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An oak is a tree or
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
in the genus ''Quercus'' (; Latin "oak tree") of the
beech Beech (''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to temperate Europe, Asia, and North America. Recent classifications recognize 10 to 13 species in two distinct subgenera, ''Engleriana'' and ''Fagus''. The ''Engle ...
family,
Fagaceae The Fagaceae are a family of flowering plants that includes beeches, chestnuts and oaks, and comprises eight genera with about 927 species. Fagaceae in temperate regions are mostly deciduous, whereas in the tropics, many species occur as evergre ...
. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''
Lithocarpus ''Lithocarpus'' is a genus in the beech family, Fagaceae. Trees in this genus are commonly known as the stone oaks and differ from ''Quercus'' primarily because they produce insect-pollinated flowers on erect spikes and the female flowers have ...
'' (stone oaks), as well as in those of unrelated species such as '' Grevillea robusta'' (silky oaks) and the Casuarinaceae (she-oaks). The genus ''Quercus'' is native to the Northern Hemisphere, and includes deciduous and evergreen species extending from cool temperate to tropical latitudes in the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...
, Asia, Europe, and North Africa. North America has the largest number of oak species, with approximately 160 species in Mexico of which 109 are endemic and about 90 in the United States. The second greatest area of oak diversity is China, with approximately 100 species.


Description

Oaks have
spiral In mathematics, a spiral is a curve which emanates from a point, moving farther away as it revolves around the point. Helices Two major definitions of "spiral" in the American Heritage Dictionary are:leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
, with lobate margins in many species; some have
serrated leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
or entire leaves with smooth margins. Many deciduous species are
marcescent Marcescence is the withering and persistence of plant organs that normally are shed, and is a term most commonly applied to plant leaves. The underlying physiological mechanism is that trees transfer water and sap from the roots to the leaves t ...
, not dropping dead leaves until spring. In spring, a single oak tree produces both male flowers (in the form of
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in ''Salix''). They contain many, usually unisexual flowers, arranged cl ...
s) and small female flowers, meaning that the trees are
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is conne ...
. The fruit is a nut called an
acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera ''Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and borne ...
or oak nut borne in a cup-like structure known as a cupule; each acorn contains one seed (rarely two or three) and takes 6–18 months to mature, depending on their species. The acorns and leaves contain tannic acid, which helps to guard from fungi and insects. The live oaks are distinguished for being evergreen, but are not actually a distinct group and instead are dispersed across the genus.


Taxonomy

Linnaeus described only five species of oak from eastern
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, based on general leaf form. These were white oak ('' Quercus alba''), chestnut oak ('' Q. montana''), red oak (''
Q. rubra ''Quercus rubra'', the northern red oak, is an oak tree in the red oak group (''Quercus'' section ''Lobatae''). It is a native of North America, in the eastern and central United States and southeast and south-central Canada. It has been introdu ...
''), willow oak ('' Q. phellos''), and water oak ('' Q. nigra''). Because he was dealing with confusing leaf forms, the ''Q. montana'' and ''Q. rubra'' specimens actually included mixed foliage of more than one species.


Classification

The most recent classification of ''Quercus'' divides the genus into two
subgenera In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
and eight sections. These divisions support the evolutionary diversification of oaks among two distinct clades: the "
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
" clade, including oaks that diversified mainly in Eurasia; and the " New World" clade, for oaks that diversified mainly in the Americas.


Subgenus ''Quercus''

* Sect. ''Quercus'' (synonyms ''Lepidobalanus'' and ''Leucobalanus''), the
white oaks The genus ''Quercus'' contains about 500 species, some of which are listed here. The genus, as is the case with many large genera, is divided into subgenera and sections. Traditionally, the genus ''Quercus'' was divided into the two subgenera ' ...
of Europe, Asia and North America. Trees or shrubs that produce nuts, specifically acorns, as fruits. Acorns mature in one year for annual trees and two years for biannual trees. Acorn is encapsulated by a spiny cupule as characterized by the family Fagaceae. Flowers in the Quercus genera produce one flower per node, with three or six styles, as well as three or six ovaries, respectively. The leaves mostly lack a bristle on their lobe tips, which are usually rounded. The type species is '' Quercus robur''. * Sect. ''Protobalanus'', the canyon live oak and its relatives, in the southwestern United States and northwest Mexico. Styles are short; the acorns mature in 18 months and taste very bitter. The inside of the acorn shell appears woolly. The leaves typically have sharp lobe tips, with bristles at the lobe tip. * Sect. ''Ponticae,'' a disjunct including just two species. Styles are short and the acorns mature in 12 months. The leaves have large stipules, high secondary venation, and are highly toothed. * Sect. ''Virentes,'' the southern live oaks of the Americas. Styles are short and the acorns mature in 12 months. The leaves are evergreen or subevergreen. * Sect. ''Lobatae'' (synonym ''Erythrobalanus''), the red oaks of North America, Central America and northern South America. Styles are long; the acorns mature in 18 months and taste very bitter. The inside of the acorn shell appears woolly. The actual nut is encased in a thin, clinging, papery skin. The leaves typically have sharp lobe tips, with spiny bristles at the lobe.


Subgenus ''Cerris''

* Sect. ''Cyclobalanopsis,'' the ring-cupped oaks of eastern and southeastern Asia. These are evergreen trees growing tall. They are distinct from subgenus ''Quercus'' in that they have acorns with distinctive cups bearing concrescent rings of scales; they commonly also have densely clustered acorns, though this does not apply to all of the species. Species of ''Cyclobalanopsis'' are common in the evergreen subtropical
laurel forest Laurel forest, also called laurisilva or laurissilva, is a type of subtropical forest found in areas with high humidity and relatively stable, mild temperatures. The forest is characterized by broadleaf tree species with evergreen, glossy and elo ...
s which extend from southern Japan, southern Korea, and Taiwan across southern China and northern Indochina to the eastern Himalayas, in association with trees of the genus '' Castanopsis'' and the laurel family (
Lauraceae Lauraceae, or the laurels, is a plant family that includes the true laurel and its closest relatives. This family comprises about 2850 known species in about 45 genera worldwide (Christenhusz & Byng 2016 ). They are dicotyledons, and occur ma ...
). * Sect. ''Cerris'', the Turkey oak and its relatives of Europe and Asia. Styles are long; acorns mature in 18 months and taste very bitter. The inside of the acorn's shell is hairless. Its leaves typically have sharp lobe tips, with bristles at the lobe tip. * Sect. ''Ilex,'' the Ilex oak and its relatives of Eurasia and northern Africa. Styles are medium-long; acorns mature in 12–24 months, appearing hairy on the inside. The leaves are evergreen, with bristle-like extensions on the teeth.


Fossil history

Potential records of ''Quercus'' have been reported from Late Cretaceous deposits in North America and East Asia, however these are not considered definitive. In a survey of the fossil record of ''Quercus'' it was concluded that "pre-
Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...
, and perhaps pre- Eocene occurrences of ''Quercus'' macroremains are generally represented by poorly preserved fossils that lack critical features needed for certain identification and need to be treated with caution." Amongst the oldest unequivocal records of ''Quercus'' are pollen from Austria, dating to the Paleocene-Eocene boundary, around 55 million years ago. The oldest records of ''Quercus'' in North America are from Oregon, dating to the Middle Eocene, around 44 million years ago, with the oldest records in Asia being from the Middle of Eocene of Japan; both forms have affinites to the '' Cyclobalanopsis'' group.


Phylogeny

The advent of molecular techniques for phylogenetic analysis transformed understanding of oak relationships, initially by uncovering molecular support for the diphyletic division of ''Quercus'' into Old World and New World clades. These techniques have proved highly useful in resolving fine-scale relationships among 2–5 oak species, particularly groups known to hybridize, but until recently the larger emphasis on this narrow approach prevented systematists from making large-scale determinations about oak history. As the capacity for sampling across wider swaths of oak species rose, so has resolution at the section and species level across the ''Quercus'' genus. Further advances in oak systematics are expected to arise from next-generation sequencing techniques, including a recent project to sequence the entire genome of ''Quercus robur'' (the pedunculate oak). The recent completion of that genome has uncovered an array of mutations that may underlie the evolution of longevity and disease resistance in oaks. In addition, the generation of
RAD-seq Restriction site associated DNA (RAD) markers are a type of genetic marker which are useful for association mapping, Quantitative trait locus, QTL-mapping, population genetics, ecological genetics and evolutionary genetics. The use of RAD markers ...
loci for hundreds of oak species has allowed for the construction of the most highly detailed oak phylogeny to date, although the high signal of
introgression Introgression, also known as introgressive hybridization, in genetics is the transfer of genetic material from one species into the gene pool of another by the repeated backcrossing of an interspecific hybrid with one of its parent species. Intr ...
(by hybridization) across the tree poses difficulties for deriving an unambiguous, unitary history of oaks. The phylogeny from Hipp et al 2019 is:


Ecology

Oaks are
keystone species A keystone species is a species which has a disproportionately large effect on its natural environment relative to its abundance, a concept introduced in 1969 by the zoologist Robert T. Paine. Keystone species play a critical role in maintaini ...
in a wide range of habitats from Mediterranean semi-desert to subtropical rainforest. For example, oak trees are important components of hardwood forests, and certain species are particularly known to grow in associations with members of the Ericaceae in
oak–heath forest An oak–heath forest is a plant community association and type of forest ecology. It is a deciduous forest type of well-drained, acidic soils, characterized by oaks (''Quercus'') and plants of the heath family (Ericaceae). It is commonly found in ...
s. A number of kinds of truffles, including the two well known varieties, the black Périgord truffle and the white Piedmont truffle, have symbiotic relationships with oak trees. Similarly many other mushrooms such as '' Ramaria flavosaponaria'' also associate with oaks. The European pied flycatcher is an example of an animal species that often depends upon oak trees. Many species of oaks are under threat of extinction in the wild, largely due to land use changes, livestock grazing and unsustainable harvesting. For example, over the past 200 years, large areas of oak forest in the highlands of Mexico, Central America and the northern Andes have been cleared for coffee plantations and cattle ranching. There is a continuing threat to these forests from exploitation for timber, fuelwood and charcoal. In the US, entire oak ecosystems have declined due to a combination of factors still imperfectly known, but thought to include fire suppression, increased consumption of acorns by growing mammal populations, herbivory of seedlings, and introduced pests. However, it has also been suggested that oaks as generally light-demanding trees with a relatively high tolerance for mechanic disturbances might depend on grazers like
bison Bison are large bovines in the genus ''Bison'' (Greek: "wild ox" (bison)) within the tribe Bovini. Two extant and numerous extinct species are recognised. Of the two surviving species, the American bison, ''B. bison'', found only in North Ame ...
and the clearances they create in order to regenerate successfully, thus missing them since they were extirpated in most regions following the European colonization. The mature trees shed varying numbers of acorns annually. Scientists suggest that shedding excess numbers allows the oaks to satiate nut gathering species which improves the chances of germination. Every four to ten years, certain oak populations will synchronize to produce almost no acorns at all, only to rain them down excessively the following year, known as a mast year. The year preceding the mast year is thought to starve off the mammal populations feeding on the supply, thereby increasing the effectiveness of the overproduction in the mast year that follows. This is necessary to the survival of any given oak species, as only one in 10,000 acorns results in an eventual tree.


Hybridization

Interspecific hybridization is quite common among oaks, but usually between species within the same section only, and most common in the white oak group. White oaks are unable to discriminate against pollination by other species in the same section. Because they are wind pollinated and they have weak internal barriers to hybridization, hybridization produces functional seeds and fertile hybrid offspring. Ecological stresses, especially near habitat margins, can also cause a breakdown of mate recognition as well as a reduction of male function (pollen quantity and quality) in one parent species. Frequent hybridization among oaks has consequences for oak populations around the world; most notably, hybridization has produced large populations of hybrids with copious amounts of
introgression Introgression, also known as introgressive hybridization, in genetics is the transfer of genetic material from one species into the gene pool of another by the repeated backcrossing of an interspecific hybrid with one of its parent species. Intr ...
, and the evolution of new species. Frequent hybridization and high levels of introgression have caused different species in the same populations to share up to 50% of their genetic information. Having high rates of hybridization and introgression produces genetic data that often does not differentiate between two clearly morphologically distinct species, but instead differentiates populations. Research suggests that the maintenance of particular loci for adaptation to ecological niches might explain the retention of species identity despite significant gene flow. The
Fagaceae The Fagaceae are a family of flowering plants that includes beeches, chestnuts and oaks, and comprises eight genera with about 927 species. Fagaceae in temperate regions are mostly deciduous, whereas in the tropics, many species occur as evergre ...
, or beech family, to which the oaks belong, is a very slow evolving
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
compared to other
angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s, and the patterns of hybridization and introgression in ''Quercus'' pose a great challenge to the concept of a species since a species is often defined as a group of "actually or potentially interbreeding populations which are reproductively isolated from other such groups." By this definition, many species of ''Quercus'' would be lumped together according to their geographic and ecological habitat, despite clear distinctions in morphology and, to a large extent, genetic data.


Diseases and pests

Sudden oak death (''Phytophthora ramorum'') is a water mould that can kill oaks within just a few weeks. Oak wilt, caused by the fungus ''Bretziella fagacearum'' is also a lethal disease of some oaks, particularly the red oaks (the white oaks can be infected but generally live longer). Other dangers include
wood-boring The term woodboring beetle encompasses many species and families of beetles whose larval or adult forms eat and destroy wood (i.e., are xylophagous). In the woodworking industry, larval stages of some are sometimes referred to as woodworms. The t ...
beetles, as well as root
rot Rot(s) or rotting may refer to: Decay Organic matter * Rot, decomposition of organic matter ** Dry rot, of wood ** Root rot ** Wet rot, of wood * Necrosis, of tissue Technology * Bit rot, data degradation ** Software rot, a form of bit rot * Di ...
in older trees which may not be apparent on the outside, often being discovered only when the trees come down in a strong gale. Oak apples are
gall Galls (from the Latin , 'oak-apple') or ''cecidia'' (from the Greek , anything gushing out) are a kind of swelling growth on the external tissues of plants, fungi, or animals. Plant galls are abnormal outgrowths of plant tissues, similar to be ...
s on oaks made by the gall wasp. The female kermes
scale Scale or scales may refer to: Mathematics * Scale (descriptive set theory), an object defined on a set of points * Scale (ratio), the ratio of a linear dimension of a model to the corresponding dimension of the original * Scale factor, a number ...
causes galls to grow on kermes oak. Oaks are used as food plants by the larvae of
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
( butterfly and moth) species such as the gypsy moth, ''Lymantria dispar'', which can defoliate oak and other broadleaved tree species in North America. A considerable number of galls are found on oak leaves, buds, flowers, roots, etc. Examples are oak artichoke gall, oak marble gall, oak apple gall, knopper gall, and spangle gall. A number of species of fungus cause powdery mildew on oak species. In Europe the species ''
Erysiphe alphitoides ''Erysiphe alphitoides'' is a species of fungus which causes powdery mildew on oak trees. Discovery and taxonomy Today oak powdery mildew is one of the most common diseases in European forests but it is thought to not always have been present ...
'' is the most common cause. A new and yet little understood disease of mature oaks, acute oak decline, has been reported in parts of the UK since 2009. The oak processionary moth (''Thaumetopoea processionea'') has become a serious threat in the UK since 2006. The caterpillars of this species defoliate the trees, and are hazardous to human health; their bodies are covered with poisonous hairs which can cause rashes and respiratory problems. In California, oaks are affected by the fungal disease foamy bark canker. The eastern grey squirrel (''Sciurus carolinensis'') is native in North America and an invasive species across Europe where they are known to strip bark off of a variety of large trees, including oaks. Bark stripping can result in the death of the leading shoot and decreased crown size.


Toxicity

The leaves and acorns of the oak tree are poisonous in large amounts to livestock including cattle, horses, sheep, and goats due to the toxin tannic acid, causing kidney damage and
gastroenteritis Gastroenteritis, also known as infectious diarrhea and gastro, is an inflammation of the gastrointestinal tract including the stomach and intestine. Symptoms may include diarrhea, vomiting, and abdominal pain. Fever, lack of energy, and dehydra ...
. Symptoms of poisoning include lack of appetite, depression, constipation, diarrhea (which may contain blood), blood in urine, and colic. The exception is the domestic pig, which may be fed entirely on acorns in the right conditions, and has traditionally been pastured in oak woodlands (such as the Spanish '' dehesa'' and the English system of
pannage Pannage (also referred to as ''Eichelmast'' or ''Eckerich'' in Germany, Switzerland, Liechtenstein, Luxembourg, Austria, Slovenia and Croatia) is the practice of releasing livestock-domestic pig, pigs in a forest, so that they can feed on falle ...
).
Acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera ''Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and borne ...
s are also edible by humans, after leaching of the tannins.


Uses

Oak wood has a density of about creating great strength and hardness. The wood is very resistant to insect and fungal attack because of its high tannin content. It also has very appealing grain markings, particularly when quartersawn. Oak planking was common on high status Viking longships in the 9th and 10th centuries. The wood was hewn from green logs, by axe and wedge, to produce radial planks, similar to quarter-sawn timber. Wide, quarter-sawn boards of oak have been prized since the Middle Ages for use in interior panelling of prestigious buildings such as the debating chamber of the House of Commons in London and in the construction of fine furniture. Oak wood, from '' Quercus robur'' and '' Quercus petraea'', was used in Europe for the construction of ships, especially naval
men of war ''Men of War'' is a real-time tactics video game franchise, based mainly in World War II. Main series Soldiers: Heroes of World War II ''Soldiers: Heroes of World War II'' is the original game of the 'Men of War' series, and uses an early GEM ...
, until the 19th century, and was the principal timber used in the construction of European timber-framed buildings. Today oak wood is still commonly used for
furniture Furniture refers to movable objects intended to support various human activities such as seating (e.g., stools, chairs, and sofas), eating (tables), storing items, eating and/or working with an item, and sleeping (e.g., beds and hammocks). Fu ...
making and flooring, timber-frame buildings, and veneer production. Japanese oak is used for professional drums made by Yamaha Drums. The higher density of oak gives the drum a brighter and louder tone compared to traditional materials such as maple and birch. In hill states of India, besides fuelwood and timber, the locals use oak wood for agricultural implements. The leaves are used as fodder for livestock during lean periods. Of the North American red oaks, the northern red oak is one of the most prized for lumber, and is marketed as red oak regardless of species. This wood has open capillaries, and air blown through an end grain piece 10 inches long can send bubbles out the other end into a glass of water. The openings give fungus easy access when the finish deteriorates, and natural red oak rots easily outdoors. However, if the wood is treated with a preservative compound, the capillaries absorb it deeply, and treated red oak will resist rot better than cured white oak heartwood, which has a closed cell structure. Shumard oak, a member of the red oak subgenus, provides timber described as "mechanically superior" to northern red oak. Cherrybark oak is another type of red oak that provides excellent timber. The standard lumber tree of the white oak group – all marketed as white oak – is '' Quercus alba''. White oak is often used to make wine barrels. The wood of the deciduous pedunculate oak and sessile oak accounts for most European oak production, but evergreen species such as
Holm oak Holm oak may refer to: * '' Quercus ilex'', tree native to South and Southeast Europe and parts of France * '' Quercus rotundifolia'', tree native to the Iberian Peninsula and Northwest Africa * ''Quercus agrifolia ''Quercus agrifolia'', the Cal ...
and cork oak also produce valuable timber. Oak bark is also rich in tannin, and is used by tanners for
tanning Tanning may refer to: * Tanning (leather), treating animal skins to produce leather * Sun tanning, using the sun to darken pale skin ** Indoor tanning, the use of artificial light in place of the sun ** Sunless tanning, application of a stain or d ...
leather. Oak
gall Galls (from the Latin , 'oak-apple') or ''cecidia'' (from the Greek , anything gushing out) are a kind of swelling growth on the external tissues of plants, fungi, or animals. Plant galls are abnormal outgrowths of plant tissues, similar to be ...
s were used for centuries as a main ingredient in iron gall ink for manuscripts, harvested at a specific time of year. In Korea, oak bark is used to make
shingles Shingles, also known as zoster or herpes zoster, is a viral disease characterized by a painful skin rash with blisters in a localized area. Typically the rash occurs in a single, wide mark either on the left or right side of the body or face. ...
for traditional roof construction.


Culinary

Barrels A barrel or cask is a hollow cylindrical container with a bulging center, longer than it is wide. They are traditionally made of wooden staves and bound by wooden or metal hoops. The word vat is often used for large containers for liquids, u ...
for
aging Ageing ( BE) or aging ( AE) is the process of becoming older. The term refers mainly to humans, many other animals, and fungi, whereas for example, bacteria, perennial plants and some simple animals are potentially biologically immortal. In ...
wines, sherry, and spirits such as
brandy Brandy is a liquor produced by distilling wine. Brandy generally contains 35–60% alcohol by volume (70–120 US proof) and is typically consumed as an after-dinner digestif. Some brandies are aged in wooden casks. Others are coloured with ...
, Irish whiskey,
Scotch whisky Scotch whisky (; sco, Scots whisky/whiskie, whusk(e)y; often simply called whisky or Scotch) is malt whisky or grain whisky (or a blend of the two), made in Scotland. All Scotch whisky was originally made from malted barley. Commercial distil ...
and
Bourbon whiskey Bourbon () is a type of barrel-aged American whiskey made primarily from corn. The name derives from the French Bourbon dynasty, although the precise source of inspiration is uncertain; contenders include Bourbon County in Kentucky and Bourbo ...
, are made from European and American oak, with single barrel whiskey fetching a premium. The use of oak in wine can add gustatory dimensions depending on the type of oak. Oak barrels, which may be charred before use, contribute to the colour, taste, and aroma of their potable contents, imparting a desirable oaky vanillin flavour. A dilemma for wine producers is to choose between French and American oakwoods. French oaks (''Quercus robur'', ''Q. petraea'') give greater refinement, and are chosen for the best, most expensive wines; while American oak contributes greater texture and resistance to ageing, but produces a more powerful bouquet. Oak wood chips are also used for
smoking Smoking is a practice in which a substance is burned and the resulting smoke is typically breathed in to be tasted and absorbed into the bloodstream. Most commonly, the substance used is the dried leaves of the tobacco plant, which have bee ...
fish, meat,
cheese Cheese is a dairy product produced in wide ranges of flavors, textures, and forms by coagulation of the milk protein casein. It comprises proteins and fat from milk, usually the milk of cows, buffalo, goats, or sheep. During production, ...
s, and other foods. The bark of the cork oak is used to produce wine stoppers (corks). This species grows around the Mediterranean Sea, with Portugal, Spain, Algeria, and Morocco producing most of the world's supply. The
bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, ...
of the white oak is dried and used in medical preparations.
Acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera ''Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and borne ...
s are used for making flour or roasted for acorn coffee.


Conservation

According to a comprehensive report by
The Morton Arboretum The Morton Arboretum, in Lisle, Illinois, United States, is a public garden, and outdoor museum with a library, herbarium, and program in tree research including the Center for Tree Science. Its grounds, covering 1,700 acres (6.9 square kilometres ...
and the International Union for Conservation of Nature (IUCN) an estimated 31% of the world's estimated 430 oak species are threatened with extinction, while the study found an estimated 41% of oak species to be of conservation concern. The countries with the highest numbers of threatened oak species according to the report are
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
with 36 species, Mexico with 32 species, Vietnam with 20 species and the USA with 16 species. While the cause of decline is still partly unknown for some species, the main causes the scientists determined were climate change and invasive pests in the US, and deforestation and urbanization in Asia. In the Himalayan region of India, oak forests are being invaded by pine forests due to the increase in temperature. The associated species of pine forest may cross frontiers and become new elements of the oak forests. In eastern North America, rare species of oak trees include scarlet oak ('' Quercus coccinea''), chinquapin oak ('' Quercus muehlenbergii''), and post oak ('' Quercus stellata'').


Culture


Symbols


National

The oak is a common
symbol A symbol is a mark, sign, or word that indicates, signifies, or is understood as representing an idea, object, or relationship. Symbols allow people to go beyond what is known or seen by creating linkages between otherwise very different conc ...
of strength and endurance and has been chosen as the national tree of many countries. In England, oaks have been a national symbol since at least the sixteenth century, often used by Shakespeare to convey heritage and power. In England today they remain a symbol of the nation's history, traditions, and the beauty of its countryside. Already an ancient Germanic symbol (in the form of the Donar Oak, for instance), certainly since the early nineteenth century, it stands for the nation of Germany and oak branches are thus displayed on some German coins, both of the former Deutsche Mark and the current euro currency. In 2004 the Arbor Day Foundation held a vote for the official National Tree of the United States of America. In November 2004, the United States Congress passed legislation designating the oak as America's National Tree. Other countries have also designated the oak as their national tree including Bulgaria, Croatia, Cyprus ( golden oak), Estonia, France, Germany, Moldova, Jordan,
Latvia Latvia ( or ; lv, Latvija ; ltg, Latveja; liv, Leţmō), officially the Republic of Latvia ( lv, Latvijas Republika, links=no, ltg, Latvejas Republika, links=no, liv, Leţmō Vabāmō, links=no), is a country in the Baltic region of ...
,
Lithuania Lithuania (; lt, Lietuva ), officially the Republic of Lithuania ( lt, Lietuvos Respublika, links=no ), is a country in the Baltic region of Europe. It is one of three Baltic states and lies on the eastern shore of the Baltic Sea. Lithuania ...
, Poland, Romania, Serbia, and Wales.


Regional and state

The oak is the emblem of
County Londonderry County Londonderry ( Ulster-Scots: ''Coontie Lunnonderrie''), also known as County Derry ( ga, Contae Dhoire), is one of the six counties of Northern Ireland, one of the thirty two counties of Ireland and one of the nine counties of Ulster. B ...
in Northern Ireland, as a vast amount of the county was covered in forests of the tree until relatively recently. The name of the county comes from the city of
Derry Derry, officially Londonderry (), is the second-largest city in Northern Ireland and the fifth-largest city on the island of Ireland. The name ''Derry'' is an anglicisation of the Old Irish name (modern Irish: ) meaning 'oak grove'. The ...
, which originally in Irish was known as ''Doire'' meaning "oak". The Irish
County Kildare County Kildare ( ga, Contae Chill Dara) is a county in Ireland. It is in the province of Leinster and is part of the Eastern and Midland Region. It is named after the town of Kildare. Kildare County Council is the local authority for the county, ...
derives its name from the town of Kildare which originally in Irish was ''Cill Dara'' meaning the Church of the Oak or Oak Church. In the United States, Iowa designated the oak as its official
state tree This is a list of U.S. state, federal district, and territory trees, including official trees of the following of the states, of the federal district, and of the territories. Table See also * List of U.S. state, district, and territorial insign ...
in 1961; and the white oak is the state tree of Connecticut, Illinois and Maryland. The northern red oak is the provincial tree of Prince Edward Island, as well as the state tree of New Jersey. The live oak is the state tree of the US state of Georgia. The oak is a national symbol from the
Basque Country Basque Country may refer to: * Basque Country (autonomous community), as used in Spain ( es, País Vasco, link=no), also called , an Autonomous communities of Spain, autonomous community of Spain (shown in pink on the map) * French Basque Country o ...
, especially in the province of Biscay. In
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
, the oak tree is an insignia of the
Department of Boyacá Department may refer to: * Departmentalization, division of a larger organization into parts with specific responsibility Government and military *Department (administrative division), a geographical and administrative division within a country, ...
. In 2008, the Flag of Boyacá Department was amended to include five oak leaves. The oak is a symbol of the East Bay of the San Francisco Bay Area; the coat-of-arms and flag of Oakland, California feature the oak and the logo of the East Bay Regional Park District is an oak leaf. The coats-of-arms of Vest-Agder, Norway, and Blekinge,
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
, feature oak trees. The coat-of-arms of the municipality
Eigersund Eigersund is a municipality in Rogaland county, Norway. It is in the traditional district of Dalane. The administrative centre of the municipality is the town of Egersund. The town was known for its pottery factory (closed in 1979) and it is amo ...
, Norway features an oak leaf.


Military

Oak leaves are traditionally an important part of German Army regalia. The Nazi party used the traditional German eagle, standing atop of a swastika inside a wreath of oak leaves. It is also known as the Iron Eagle. During the Third Reich of Nazi Germany, oak leaves were used for military valor decoration on the Knights Cross of the Iron Cross. They also symbolize rank in the United States Armed Forces. A gold oak leaf indicates an O-4 (
major Major (commandant in certain jurisdictions) is a military rank of commissioned officer status, with corresponding ranks existing in many military forces throughout the world. When used unhyphenated and in conjunction with no other indicators ...
or lieutenant commander), whereas a silver oak leaf indicates an O-5 (
lieutenant colonel Lieutenant colonel ( , ) is a rank of commissioned officers in the armies, most marine forces and some air forces of the world, above a major and below a colonel. Several police forces in the United States use the rank of lieutenant colone ...
or
commander Commander (commonly abbreviated as Cmdr.) is a common naval officer rank. Commander is also used as a rank or title in other formal organizations, including several police forces. In several countries this naval rank is termed frigate captain. ...
). Arrangements of oak leaves, acorns and sprigs indicate different branches of the United States Navy
staff corps A military staff or general staff (also referred to as army staff, navy staff, or air staff within the individual services) is a group of officers, enlisted and civilian staff who serve the commander of a division or other large military u ...
officers. Oak leaves are embroidered onto the covers (hats) worn by field grade officers and flag officers in the United States armed services. If a member of the United States Army or Air Force earns multiple awards of the same medal, then instead of wearing a ribbon or medal for each award, he or she wears one metal representation of an " oak leaf cluster" attached to the appropriate ribbon for each subsequent award.


Political

The oak tree is used as a symbol by a number of political parties. It is the symbol of Toryism (on account of the Royal Oak) and the Conservative Party in the United Kingdom, and formerly of the Progressive Democrats in Ireland and the Democrats of the Left in Italy. In the cultural arena, the oakleaf is the symbol of the National Trust (UK), The Woodland Trust, and
The Royal Oak Foundation The Royal Oak Foundation is an alliance of American citizens supporting the mission of the National Trust of England, Wales and Northern Ireland, which is Britain's largest heritage organisation. The foundation is headquartered in New York City. ...
.


Religion

The prehistoric Indo-European tribes worshiped the oak and connected it with a thunder or lightning god, and this tradition descended to many classical cultures. In Greek mythology, the oak is the tree sacred to Zeus, king of the gods. In Zeus's
oracle An oracle is a person or agency considered to provide wise and insightful counsel or prophetic predictions, most notably including precognition of the future, inspired by deities. As such, it is a form of divination. Description The word '' ...
in Dodona, Epirus, the sacred oak was the centerpiece of the precinct, and the priests would divine the pronouncements of the god by interpreting the rustling of the oak's leaves. Mortals who destroyed such trees were said to be punished by the gods since the ancient Greeks believed beings called
hamadryad A hamadryad (; grc, αμαδρυάδα, hamadryáda) is a Greek mythological being that lives in trees. It is a particular type of dryad which, in turn, is a particular type of nymph. Hamadryads are born bonded to a certain tree. Some maintain t ...
s inhabit them. In
Celtic polytheism Ancient Celtic religion, commonly known as Celtic paganism, was the religion of the ancient Celtic peoples of Europe. Because the ancient Celts did not have writing, evidence about their religion is gleaned from archaeology, Greco-Roman accounts ...
, the name of the oak tree was part of the Proto-Celtic word for 'druid': ''*derwo-weyd-'' > ''*-''; however, Proto-Celtic ''*derwo-'' (and ''*dru-'') can also be adjectives for 'strong' and 'firm', so Ranko Matasovic interprets that ''*-'' may mean 'strong knowledge'. As in other Indo-European faiths,
Taranis In Celtic mythology, Taranis (Proto-Celtic: *''Toranos'', earlier ''*Tonaros''; Latin: Taranus, earlier Tanarus) is the god of thunder, who was worshipped primarily in Gaul, Hispania, Britain, and Ireland, but also in the Rhineland and Danube reg ...
, being a thunder god, was associated with the oak tree. "Tree" and ''drus'' may also be cognate with "Druid," the Celtic priest to whom the oak was sacred. There has even been a study that shows that oaks are more likely to be struck by lightning than any other tree of the same height. In
Norse mythology Norse, Nordic, or Scandinavian mythology is the body of myths belonging to the North Germanic peoples, stemming from Old Norse religion and continuing after the Christianization of Scandinavia, and into the Nordic folklore of the modern period ...
, the oak was sacred to the thunder god, Thor. Thor's Oak was a sacred tree of the Germanic Chatti tribe. In Baltic and Slavic mythology, the oak was the sacred tree of Latvian god Pērkons,
Lithuanian Lithuanian may refer to: * Lithuanians * Lithuanian language * The country of Lithuania * Grand Duchy of Lithuania * Culture of Lithuania * Lithuanian cuisine * Lithuanian Jews as often called "Lithuanians" (''Lita'im'' or ''Litvaks'') by other Jew ...
Perkūnas, Prussian Perkūns and Slavic Perun, the god of thunder and one of the most important deities. The oak also appears in the Hebrew tradition. In the Bible, the oak tree at Shechem is the site where Jacob buries the foreign gods of his people (Gen. 35:4). Also, Joshua erects a stone under an oak tree as the first covenant of Yahweh, the god of Israel (Josh. 24.25–7). In Isaiah 61, the prophet refers to the Israelites as "Oaks of Righteousness". Absalom's long hair (2 Samuel 18:9) gets caught in an oak tree, and allows Joab to kill him. Veneration of the oak survives in
Serbian Orthodox Church The Serbian Orthodox Church ( sr-Cyrl, Српска православна црква, Srpska pravoslavna crkva) is one of the autocephalous (ecclesiastically independent) Eastern Orthodox Christian denomination, Christian churches. The majori ...
tradition. Christmas celebrations include the ''badnjak'', a branch taken from a young and straight oak ceremonially felled early on Christmas Eve morning, similar to a yule log. In recent times, only the branches are collected, brought home, and ceremoniously thrown into a stove or church bonfire. In another tradition, a z''apis'' (lit. "inscription") is an old, isolated oak on a hilltop or promontory, often inscribed with a cross by a parish priest. Reverence for ' probably originated in pre-Christian times, and they long remained places of public gathering and even of Christian worship where churches were not available. For example, in 1815, at a ''zapis'' assembly in Takovo, ''knez'' Miloš Obrenović declared the start of the Second Serbian Uprising. Even in modern times, cutting down ''zapis'' oaks can result in public outcry, even for projects like road building.


History

Several oak trees, such as the Royal Oak in Britain, the Charter Oak in the United States, and the
Guernica oak ''Gernikako Arbola'' ("the Tree of Gernika" in Basque) is an oak tree that symbolizes traditional freedoms for the Biscayan people, and by extension for the Basque people as a whole. The Lords of Biscay (including kings of Castile and Carlist ...
in the
Basque Country Basque Country may refer to: * Basque Country (autonomous community), as used in Spain ( es, País Vasco, link=no), also called , an Autonomous communities of Spain, autonomous community of Spain (shown in pink on the map) * French Basque Country o ...
are of great historical or cultural importance. " The Proscribed Royalist, 1651", a famous painting by John Everett Millais, depicted a Royalist fleeing from Cromwell's forces and hidden in an oak. Millais painted the picture in Hayes, Kent, from a local oak tree that became known as the Millais Oak. Approximately 50 km west of Toronto, Canada is the town of Oakville, Ontario, famous for its history as a shipbuilding port on Lake Ontario. The city of Raleigh, N.C., is known as the "City of Oaks". The Jurupa Oak tree – a clonal colony of '' Quercus palmeri'' or Palmer's oak found in Riverside County, California – is an estimated 13,000 years old. Large groups of very old oak trees are rare. One venerable group found in Poland, is about 480 years old, as assessed by dendrochronology. In the Roman Republic, a crown of oak leaves was given to those who had saved the life of a citizen in battle; it was called the " civic oak crown".


Famous specimens

: ''Individual oak trees'' * The Emancipation Oak is designated one of the 10 Great Trees of the World by the National Geographic Society and is part of the National Historic Landmark district of Hampton University in Virginia. * The Ivenack Oak which is one of the largest trees in Europe is located in Mecklenburg-Vorpommern, Germany, and is approximately 800 years old. * The
Bowthorpe Oak Bowthorpe Oak in Manthorpe near Bourne, Lincolnshire, England is perhaps England's oldest oak tree with an estimated age of over 1,000 years. The tree has a circumference of about and has a hollow trunk. The tree can be found on Bowthorpe Pa ...
, located in Bourne, Lincolnshire, is thought to be 1,000 years old. It was featured in the Guinness Book of World Records and was filmed for a TV documentary for its astonishing longevity. * The Minchenden (or Chandos) Oak, in Southgate, London, is said to be the largest oak tree in England (already in girth in the nineteenth century), and is perhaps 800 years old. * The
Seven Sisters Oak Seven Sisters Oak, located in Mandeville, Louisiana, is a notably large southern live oak in Louisiana. The tree was originally registered at the Live Oak Society as "Doby’s Seven Sisters" (No. 200) because the Doby family owned the property wh ...
is the largest certified southern live oak tree. Located in Mandeville, Louisiana, it is estimated to be up to 1,500 years old with a trunk that measures . * The Major Oak is an 800- to 1000-year-old tree located in Sherwood Forest, Nottinghamshire. According to folklore, it was used by Robin Hood for shelter. * Friendship Oak is a 500-year-old southern live oak located in Long Beach, Mississippi. * The Crouch Oak is believed to have originated in the 11th Century and is located in Addlestone, Surrey. It is an important symbol of the town with many local businesses adopting its name. It used to mark the boundary of Windsor Great Park. Legend says that Queen Elizabeth I stopped by it and had a picnic. * The Angel Oak is a southern live oak located in Angel Oak Park on
John's Island Johns Island is an island in Charleston County, South Carolina, United States, and is the largest island in the state of South Carolina. Johns Island is bordered by the Wadmalaw, Seabrook, Kiawah, Edisto, Folly, and James islands; the Stono an ...
near
Charleston, South Carolina Charleston is the largest city in the U.S. state of South Carolina, the county seat of Charleston County, and the principal city in the Charleston–North Charleston metropolitan area. The city lies just south of the geographical midpoint o ...
. The Angel Oak is estimated to be in excess of 400–500 years old, stands tall, and measures in circumference. * The Kaiser's Oak, located at the village of Gommecourt in Artois, France, named in honour of Kaiser Wilhelm II, symbolically marked from late 1914 to April 1917 the furthest point in the West of the German Imperial Army during World War I. * The Wye Oak in Maryland was the United States' largest white oak tree before it blew down in a storm in 2002, at an estimated age of 460 years. * The Bland Oak in
Sydney Sydney ( ) is the capital city of the state of New South Wales, and the most populous city in both Australia and Oceania. Located on Australia's east coast, the metropolis surrounds Sydney Harbour and extends about towards the Blue Mountain ...
,
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, planted in the 1840s, was the largest tree in Australia until it was split in a storm early on New Year Day 1941. * The Treaty Oak in
Austin Austin is the capital city of the U.S. state of Texas, as well as the seat and largest city of Travis County, with portions extending into Hays and Williamson counties. Incorporated on December 27, 1839, it is the 11th-most-populous city ...
, Texas, is a Texas live oak, and the last surviving member of the Council Oaks, a grove of 14 trees that served as a sacred meeting place for Comanche and Tonkawa tribes prior to European settlement of the area. * The Wilberforce Oak was a pollard pedunculate oak in Keston, Kent, under which William Wilberforce resolved to propose the abolition of the slave trade to the House of Commons in 1787. The original was a hollow shell by 1969; its young replacement then blew down in the Great Storm of 1987, and the third generation oak now stands in its place. File:Tamme-Lauri Tamm suvel.jpg, Tamme-Lauri oak is the thickest and oldest tree in Estonia. File:Gustave Courbet - Le Gros Chêne (1843).jpg, ''The Big Oak'', by Gustave Courbet (1843). File:Dąb Blaszak (Blaszany) w Henrykowie Gm. Szprotawa.jpg, ''Tin Oak'' in Henryków Poland; the name refers to the steel fittings that fasten the damage to the tree.


References


Bibliography

* Byfield, Liz (1990) ''An Oak Tree'', Collins book bus, London: Collins Educational, * Philips, Roger. ''Trees of North America and Europe'', Random House, New York , 1979. * Logan, William B. (2005) ''Oak: The Frame of Civilization'', New York; London: W. W. Norton, * Paterson, R.T . (1993) ''Use of Trees by Livestock'', 5: ''Quercus'', Chatham: Natural Resources Institute, * Royston, Angela (2000) ''Life Cycle of an Oak Tree'', Heinemann first library, Oxford : Heinemann Library, * Savage, Stephen (1994) ''Oak Tree'', Observing nature series, Hove: Wayland, * Tansley, Arthur G., Sir (1952) ''Oaks and Oak Woods'', Field study books, London: Methuen. * Żukow-Karczewski, Marek (1988) "Dąb – król polskich drzew" ("Oak – the king of the Polish trees"), ''AURA: A Monthly for the Protection and Shaping of Human Environment'', 9, 20–21.


External links


''Flora of China'' – ''Cyclobalanopsis''

Oak diseases

Flora Europaea: ''Quercus''

Oaks from Bialowieza Forest

Common Oaks of Florida

Oaks of the world

The Global Trees Campaign
The Red List of Oaks and Global Survey of Threatened Quercus
Latvia – the land of oaks
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