodonta. Borhyaenids are not true marsupials, but members of a sister taxon, Sparassodonta. Like most metatherians, borhyaenids and other sparassodonts are thought to have had a pouch to carry their offspring around. Borhyaenids had strong and powerful jaws, like those of the unrelated placentalians ''Hyaenodon'' and ''Andrewsarchus'', for crushing bones. Borhyaenids grew up to an average of long. ''Borhyaena'' is the type genus of this group. Originally, the borhyaenids were one of the most diverse groups of sparassodonts, including all species not originally included in the Thylacosmilidae. However, in recent years, with the elevation of most sparassodont subfamilies to family rank and the discovery that borhyaenids are more closely related to proborhyaenids and thylacosmilids than other sparassodonts, the family has been reduced to seven species in four genera. Th ...
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Colhuehuapian
The Colhuehuapian age is a period of geologic time (21.0–17.5 Ma) within the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Deseadan and precedes the Santacrucian age. Etymology The age is named after the Colhué Huapí Member of the Sarmiento Formation in the Golfo San Jorge Basin, Patagonia, Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th .... Formations Fossils References Bibliography ;Colhué Huapí Member * * ;Abanico Formation * * * * * * ;Biblián Formation * ;Castillo Formation * * * * * ;Cerro Bandera Formation * * ;Chichinales Formation * * ;Chilcatay Formation * * * * * * * * * * ;Cura-Mallín Group * * * ;Gaiman Formation * ...
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Anachlysictis
''Anachlysictis gracilis'' is an extinct carnivorous mammal belonging to the group Sparassodonta, which were metatherians (a group including marsupials and their close relatives) that inhabited South America during the Cenozoic. ''Anachlysictis'' is the first record of such borhyaenoids in northern South America, and also the most primitive known member of the family Thylacosmilidae, a group of predators equipped with "saber teeth". It was also the only confirmed record of a thylacosmilid that did not belong to the genus ''Thylacosmilus'' until the official publication of ''Patagosmilus'' in 2010. This species was found in the Villavieja Formation in the area of La Venta in Colombia, a famous fossil deposit in the Middle Miocene (Laventan; 13.8-11.8 million years ago),''Anachlysictis gracilis' ...
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Patagosmilus
''Patagosmilus'' ("Patagonian knife" in Greek) is an extinct genus of meat-eating metatherian mammal of the family Thylacosmilidae, that lived in the Middle Miocene in South America.''Patagosmilus''
at .org
Like other representatives of this family, such as '' Thylacosmilus atrox'' and '' Anachlysictis gracilis'', it was characterized by its elongated s of the upper jaw, similar to th ...
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La Venta (Colombia)
La Venta is a fossil locality located in the modern departments of Tolima and Huila in Colombia. This site is one of the richest Neogene fossil assemblages in South America and represents the best-known Cenozoic fossil site outside of Argentina. It provides a glimpse of what life in the region was like before the main wave of the Great American Interchange. Geology The fossils occur in Middle Miocene rocks of the Honda Group, which is divided into the younger Villavieja Formation and the older La Victoria Formation. The La Venta fauna contained ancient species of animal genera and families still alive today, as well as some entirely extinct prehistoric lineages. These animals lived some 13.8 to 12 million years ago in the Laventan age, which was named after La Venta. At that time, the climate of the region was wetter than today and there was much forest of trees similar (and probably related) to the ''sapino'' (''Goupia glabra'') of our time. List of fauna Fossil animals f ...
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Eomakhaira
''Eomakhaira'' is an extinct genus of thylacosmilid sparassodont known from the Oligocene Abanico Formation of Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a .... It contains a single species, ''Eomakhaira molossus.'' References Sparassodonts Prehistoric mammal genera {{paleo-mammal-stub ...
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Smilodon
''Smilodon'' is a genus of the extinct machairodont subfamily of the felids. It is one of the most famous prehistoric mammals and the best known saber-toothed cat. Although commonly known as the saber-toothed tiger, it was not closely related to the tiger or other modern cats. ''Smilodon'' lived in the Americas during the Pleistocene epoch (2.5 Year#mya, mya – 10,000 years ago). The genus was named in 1842 based on fossils from Brazil; the generic name means "scalpel" or "two-edged knife" combined with "tooth". Three species are recognized today: ''S. gracilis'', ''S. fatalis'', and ''S. populator''. The two latter species were probably descended from ''S. gracilis'', which itself probably evolved from ''Megantereon''. The hundreds of individuals obtained from the La Brea Tar Pits in Los Angeles constitute the largest collection of ''Smilodon'' fossils. Overall, ''Smilodon'' was more robustly built than any Neontology, extant cat, with particularly well-d ...
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Great American Biotic Interchange
The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which land and freshwater fauna migrated from North America via Central America to South America and vice versa, as the volcanic Isthmus of Panama rose up from the sea floor and bridged the formerly separated continents. Although earlier dispersals had occurred, probably over water, the migration accelerated dramatically about 2.7 million years ( Ma) ago during the Piacenzian age. It resulted in the joining of the Neotropic (roughly South American) and Nearctic (roughly North American) biogeographic realms definitively to form the Americas. The interchange is visible from observation of both biostratigraphy and nature (neontology). Its most dramatic effect is on the zoogeography of mammals, but it also gave an opportunity for reptiles, amphibians, a ...
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Laventan
The Laventan ( es, Laventense) age is a period of geologic time (13.8 to 11.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes the Mayoan age.Madden et al., 1997 Etymology The age is named after the Miocene Lagerstätte La Venta, where a rich biodiversity from the Middle Miocene has been recovered from the Honda Group. Formations Fossil content Correlations The Laventan (13.8 to 11.8 Ma) correlates with: * NALMA ** latest Barstovian (15.97-13.65 Ma)Barstovian
at Fossilworks.org
** early

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Honda Group, Colombia
The Honda Group ( es, Grupo Honda, Tsh, Ngh) is a geological group of the Upper and Middle Magdalena Basins and the adjacent Central and Eastern Ranges of the Colombian Andes. The group, in older literature also defined as formation, is in its present-day type section in the Tatacoa Desert in the department of Huila subdivided into two main formations; La Victoria and Villavieja. The group was originally defined in and named after Honda, Tolima, but has been redefined based on the many fossil finds in the Tatacoa Desert, to the south. In the original type section of its occurrence, the thick group is subdivided into three formations, from old to young; Cambrás, San Antonio and Los Limones. The group dates to the Neogene period; in its broadest definition from the Late Oligocene to Late Miocene, and in the redefined type section restricted to the Laventan age of the South American Land Mammal Ages (SALMA), equivalent to the Middle Miocene Serravallian epoch. The Honda Gr ...
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