List Of Utricularia Species
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There are around 240 species in the genus ''
Utricularia ''Utricularia'', commonly and collectively called the bladderworts, is a genus of carnivorous plants consisting of approximately 233 species (precise counts differ based on classification opinions; a 2001 publication lists 215 species).Salmon, Br ...
'', belonging to the bladderwort family (
Lentibulariaceae Lentibulariaceae is a family of carnivorous plants containing three genera: ''Genlisea'', the corkscrew plants; ''Pinguicula'', the butterworts; and ''Utricularia'', the bladderworts. The genera ''Polypompholyx'' (two species of pink petticoats ...
). It is the largest genus of
carnivorous plants Carnivorous plants are plants that derive some or most of their nutrients from trapping and consuming animals or protozoans, typically insects and other arthropods. Carnivorous plants still generate some of their energy from photosynthesis ...
and has a worldwide distribution, being absent only from Antarctica and the oceanic islands. This genus was considered to have 250 species until Peter Taylor reduced the number to 214 in his exhaustive study, ''
The genus Utricularia - a taxonomic monograph ''The Genus Utricularia: A Taxonomic Monograph'' is a monograph by Peter Taylor on the carnivorous plant genus ''Utricularia'', the bladderworts. It was published in 1989 by Her Majesty's Stationery Office (HMSO) as the fourteenth entry in the '' ...
'', published by
HMSO The Office of Public Sector Information (OPSI) is the body responsible for the operation of His Majesty's Stationery Office (HMSO) and of other public information services of the United Kingdom. The OPSI is part of the National Archives of the U ...
(1989). Taylor's classification is generally accepted, though his division of the genus into two subgenera was soon seen as obsolete. Molecular genetic studies have mostly confirmed Taylor's sections with some modifications (Jobson et al., 2003), but reinstalled the division of the genus in three subgenera. This list follows the subgeneric classification ''sensu'' Müller & Borsch (2005), updated with new information in Müller et al. (2006). For a list of known ''Utricularia'' species by common name, see ''Utricularia'' species by common name.


Subgenus '' Bivalvaria''


Section '' Aranella''

*'' Utricularia blanchetii'' *'' Utricularia costata'' *'' Utricularia fimbriata'' *'' Utricularia laciniata'' *'' Utricularia longeciliata'' *'' Utricularia parthenopipes'' *'' Utricularia purpureocaerulea'' *'' Utricularia rostrata'' *''
Utricularia sandwithii ''Utricularia sandwithii'' is a small, probably perennial, carnivorous plant that belongs to the genus ''Utricularia''. ''U. sandwithii'' is endemic to Brazil, Guyana, Suriname, and Venezuela. It grows as a terrestrial plant in damp, sandy s ...
'' *''
Utricularia simulans ''Utricularia simulans'', the fringed bladderwort, is a small to medium-sized, probably perennial, species of carnivorous plant in the family Lentibulariaceae. ''Utricularia simulans'' is native to tropical Africa and the Americas. It grows ...
''


Section '' Australes''

*''
Utricularia delicatula ''Utricularia delicatula'' is a terrestrial species of bladderwort and is unique within its genus in being endemic to New Zealand.Bruce Salmon (2001). ''Carnivorous Plants of New Zealand''. Ecosphere Publications The specific epithet is Latin f ...
'' *'' Utricularia lateriflora'' *''
Utricularia simplex ''Utricularia simplex'', commonly known as bluecoats, is a very small perennial carnivorous plant that belongs to the genus ''Utricularia''. ''U. simplex'' is endemic to Western Australia. It grows as a terrestrial plant in peaty soils in h ...
''


Section '' Avesicarioides''

*''
Utricularia rigida ''Utricularia rigida'' is a small to medium-sized perennial, rheophytic carnivorous plant that belongs to the genus ''Utricularia''. ''U. rigida'' is endemic to western tropical Africa, where it can be found in Côte d'Ivoire, Guinea, Guine ...
'' *''
Utricularia tetraloba ''Utricularia tetraloba'' is a very small, probably perennial, rheophytic carnivorous plant that belongs to the genus ''Utricularia''. ''U. tetraloba'' is endemic to Guinea and Sierra Leone. It grows as a rheophyte on rocks in shallow runni ...
''


Section '' Benjaminia''

*''
Utricularia nana ''Utricularia nana'' is a carnivorous plant of the Lentibulariaceae family and is the only species in the section ''Benjaminia''. It is native to the South American countries of Venezuela, Guyana, Suriname, French Guinea, Brazil and Paraguay. ...
''


Section '' Calpidisca''

*''
Utricularia arenaria ''Utricularia arenaria'' is a small annual carnivorous plant that belongs to the genus ''Utricularia''. It is native to tropical and southern Africa, where it can be found in Angola, Burundi, Cameroon, Côte d'Ivoire, the Democratic Republic of t ...
'' *''
Utricularia bisquamata ''Utricularia bisquamata'' is a small annual carnivorous plant that belongs to the genus ''Utricularia''. It is native to southern Africa, where it can be found in Angola, Lesotho, Madagascar, Namibia, and South Africa. ''U. bisquamata'' gro ...
'' *''
Utricularia firmula ''Utricularia firmula'' is a small annual carnivorous plant that belongs to the genus ''Utricularia''. It is native to tropical and southern Africa, where it can be found in Angola, Cameroon, Côte d'Ivoire, the Democratic Republic of the Congo, ...
'' *''
Utricularia livida ''Utricularia livida'', the leaden bladderwort, is a species of flowering plant in the bladderwort family, native to central and southern Africa, and Mexico. Growing to tall and broad, it is a carnivorous perennial. It was originally described ...
'' *''
Utricularia microcalyx ''Utricularia microcalyx'' is a small to medium-sized annual carnivorous plant that belongs to the genus ''Utricularia''. It is native to tropical Africa, where it can be found in the Democratic Republic of the Congo and Zambia. ''U. microca ...
'' *'' Utricularia odontosepala'' *'' Utricularia pentadactyla'' *''
Utricularia sandersonii ''Utricularia sandersonii'', Sanderson's bladderwort, is a species of flowering plant in the bladderwort family. Originally described and published by the British botanist Daniel Oliver in 1865, it is a carnivorous evergreen perennial, endemic t ...
'' *'' Utricularia troupinii'' *'' Utricularia welwitschii''


Section '' Enskide''

*'' Utricularia chrysantha'' *'' Utricularia fulva''


Section '' Lloydia''

*'' Utricularia pubescens''


Section '' Minutae''

*'' Utricularia simmonsii''Lowrie et al. (2008) notes the placement of section ''Minutae'' appears, morphologically, to be closer to subgenus ''Bivalvaria''. This was confirmed by a molecular study (Reut & Jobson, 2010)


Section '' Nigrescentes''

*'' Utricularia bracteata'' *'' Utricularia caerulea'' *'' Utricularia warburgii''


Section '' Oligocista''

*'' Utricularia adpressa'' *'' Utricularia albocaerulea'' *'' Utricularia andongensis'' *'' Utricularia arcuata'' *'' Utricularia babui'' *'' Utricularia bifida'' *'' Utricularia bosminifera'' *'' Utricularia cecilii'' *'' Utricularia chiribiquitensis'' *'' Utricularia circumvoluta'' *'' Utricularia delphinioides'' *'' Utricularia densiflora'' *'' Utricularia erectiflora'' *'' Utricularia foveolata'' *'' Utricularia graminifolia'' *'' Utricularia heterosepala'' *'' Utricularia involvens'' *'' Utricularia jackii'' *'' Utricularia laxa'' *'' Utricularia lazulina'' *'' Utricularia letestui'' *'' Utricularia lloydii'' *'' Utricularia macrocheilos'' *'' Utricularia malabarica'' *'' Utricularia meyeri'' *'' Utricularia micropetala'' *'' Utricularia odorata'' *'' Utricularia pierrei'' *'' Utricularia pobeguinii'' *'' Utricularia polygaloides'' *'' Utricularia praeterita'' *'' Utricularia prehensilis'' *'' Utricularia recta'' *'' Utricularia reticulata'' *'' Utricularia scandens'' *'' Utricularia smithiana'' *'' Utricularia spiralis'' *'' Utricularia subramanyamii'' *'' Utricularia tortilis'' *'' Utricularia uliginosa'' *'' Utricularia vitellina'' *'' Utricularia wightiana''


Section '' Phyllaria''

*'' Utricularia brachiata'' *'' Utricularia christopheri'' *'' Utricularia corynephora'' *'' Utricularia forrestii'' *'' Utricularia furcellata'' *'' Utricularia garrettii'' *'' Utricularia inthanonensis'' *'' Utricularia kumaonensis'' *'' Utricularia moniliformis'' *'' Utricularia multicaulis'' *'' Utricularia phusoidaoensis'' *'' Utricularia pulchra'' *'' Utricularia salwinensis'' *'' Utricularia spinomarginata'' *'' Utricularia steenisii'' *'' Utricularia striatula''


Section '' Stomoisia''

*'' Utricularia cornuta'' *''
Utricularia juncea ''Utricularia juncea'', the southern bladderwort, is a small to medium-sized, probably perennial carnivorous plant that belongs to the genus ''Utricularia''. ''U. juncea'' is native to Central, South, and North America. It grows as a terrestrial ...
''


Subgenus '' Polypompholyx''


Section '' Pleiochasia''

*'' Utricularia albiflora'' *'' Utricularia ameliae'' *'' Utricularia barkeri'' *'' Utricularia beaugleholei'' *'' Utricularia benthamii'' *'' Utricularia blackmanii'' *''
Utricularia dichotoma ''Utricularia dichotoma'', commonly known as fairy aprons, is a variable, perennial species of terrestrial bladderwort. It is a widespread species with mauve or purple fan-shaped flowers on a slender stalk and usually grows in wet locations. ...
'' *'' Utricularia fenshamii'' *'' Utricularia fistulosa'' *'' Utricularia grampiana'' *'' Utricularia hamiltonii'' *'' Utricularia helix'' *'' Utricularia inaequalis'' *'' Utricularia menziesii'' *'' Utricularia paulineae'' *'' Utricularia petertaylorii'' *'' Utricularia singeriana'' *'' Utricularia terrae-reginae'' *'' Utricularia triflora'' *'' Utricularia tubulata'' *'' Utricularia violacea'' *'' Utricularia volubilis''


Section '' Lasiocaules''Section ''Pleiochasia'' was divided into two sections (''Pleiochasia'' and ''Lasiocaules'') based upon molecular data in a phylogenetic study on subgenus ''Polypompholyx'' (Jobson ''et al.'', 2017).

* '' Utricularia albertiana'' * '' Utricularia antennifera'' * '' Utricularia arnhemica'' * '' Utricularia bidentata'' *'' Utricularia capilliflora'' *'' Utricularia cheiranthos'' *'' Utricularia dunlopii'' *''
Utricularia dunstaniae ''Utricularia dunstaniae'' is an annual terrestrial carnivorous plant that belongs to the genus ''Utricularia'' (family Lentibulariaceae). Its distribution ranges from northern Western Australia to the Northern Territory.Taylor, Peter. (1989). ...
'' *'' Utricularia georgei'' *''
Utricularia holtzei ''Utricularia holtzei'' is an annual affixed aquatic carnivorous plant that belongs to the genus ''Utricularia'' (family Lentibulariaceae). It is endemic to the Northern Territory.Taylor, Peter. (1989). '' The genus Utricularia - a taxonomic mo ...
'' *'' Utricularia kamienskii'' *'' Utricularia kenneallyi'' *'' Utricularia kimberleyensis'' *'' Utricularia lasiocaulis'' *'' Utricularia leptorhyncha'' *'' Utricularia lowriei'' *'' Utricularia magna'' *'' Utricularia papilliscapa'' *'' Utricularia quinquedentata'' *'' Utricularia rhododactylos'' *'' Utricularia tridactyla'' * '' Utricularia uniflora'' *'' Utricularia wannanii''


Section '' Polypompholyx''

*'' Utricularia multifida'' *'' Utricularia tenella''


Section '' Tridentaria''

*'' Utricularia westonii''


Subgenus ''

Utricularia ''Utricularia'', commonly and collectively called the bladderworts, is a genus of carnivorous plants consisting of approximately 233 species (precise counts differ based on classification opinions; a 2001 publication lists 215 species).Salmon, Br ...
''


Section '' Avesicaria''

*'' Utricularia neottioides'' *'' Utricularia oliveriana''


Section ''

Candollea ''Stylidium'' (also known as triggerplants or trigger plants) is a genus of dicotyledonous plants that belong to the family Stylidiaceae. The genus name ''Stylidium'' is derived from the Greek ''στύλος'' or ''stylos'' (column or pillar), wh ...
''

*'' Utricularia podadena''


Section ''

Chelidon Chelidon (fl. 74 BC) was a Roman courtesan, famed for her influence during the praetorship of Gaius Verres. She was a freedwoman and a successful professional high class courtesan. She was introduced to Gaius Verres by the courtesan Pippa and Ter ...
''

*'' Utricularia mannii''


Section '' Choristothecae''

*'' Utricularia choristotheca'' *'' Utricularia determannii''


Section '' Foliosa''

*''
Utricularia amethystina ''Utricularia amethystina'', the Florida purple bladderwort, is a variable species of terrestrial bladderwort native to Bolivia, Brazil, Guyana, Peru, and south-west Florida Florida is a state located in the Southeastern region of the Un ...
'' *'' Utricularia calycifida'' *'' Utricularia hintonii'' *'' Utricularia hispida'' *''
Utricularia huntii ''Utricularia huntii'' is a medium-sized perennial terrestrial carnivorous plant that belongs to the genus '' Utricularia''. ''U. huntii'' is endemic to Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portugues ...
'' *'' Utricularia longifolia'' *'' Utricularia panamensis'' *'' Utricularia petersoniae'' *'' Utricularia praelonga'' *'' Utricularia regia'' *'' Utricularia schultesii'' *'' Utricularia tricolor'' *'' Utricularia tridentata''


Section '' Kamienskia''

*''
Utricularia mangshanensis ''Utricularia mangshanensis'' is a carnivorous plant belonging to the genus ''Utricularia''. This species is known only from the Mangshan Mountains of Hunan Province, China China, officially the People's Republic of China (PRC), is a co ...
'' *''
Utricularia peranomala ''Utricularia peranomala'' is a very small, annual, bryophilous lithophytic carnivorous plant that belongs to the genus ''Utricularia''. ''U. peranomala'' is endemic to China, where it is only known from the type location in the hilly nort ...
''


Section '' Lecticula''

*'' Utricularia resupinata'' *'' Utricularia spruceana''


Section '' Martinia''

*'' Utricularia tenuissima''


Section '' Meionula''

*'' Utricularia geoffrayi'' *'' Utricularia hirta'' *'' Utricularia minutissima''


Section '' Mirabiles''

*''
Utricularia heterochroma ''Utricularia heterochroma'' is a very small, possibly perennial, rheophytic carnivorous plant that belongs to the genus ''Utricularia''. ''U. heterochroma'' is endemic to Venezuela, where it is known from the type location at Ptari-tepui ...
'' *''
Utricularia mirabilis ''Utricularia mirabilis'' is a small, perennial, rheophytic carnivorous plant that belongs to the genus ''Utricularia''. ''U. mirabilis'' is endemic to Venezuela, where it is known from the type location near Campo Grande. It grows as a rh ...
''


Section '' Nelipus''

*'' Utricularia biloba'' *'' Utricularia leptoplectra'' *'' Utricularia limosa''


Section '' Oliveria''

*'' Utricularia appendiculata''


Section '' Orchidioides''

*'' Utricularia alpina'' *'' Utricularia asplundii'' *'' Utricularia buntingiana'' *'' Utricularia campbelliana'' *'' Utricularia cornigera'' *'' Utricularia endresii'' *'' Utricularia geminiloba'' *'' Utricularia humboldtii'' *'' Utricularia jamesoniana'' *'' Utricularia nelumbifolia'' *'' Utricularia nephrophylla'' *'' Utricularia praetermissa'' *'' Utricularia quelchii'' *'' Utricularia reniformis'' *'' Utricularia unifolia'' *'' Utricularia uxoris''


Section '' Setiscapella''

*'' Utricularia flaccida'' *'' Utricularia nervosa'' *'' Utricularia nigrescens'' *'' Utricularia physoceras'' *''
Utricularia pusilla ''Utricularia pusilla'', the tiny bladderwort, is an annual, terrestrial carnivorous plant that belongs to the genus ''Utricularia'' (family Lentibulariaceae). Its distribution includes ranges in Mexico, the Caribbean and Central and South Americ ...
'' *'' Utricularia stanfieldii'' *'' Utricularia subulata'' *'' Utricularia trichophylla'' *'' Utricularia triloba''


Section '' Sprucea''

*'' Utricularia viscosa''


Section '' Steyermarkia''

*'' Utricularia aureomaculata'' *'' Utricularia cochleata'' *'' Utricularia steyermarkii''


Section '' Stylotheca''

*'' Utricularia guyanensis''


Section ''

Utricularia ''Utricularia'', commonly and collectively called the bladderworts, is a genus of carnivorous plants consisting of approximately 233 species (precise counts differ based on classification opinions; a 2001 publication lists 215 species).Salmon, Br ...
''

*'' Utricularia aurea'' *''
Utricularia australis ''Utricularia australis'' is a medium-sized, perennial species of aquatic bladderwort.Bruce Salmon (2001) ''"Carnivorous Plants of New Zealand"'' Ecosphere Publications This species has a vast geographic range, being found throughout Europe, in ...
'' *'' Utricularia benjaminiana'' *'' Utricularia biovularioides'' *'' Utricularia bremii'' *''
Utricularia breviscapa ''Utricularia breviscapa'' is a small to medium suspended aquatic carnivorous plant that belongs to the genus ''Utricularia'' (family Lentibulariaceae). It is probably an annual plant. Its native distribution includes the Antilles and South Ame ...
'' *'' Utricularia chiakiana'' *'' Utricularia corneliana'' *'' Utricularia cymbantha'' *''
Utricularia dimorphantha ''Utricularia dimorphantha'' is a medium-sized suspended aquatic carnivorous plant that belongs to the genus ''Utricularia'' (family Lentibulariaceae). It is a perennial plant. It is endemic to Japan.Taylor, Peter. (1989). ''The genus Utricular ...
'' *'' Utricularia floridana'' *'' Utricularia foliosa'' *'' Utricularia geminiscapa'' *''
Utricularia gibba ''Utricularia gibba'', commonly known as the humped or floating bladderwort, is a small, mat-forming species of carnivorous aquatic bladderwort. It is found on all continents except Antarctica. ''U. gibba'' has an exceptionally small genome for ...
'' *'' Utricularia hydrocarpa'' *'' Utricularia incisa'' *''
Utricularia inflata ''Utricularia inflata'', commonly known as the swollen bladderwort, inflated bladderwort, or large floating bladderwort, is a large suspended aquatic carnivorous plant that belongs to the genus ''Utricularia''. It is a perennial that is native to ...
'' *'' Utricularia inflexa'' *'' Utricularia intermedia'' *'' Utricularia macrorhiza'' *'' Utricularia minor'' *'' Utricularia muelleri'' *'' Utricularia naviculata'' *''
Utricularia ochroleuca ''Utricularia ochroleuca'', the yellowishwhite bladderwort, pale bladderwort, or cream-flowered bladderwort, is a small, perennial carnivorous plant that belongs to the genus ''Utricularia''. It is usually found affixed to the substrate. ''U. o ...
'' *'' Utricularia olivacea'' *''
Utricularia perversa ''Utricularia perversa'' is a medium-sized suspended aquatic carnivorous plant that belongs to the genus ''Utricularia''. It is probably most closely related to '' U. radiata''. ''U. perversa'' is endemic to the Mexican states of Chihuahua and ...
'' *'' Utricularia platensis'' *'' Utricularia poconensis'' *'' Utricularia punctata'' *''
Utricularia radiata ''Utricularia radiata'', the little floating bladderwort, is a medium-sized suspended aquatic carnivorous plant that belongs to the genus ''Utricularia''. ''U. radiata'' is endemic to North America.Taylor, Peter. (1989). ''The genus Utricularia - ...
'' *'' Utricularia raynalii'' *'' Utricularia reflexa'' *'' Utricularia stellaris'' *'' Utricularia striata'' *''
Utricularia stygia ''Utricularia stygia'', the arctic bladderwort or Northern bladderwort, is an affixed aquatic carnivorous plant that belongs to the genus ''Utricularia''. ''U. stygia'' is native to northern Europe and North America. This species was originally ...
'' *''
Utricularia vulgaris ''Utricularia vulgaris'' (greater bladderwort or common bladderwort) is an aquatic species of bladderwort found in Asia and Europe. The plant is free-floating and does not put down roots. Stems can attain lengths of over one metre in a single g ...
'' *'' Utricularia warmingii''


Section '' Vesiculina''

*'' Utricularia cucullata'' *'' Utricularia myriocista'' *'' Utricularia purpurea''


See also

*'' Utricularia capillacea'' *'' Utricularia linearis''


Notes


References

*Lowrie A, Cowie ID, and Conran JG. (2008). A new species and section of ''Utricularia'' (Lentibulariaceae) from northern Australia. ''Telopea'', 12(1): 31-46. * Jobson RW, Playford J, Cameron KM, Albert VA. (2003). Molecular phylogenetics of Lentibulariaceae inferred from plastid ''rps''16 intron and ''trn''LF DNA sequences: implications for character evolution and biogeography. ''Systematic Botany'', 28(1): 157-171. *Jobson RW, Baleeiro PC, Reut MS (2017). Molecular phylogeny of subgenus ''Polypompholyx'' (''Utricularia''; Lentibulariaceae) based on three plastid markers: diversification and proposal for a new section. ''Australian Systematic Botany'', 30: 259-278. https://doi.org/10.1071/SB17003 *Jobson RW, Baleeiro PC, Barrett MD (2018). Six new species of ''Utricularia'' (Lentibulariaceae) from Northern Australia. ''Telopea'', 21: 57-77. https://doi.org/10.7751/telopea12630 *Müller KF and Borsch T. (2005). Phylogenetics of ''Utricularia'' (Lentibulariaceae) and molecular evolution of the ''trn''K intron in a lineage with high substitutional rates. ''Plant Systematics and Evolution'', 250: 39-67. *Müller KF, Borsch T, Legendre L, Porembski S, and Barthlott W. (2006). Recent progress in understanding the evolution of carnivorous Lentibulariaceae (Lamiales). ''Plant Biology'', 8: 748-757. *Taylor, Peter. (1989). ''
The genus Utricularia - a taxonomic monograph ''The Genus Utricularia: A Taxonomic Monograph'' is a monograph by Peter Taylor on the carnivorous plant genus ''Utricularia'', the bladderworts. It was published in 1989 by Her Majesty's Stationery Office (HMSO) as the fourteenth entry in the '' ...
''. Kew Bulletin Additional Series XIV: London.


External links


A large ''Utricularia'' photo gallery
{{CarnivorousPlants *
Utricularia ''Utricularia'', commonly and collectively called the bladderworts, is a genus of carnivorous plants consisting of approximately 233 species (precise counts differ based on classification opinions; a 2001 publication lists 215 species).Salmon, Br ...
Utricularia, List