Mexcala Vicina
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''Mexcala vicina'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of jumping spider in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Mexcala ''Mexcala'' is a genus of jumping spiders that was first described by George and Elizabeth Peckham in 1902. The name is probably derived from the Nahuatl ''mezcal''. Species it contains twenty-one species, found only in Africa, Yemen, and Iran: ...
'' that is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to South Africa, found in
Mpumalanga Mpumalanga () is a province of South Africa. The name means "East", or literally "The Place Where the Sun Rises" in the Swazi, Xhosa, Ndebele and Zulu languages. Mpumalanga lies in eastern South Africa, bordering Eswatini and Mozambique. It ...
. The spider was first defined in 2009 by Wanda Wesołowska, one of over 500 that the arachnologist described during her career. It mimics ants and ant-like wasps, living alongside and preying upon them. The spider is medium-sized, with a dark brown
carapace A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tor ...
typically long and a brown
abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. ...
between long. The abdomen has a pattern made up of three dark streaks in a triangular shape. The male has a straight embolus. The female has not been identified.


Taxonomy

''Mexcala vicina'' is a jumping spider that was first described by the Polish arachnologist Wanda Wesołowska in 2009, one of over 500
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
she identified during her career. She allocated the species to the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Mexcala ''Mexcala'' is a genus of jumping spiders that was first described by George and Elizabeth Peckham in 1902. The name is probably derived from the Nahuatl ''mezcal''. Species it contains twenty-one species, found only in Africa, Yemen, and Iran: ...
'', first raised by George and Elizabeth Peckham in 1902, as part of a thorough revision of the genus. The genus was a member of the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...
Heliophaninae Chrysillini is a tribe of jumping spider in the family Salticidae. In Maddison's 2015 revision of the family, the subfamily Heliophaninae was reclassified as a junior synonym of Chrysillini. Genera * '' Afraflacilla'' * '' Augustaea'' * '' ...
alongside '' Pseudicius'' and '' Cosmophasis'', which was absorbed into
Chrysillini Chrysillini is a tribe of jumping spider in the family Salticidae. In Maddison's 2015 revision of the family, the subfamily Heliophaninae was reclassified as a junior synonym of Chrysillini. Genera * '' Afraflacilla'' * '' Augustaea'' * '' ...
by
Wayne Maddison Wayne Paul Maddison , is a professor and Canada Research Chair at the departments of zoology and botany at the University of British Columbia, and the Director of the Spencer Entomological Collection at the Beaty Biodiversity Museum. His research ...
in 2015. The tribe is a member of the
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
Saltafresia within the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Salticoida Salticoida is an unranked clade of the jumping spider family Salticidae. It is the larger and more widespread of the two subdivisions of the "typical" jumping spiders (subfamily Salticinae), occurring effectively world-wide. Its sister clade is ...
. A year later, in 2016,
Jerzy Prószyński Jerzy Prószyński (born 1935 in Warsaw) is a Polish arachnologist specializing in systematics of jumping spiders (family Salticidae). He is a graduate of the University of Warsaw, a long-term employee of the Siedlce University of Natural Scienc ...
allocated the genus to the Heliophanines group of genera, which was named after the genus ''
Heliophanus ''Heliophanus'' is a genus of the spider family Salticidae (jumping spiders). Most of the almost 170 described species occur in Africa, with many others found in the Palearctic region from Europe to Japan. Species , the World Spider Catal ...
''. The genera share characteristics, including having a rather uniform, mainly dark appearance. The species name is derived from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
word for neighbour and refers to its relationship with other spiders in the genus.


Description

Like all ''Mexcala'' spiders, the species is slender and medium-sized. The male has a dark brown
carapace A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tor ...
covered in dense brown hairs that is typically long.The eye field is darker with black markings around the eyes. The chelicerae have two small teeth at the front and one at the rear. The spider has a low brown clypeus and light brown labium, sternum and
spinneret A spinneret is a silk-spinning organ of a spider or the larva of an insect. Some adult insects also have spinnerets, such as those borne on the forelegs of Embioptera. Spinnerets are usually on the underside of a spider's opisthosoma, and are ...
s. The
abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. ...
is between long, brown and hairy with a triangular pattern of three darker streaks on the back. The underside is plain brown. It has light brown
legs A leg is a weight-bearing and locomotive anatomical structure, usually having a columnar shape. During locomotion, legs function as "extensible struts". The combination of movements at all joints can be modeled as a single, linear element ...
. The pedipalps are yellow. The
palpal bulb The two palpal bulbs – also known as palpal organs and genital bulbs – are the copulatory organs of a male spider. They are borne on the last segment of the pedipalps (the front "limbs" of a spider), giving the spider an appearance often descr ...
has a wide tibial apophysis that has a hooked end. The embolus is straight and fixed to the tegulum. The female has not been described.


Behaviour

Like many jumping spiders, Wesołowska and Tamás Szűts noted that ''Mexcala'' spiders mimic ants. The species lives amongst the species of ant and ant-like wasps that it mimics. Like other jumping spiders, it is mainly a diurnal hunter that uses its good eyesight to spot its prey. It attacks from the front and captures its prey behind the head. The chelicerae have short thick spines on them which may be used for digging holes to act as underground hiding places. It uses visual displays during courtship and transmits vibratory signals through silk to communicate to other spiders.


Distribution

''Mexcala'' spiders can be found across Africa and the Arabian peninsula. ''Mexcala vicina'' lives in Cameroon and Republic of the Congo. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
was found near Batouri in Cameroon in 1976. The first example found in Congo was discovered in 1956.


References


Citations


Bibliography

* * * * * * * * {{Taxonbar, from=Q2286170 Arthropods of Cameroon Fauna of the Republic of the Congo Salticidae Spiders described in 2009 Spiders of South Africa Taxa named by Wanda Wesołowska