Birds
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
evolved from certain
feathered theropod
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaurs, and there is no real dividing line between birds and
non-avian dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutiona ...
s except that some of the former survived the
Cretaceous–Paleogene extinction event
The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...
while the latter did not. For the purposes of this article, a 'bird' is considered to be any member of the clade
Aves
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
in the broadest sense.
[Sereno (2005)] Some dinosaur groups which may or may not be true birds are listed below under
Proto-birds.
This page contains a listing of prehistoric bird taxa only known from completely
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
ized specimens. These extinctions took place before the Late
Quaternary
The Quaternary ( ) is the current and most recent of the three periods of the Cenozoic Era in the geologic time scale of the International Commission on Stratigraphy (ICS). It follows the Neogene Period and spans from 2.58 million years ...
and thus took place in the absence of significant human interference. While the earliest
hominids
The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); ''Gorilla'' (the ...
had been eating birds and especially their eggs,
human population
Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
and
technology
Technology is the application of knowledge to reach practical goals in a specifiable and reproducible way. The word ''technology'' may also mean the product of such an endeavor. The use of technology is widely prevalent in medicine, science, ...
was simply insufficient to seriously affect healthy bird populations until the
Upper Paleolithic Revolution
Behavioral modernity is a suite of behavioral and cognitive traits that distinguishes current ''Homo sapiens'' from other anatomically modern humans, hominins, and primates. Most scholars agree that modern human behavior can be characterized b ...
. Rather, reasons for the extinctions listed here are stochastic abiotic events such as
bolide
A bolide is normally taken to mean an exceptionally bright meteor, but the term is subject to more than one definition, according to context. It may refer to any large crater-forming body, or to one that explodes in the atmosphere. It can be a ...
impacts,
climate changes due to orbital shifts, mass volcanic eruptions etc. Alternatively, species may have gone extinct due to
evolution
Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
ary displacement by successor or competitor taxa – it is notable that an extremely large number of seabirds have gone extinct during the mid-
Tertiary
Tertiary ( ) is a widely used but obsolete term for the geologic period from 66 million to 2.6 million years ago.
The period began with the demise of the non-avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start ...
; this seems at least partly due to competition by the contemporary radiation of
marine mammal
Marine mammals are aquatic mammals that rely on the ocean and other marine ecosystems for their existence. They include animals such as seals, whales, manatees, sea otters and polar bears. They are an informal group, unified only by their ...
s.
The relationships of these
taxa
In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
are often hard to determine, as many are known only from very fragmentary remains and due to the complete fossilization precluding analysis of information from
DNA,
RNA
Ribonucleic acid (RNA) is a polymeric molecule essential in various biological roles in coding, decoding, regulation and expression of genes. RNA and deoxyribonucleic acid ( DNA) are nucleic acids. Along with lipids, proteins, and carbohydra ...
or protein sequencing. The taxa listed in this article should be classified with the
Wikipedia conservation status category "Fossil".
Before the late 19th century, when
mineral
In geology and mineralogy, a mineral or mineral species is, broadly speaking, a solid chemical compound with a fairly well-defined chemical composition and a specific crystal structure that occurs naturally in pure form.John P. Rafferty, ed. (2 ...
s were still considered one of the
kingdom
Kingdom commonly refers to:
* A monarchy ruled by a king or queen
* Kingdom (biology), a category in biological taxonomy
Kingdom may also refer to:
Arts and media Television
* ''Kingdom'' (British TV series), a 2007 British television drama s ...
s of
binomial nomenclature
In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
, fossils were often treated according to a parallel
taxonomy
Taxonomy is the practice and science of categorization or classification.
A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
. Rather than assigning them to animal or plant genera, they were treated as mineral genera and given
binomial name
In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
s typically using ''Osteornis'' ("bone-bird") or ''Ornitholithus'' ("bird fossil") as "
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
". The latter name, however, is still in use for an
oogenus
Egg fossils are the fossilized remains of eggs laid by ancient animals. As evidence of the physiological processes of an animal, egg fossils are considered a type of trace fossil. Under rare circumstances a fossil egg may preserve the remains of ...
of
fossil bird eggs. Also, other animals (in particular
pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 ...
s) were placed in these "genera". In sources pre-dating the
Linnean system, the above terms are also seen in the more extensive descriptions used to name
taxa
In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
back then.
Taxonomic list of fossil prehistoric birds
Higher-level
taxa
In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
are presented in likely or suspected
phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
order.
Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
-level taxa and lower are sorted chronologically, in ascending order (i.e., older taxa first).
The higher-level groups of non-
Neornithes
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
are arranged based on the
phylogeny
A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spe ...
proposed by
Luis Chiappe, updated and expanded to incorporate recent research. The categories are inclusive in ascending order.
''Taxonomic assignments, especially in the pygostylian to early neornithine genera, are still very provisional and subject to quite frequent change.''
Basal
Avialae
Avialae ("bird wings") is a clade containing the only living dinosaurs, the birds. It is usually defined as all theropod dinosaurs more closely related to birds (Aves) than to deinonychosaurs, though alternative definitions are occasionally use ...
(extinct)
The most primitive "birds", usually still possessing a long bony tail with generally unfused vertebrae. Not all of these may be on the line of bird ancestors; whether they are not closer to other
theropods
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ca ...
groups than to the
Avialae
Avialae ("bird wings") is a clade containing the only living dinosaurs, the birds. It is usually defined as all theropod dinosaurs more closely related to birds (Aves) than to deinonychosaurs, though alternative definitions are occasionally use ...
remains to be thoroughly tested (see ''
Xiaotingia
''Xiaotingia'' is a genus of anchiornithid theropod dinosaur from Middle Jurassic or early Late Jurassic deposits of western Liaoning, China, containing a single species, ''Xiaotingia zhengi''.
Discovery
''Xiaotingia'' is known from the hol ...
'').
* †''
Anchiornis huxleyi
''Anchiornis'' is a genus of small, four-winged paravian dinosaurs, with only one known species, the type species ''Anchiornis huxleyi'', named for its similarity to modern birds. The Latin name ''Anchiornis'' derives from a Greek word meaning " ...
''
* †''
Aurornis xui
''Aurornis'' is an extinct genus of anchiornithid theropod dinosaurs from the Jurassic period of China. The genus ''Aurornis'' contains a single known species, ''Aurornis xui'' (). ''Aurornis xui'' may be the most basal ("primitive") avialan d ...
''
* †''
Balaur bondoc
''Balaur bondoc'' is a species of theropod dinosaur from the late Cretaceous period, in what is now Romania. It is the type species of the monotypic genus ''Balaur'', after the ''balaur'' (), a dragon of Romanian folklore. The specific name ''b ...
''
* †''
Jixiangornis orientalis
''Jixiangornis'' is a genus of basal avialans from the Early Cretaceous. Like later avialans, it had no teeth, but it also had a long tail, unlike modern birds. Since teeth were still present in some more derived short-tailed avialans, ''Jixian ...
''
* †''
Xiaotingia zhengi''
* †
Yandangornithiformes
** †
Yandangornithidae
*** †''
Yandangornis longicaudatus'' (Late Cretaceous)
* †
Archaeopterygiformes
** †
Archaeopterygidae
Archaeopterygidae is a group of maniraptoran dinosaurs, known from the latest Jurassic and earliest Cretaceous of Europe. In most current classifications, it contains only the genera ''Archaeopteryx'' and ''Wellnhoferia''. As its name suggests, ...
rchaeornithidae ; Archaeopteridae ; Archornithidae '
a''
*** †''
Wellnhoferia grandis
''Wellnhoferia'' (named after Peter Wellnhofer) is a genus of early prehistoric bird-like theropod dinosaur closely related to '' Archaeopteryx''. It lived in what is now Germany, during the Late Jurassic. While ''Wellnhoferia'' was similar to '' ...
'' (Late Jurassic) – possible synonym of ''Archaeopteryx''
*** †''
Archaeopteryx''
'Archaeornis''_;_''Griphornis.html" ;"title="Archaeornis.html" ;"title="'Archaeornis">'Archaeornis'' ; ''Griphornis">Archaeornis.html" ;"title="'Archaeornis">'Archaeornis'' ; ''Griphornis'' ; ''Griphosaurus'' ] (Late Jurassic)
* †Jeholornithiformes
** †Jeholornithidae
*** †''Dalianraptor, Dalianraptor cuhe''
*** †''Jeholornis'' [''Shenzhouraptor'' ] (Early Cretaceous)
Basal
Pygostylia
Pygostylia is a group of avialans which includes the Confuciusornithidae and all of the more advanced species, the Ornithothoraces.
Definition
The group Pygostylia was intended to encompass all avialans with a short, stubby tail, as opposed to ...
(extinct)
The earliest birds with a modern
pygostyle
Pygostyle describes a skeletal condition in which the final few caudal vertebrae are fused into a single ossification, supporting the tail feathers and musculature. In modern birds, the rectrices attach to these. The pygostyle is the main compone ...
: a reduction and fusion of the tail vertebrae; possibly a
paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
group. Two types of pygostyle are known, a rod-shaped one found in
Confuciusornithidae
Confuciusornithidae is an extinct family of pygostylian avialans known from the Early Cretaceous, found in northern China. They are commonly placed as a sister group to Ornithothoraces, a group that contains all extant birds along with their clo ...
,
Enantiornithes and some non-avian theropods such as ''
Nomingia
''Nomingia'' is a genus of oviraptorid theropod dinosaur hailing from the Late Cretaceous Bugin Tsav Beds of Mongolia.
Discovery and naming
The remains, consisting of most of the vertebral column, pelvic girdle and left tibio-tarsus, holot ...
'', and a
plowshare
In agriculture, a plowshare ( US) or ploughshare ( UK; ) is a component of a plow (or plough). It is the cutting or leading edge of a moldboard which closely follows the coulter (one or more ground-breaking spikes) when plowing.
The plowsh ...
-shaped one, only known in the lineage leading to modern birds. It is not certain that the pygostyles found in birds are indeed
synapomorphies.
* †''
Chongmingia zhengi''
* †
Confuciusornithiformes
Confuciusornithidae is an extinct family of pygostylian avialans known from the Early Cretaceous, found in northern China. They are commonly placed as a sister group to Ornithothoraces, a group that contains all extant birds along with their clos ...
** †''
Proornis coreae'' (nomen nudum)
** †''
Zhongornis haoae'' - confuciusornithid?
** †
Confuciusornithidae
Confuciusornithidae is an extinct family of pygostylian avialans known from the Early Cretaceous, found in northern China. They are commonly placed as a sister group to Ornithothoraces, a group that contains all extant birds along with their clo ...
*** †''
Changchengornis hengdaoziensis'' (Early Cretaceous)
*** †''
Eoconfuciusornis zhengi'' (Early Cretaceous)
*** †''
Evgenavis nobilis ''
*** †''
Confuciusornis
''Confuciusornis'' is a genus of basal crow-sized avialan from the Early Cretaceous Period of the Yixian and Jiufotang Formations of China, dating from 125 to 120 million years ago. Like modern birds, ''Confuciusornis'' had a toothless beak, b ...
''
'Jinzhouornis''_Hou_et_al._2002.html" ;"title="Jinzhouornis.html" ;"title="'Jinzhouornis">'Jinzhouornis'' Hou et al. 2002">Jinzhouornis.html" ;"title="'Jinzhouornis">'Jinzhouornis'' Hou et al. 2002(Early Cretaceous)
* †Omnivoropterygiformes [Sapeornithiformes ]
** †Omnivoropterygidae [Sapeornithidae ]
*** †''Omnivoropteryx, Omnivoropteryx sinousaorum'' (Early Cretaceous)
*** †''Sapeornis, Sapeornis chaoyangensis''
'Didactylornis_jii''_;_''Sapeornis_angustis.html" ;"title="Didactylornis_jii.html" ;"title="'Didactylornis jii">'Didactylornis jii'' ; ''Sapeornis angustis">Didactylornis_jii.html" ;"title="'Didactylornis jii">'Didactylornis jii'' ; ''Sapeornis angustis'' ; ''Shenshiornis primita'' ] (Early Cretaceous)
Enantiornithes (extinct)
![Iberomesornis-model-2](https://upload.wikimedia.org/wikipedia/commons/b/bf/Iberomesornis-model-2.jpg)
The taxonomic list of enantiornithine groups presented here follows a summary published by Thomas R. Holz, Jr. in 2011.
†
Enantiornithes
* Basal
Enantiornithes and Enantiornithes ''incerta sedis''
**†''
Cerebavis cenomanica'' (Late Cretaceous)
**†''
Cruralispennia multidonta''
**†''
Dalingheornis liweii'' (Early Cretaceous)
**†''
Dunhuangia cuii''
**†''
Elsornis keni'' (Late Cretaceous)
**†''
Feitianius paradisi''
**†''
Holbotia ponomarenkoi''
'Holbotia ponomarenkoi'' **†''
Houornis caudatus''
'Cathayornis_caudatus''_.html" ;"title="Cathayornis_caudatus.html" ;"title="'Cathayornis caudatus">'Cathayornis caudatus'' ">Cathayornis_caudatus.html" ;"title="'Cathayornis caudatus">'Cathayornis caudatus'' **†Laevisoolithidae [Subtiliolithidae]
otaxa***†''Laevisoolithus, Laevisoolithus sochavai''
otaxa***†''
Subtiliolithus microtuberculatus''
otaxa***†''
Tipoolithus achloujensis''
otaxa**†''
Linyiornis amoena''
**†''
Monoenantiornis sihedangia''
**†''
Paraprotopteryx gracilis'' (Early Cretaceous)
**†''
Parvavis chuxiongensis''
**†''
Piscivorenantiornis inusitatus''
**†''
Pterygornis dapingfangensis''
**†''
Xiangornis shenmi'' (Jiufotang Early Cretaceous of Liaoning)
**†''
Yuanjiawaornis viriosus''
**†
Kurzholiidae
***†''
Kuszholia mengi'' (Late Cretaceous)
**†
Iberomesornithiformes
***†
Iberomesornithidae
****†''
Iberomesornis romerali'' (Early Cretaceous)
**†
Liaoningornithiformes
***†
Liaoningornithidae
****†''
Eoalulavis hoyasi
''Eoalulavis'' (from the Ancient Greek: ''Éōs'', "dawn"; ''alula'', "bastard wing"; ''avis'', "bird") is a monotypic genus of enantiornithean bird that lived during the Barremian, in the Lower Cretaceous around 125 million years ago. The only ...
'' (Early Cretaceous)
****†''
Liaoningornis longidigitris''
* Basal
Euenantiornithes
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and cla ...
**†''
Catenoleimus anachoretus'' (Late Cretaceous)
**†''
Cratoavis cearensis''
**†''
Elbretornis bonapartei'' (Late Cretaceous)
**†''
Eocathayornis walkeri''
'Eocathayornis walkeri'' (Early Cretaceous)
**†''
Flexomornis howei'' (Late Cretaceous)
**†''
Fortunguavis xiaotaizicus''
**†''
Grabauornis lingyuanensis''
**†''
Huoshanornis huji '' (Late Cretaceous)
**†''
Largirostrornis sexdentoris'' (Early Cretaceous)
**†''
Lectavis bretincola'' (Late Cretaceous)
**†''
Martinavis
''Martinavis'' is a genus of enantiornithine birds which existed in what is now southern France, North America and Salta Province, Argentina during the late Cretaceous period. It was named by Cyril A. Walker, Eric Buffetaut and Gareth J. Dyke ...
'' (Late Cretaceous)
***†''
M. cruzyensis ''
***†''
M. vincei ''
***†''
M. minor''
***†''
M. saltariensis''
***†''
M. whetstonei''
**†''
Musivavis amabilis'' (Early Cretaceous)
**†''
Yungavolucris brevipedalis'' Chiappe 1993 (Late Cretaceous)
**†
Liaoxiornithiformes
***†
Liaoxiornithidae
****†''
Liaoxiornis delicatus''
'Lingyuanornis_parvus''_.html" ;"title="Lingyuanornis_parvus.html" ;"title="'Lingyuanornis parvus">'Lingyuanornis parvus'' ">Lingyuanornis_parvus.html" ;"title="'Lingyuanornis parvus">'Lingyuanornis parvus'' (Late Cretaceous)
**†Pengornithidae
***†''Chiappeavis, Chiappeavis magnapremaxillo''
***†''Eopengornis, Eopengornis martini''
***†''Pengornis, Pengornis houi'' (Early Cretaceous)
***†''Parapengornis, Parapengornis eurycaudatus''
**†
Protopterygiformes
***†
Protopterygidae
****†''
Jibeinia luanhera''
'Jibeinia luanhera'' (Early Cretaceous)
****†''
Protopteryx fengningensis
''Protopteryx'' is an extinct bird and the most basal enantiornithean, from the Cretaceous period.Zhou, Zhonghe. (2002). "A new and primitive enantiornithine bird from the Early Cretaceous of China." ''Journal of Vertebrate Paleontology'', 22(1): ...
'' (Early Cretaceous)
****†''
Hebeiornis fengningensis''
*†
Eoenantiornithiformes
**†
Eoenantiornithidae
***†''
Dapingfangornis sentisorhinus'' (Early Cretaceous)
***†''
Eoenantiornis buhleri''
**†
Bohaiornithidae
Bohaiornithidae is a group of early predatory enantiornitheans from the early Cretaceous Period of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125–120 million years ago. B ...
***†''
Beiguornis khinganensis''
***†''
Bohaiornis guoi
''Bohaiornis'' is a genus of enantiornithean birds. Fossils have been found from the Lower Cretaceous Jiufotang Formation of western Liaoning, China. The only known species, ''Bohaiornis guoi'', was named by Dongyu Hu, Li Li, Lianhaim Hou and ...
''
***†''
Longusunguis kurochkini''
***†''
Parabohaiornis martini''
***†''
Shenqiornis mengi'' (Early Cretaceous)
***†''
Sulcavis geeorum'' (Early Cretaceous)
***†''
Zhouornis hani''
*†
Longipterygiformes
Longipterygidae is a family of early enantiornithean avialans from the Early Cretaceous epoch of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125-120 million years ago.
Desc ...
oluochiiformes ; Longirostraviformes **†
Longipterygidae oluochiidae ; Longirostravidae ***†''
Boluochia zhengi'' (Early Cretaceous)
***†''
Camptodontus yangi''
(Early Cretaceous)
***†''
Longipteryx chaoyangensis'' (Early Cretaceous)
***†''
Longirostravis hani'' (Early Cretaceous)
***†''
Otogornis genghisi'' (Early Cretaceous)
***†''
Rapaxavis pani'' (Early Cretaceous)
***†''
Shanweiniao cooperorum'' (Early Cretaceous)
***†''
Shengjingornis yangi'' (Early Cretaceous)
*†
Cathayornithiformes inornithiformes ; Euornithiformes **†
Alethoalaornithidae
***†''
Alethoalaornis agitornis'' (Early Cretaceous)
**†
Concornithidae
***†''
Concornis lacustris'' (Early Cretaceous)
***†''
Qiliania graffini'' (Early Cretaceous)
***†''
Noguerornis gonzalezi'' (Early Cretaceous)
**†
Cathyornithidae inornithidae ***†''
Gracilornis jiufotangensis'' (Early Cretaceous)
***†''
Longchengornis sanyanensis'' (Early Cretaceous)
***†''
Sinornis santensis'' (Early Cretaceous)
***†''
Cathayornis
''Cathayornis'' is a genus of enantiornithean birds from the Jiufotang Formation of Liaoning, People's Republic of China. It is known definitively from only one species, ''Cathayornis yandica'', one of the first Enantiornithes found in China. Se ...
'' (Early Cretaceous)
*†
Enantiornithiformes obipterygiformes ; Alexornithiformes **†
Avisauridae
***†''
Cuspirostrisornis houi'' (Early Cretaceous)
***†''
Avisaurus'' (Late Cretaceous)
***†''
Bauxitornis mindszentyae''
(Late Cretaceous)
***†''
Enantiophoenix electrophyla'' (Late Cretaceous)
***†''
Halimornis thompsonae'' (Late Cretaceous)
***†''
Gettyia
''Gettyia'' is an extinct genus of avisaurid enantiornithean bird from the Late Cretaceous of North America.
Description
''Gettyia'' is known from a single right tarsometatarsus. This bone was similar to that of other North American avisau ...
'' (Late Cretaceous)
***†''
Intiornis inexpectatus''
(Late Cretaceous)
***†''
Mirarce
''Mirarce'' (meaning "wonderful winged messenger") is a genus of enantornithe bird from the Late Cretaceous of Utah. It contains a single species, ''M. eatoni''.Science NewsMirarce eatoni: Newly-Discovered Cretaceous Bird Lived Among Dinosaurs, ...
'' (Late Cretaceous)
***†''
Mystiornis cyrili'' (Early Cretaceous)
***†''
Neuquenornis volans'' (Late Cretaceous)
***†''
Soroavisaurus australis'' (Late Cretaceous)
**†
Gobipterygidae
Gobipterygidae is a family of extinct enantiornithine birds known from the Cretaceous of Asia.Holtz, Thomas R. Jr. (2012) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2011 Appendix./ref>
Referen ...
***†''
Gobipipus reshetovi''
***†''
Gobipteryx minuta
''Gobipteryx'' (from Gobi eferring to the Gobi Desert where it was first discovered and Greek pteryx “wing”) is a genus of prehistoric bird from the Campanian Age of the Late Cretaceous Period.Elżanowski, A. (1974): Preliminary note on th ...
'' (Late Cretaceous)
***†''
Vescornis hebeiensis'' (Early Cretaceous)
**†
Enantiornithidae
***†''
Enantiornis leali'' (Late Cretaceous)
***†''
Gurilynia nessovi'' (Late Cretaceous)
***†''
Wyleyia valdensis'' (Late Cretaceous)
***†''
Nanantius'' (Early-?Late Cretaceous)
**†
Alexornithidae
***†''
Abavornis bonaparti'' (Late Cretaceous)
***†''
Alexornis antecedens
''Alexornis'' is a genus of enantiornithine birds from the Bocana Roja Formation of Baja California, Mexico.Tykoski, R. S., & Fiorillo, A. R. (2010). An enantiornithine bird from the lower middle Cenomanian of Texas. ''Journal of Vertebrate Pal ...
'' (Late Cretaceous)
***†''
Explorornis
''Explorornis'' ("discovered bird", from Latin ''explōrō'' "to discover" and Ancient Greek ''ornis'' (''όρνις'') "bird") is a genus of Mesozoic birds which lived during the mid-late Turonian stage, around 90 million years ago, in the Biss ...
'' (Late Cretaceous)
***†''
Incolornis'' (Late Cretaceous)
***†''
Kizylkumavis cretacea'' (Late Cretaceous)
***†''
Lenesornis maltschevskyi''
' 'Ichthyornis_maltschevskyi''_">Ichthyornis_maltschevskyi.html"_;"title="'Ichthyornis_maltschevskyi">'Ichthyornis_maltschevskyi''_(Late_Cretaceous)
***†''Sazavis.html" ;"title="Ichthyornis_maltschevskyi''_.html" ;"title="Ichthyornis_maltschevskyi.html" ;"title="'Ichthyornis maltschevskyi">'Ichthyornis maltschevskyi'' ">Ichthyornis_maltschevskyi.html" ;"title="'Ichthyornis maltschevskyi">'Ichthyornis maltschevskyi'' (Late Cretaceous)
***†''Sazavis">Sazavis prisca'' (Early Cretaceous)
***†''Zhyraornis'' (Late Cretaceous)
Note that Holtz (2011) also included ''Zhyraornis'' in his classification of euenantiornithines, though this genus is more often classified as an ornithurae, ornithuran.
[Kurochkin (2006)] Holtz also placed ''
Liaoningornis'' as an
ornithuromorph
Euornithes (from Greek ' meaning "true birds") is a natural group which includes the most recent common ancestor of all avialans closer to modern birds than to ''Sinornis''.
Description
Clarke ''et al''. (2006) found that the most primitive know ...
, though more recent studies have placed it as a close relative of ''Eoalulavis''.
[O’Connor, J.K. (2012)]
Basal
Euornithes
Euornithes (from Greek ' meaning "true birds") is a natural group which includes the most recent common ancestor of all avialans closer to modern birds than to '' Sinornis''.
Description
Clarke ''et al''. (2006) found that the most primitive kn ...
(extinct)
Also called "basal
Ornithuromorpha
Euornithes (from Greek ' meaning "true birds") is a natural group which includes the most recent common ancestor of all avialans closer to modern birds than to ''Sinornis''.
Description
Clarke ''et al''. (2006) found that the most primitive know ...
". Essentially modern birds, except many still possess a few primitive features such as teeth or wing claws. These have the
plowshare
In agriculture, a plowshare ( US) or ploughshare ( UK; ) is a component of a plow (or plough). It is the cutting or leading edge of a moldboard which closely follows the coulter (one or more ground-breaking spikes) when plowing.
The plowsh ...
-shaped pygostyle and proper tail fan as seen in most living birds. The
taxonomy
Taxonomy is the practice and science of categorization or classification.
A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
of this group is confusing; the name "Ornithurae" was first proposed by
Ernst Haeckel
Ernst Heinrich Philipp August Haeckel (; 16 February 1834 – 9 August 1919) was a German zoologist, naturalist, eugenicist, philosopher, physician, professor, marine biologist and artist. He discovered, described and named thousands of new sp ...
in 1866 and has been revised in meaning several times since.
The following is a list of primitive euornithian genera and those that cannot be confidently referred to any subgroups, following Holtz (2011).
[Winter 2011 Appendix.]
/ref>
*†'' Archaeorhynchus spathula''
*†'' Changmaornis houi''
*†'' Changzuiornis ahgmi''
*†'' Gargantuavis philoinos''
*†''Hollanda luceria
''Hollanda'' is a genus of small ground birds known from fossils found in the Barun Goyot Formation of Mongolia. Found at Khermeen Tsav, it dates from the late Cretaceous period (Campanian stage), about 75 million years ago. Known only from part ...
'' ' 'Hollandornis_birdus''_">Hollandornis_birdus.html"_;"title="'Hollandornis_birdus">'Hollandornis_birdus''_*†'' ''_.html"_;"title="Hollandornis_birdus.html"_;"title="'Hollandornis_birdus">'Hollandornis_birdus''_">Hollandornis_birdus.html"_;"title="'Hollandornis_birdus">'Hollandornis_birdus''_*†''Horezmavis">Horezmavis_eocretaceous''_
*†''Hulsanpes.html" ;"title="Horezmavis.html" ;"title="Hollandornis_birdus''_.html" ;"title="Hollandornis_birdus.html" ;"title="'Hollandornis birdus">'Hollandornis birdus'' ">Hollandornis_birdus.html" ;"title="'Hollandornis birdus">'Hollandornis birdus'' *†''Horezmavis">Horezmavis eocretaceous''
*†''Hulsanpes">Hulsanpes perlei'' (?Troodontidae)
*†''Iteravis, Iteravis huchzermeyeri''
*†''Jianchangornis, Jianchangornis microdonta''
*†''Jiuquanornis, Jiuquanornis niui''
*†''Juehuaornis, Juehuaornis zhangi''
*†''Palaeopteryx thomsoni'' (dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
)
*†'' Platanavis nana''
*†'' Schizooura lii''
*†'' Tingmiatornis arctica''
*†'' Vorona berivotrensis''
*†'' Xinghaiornis lini''
*†'' Yumenornis huangi''
*†'' Zhongjianornis yangi''
*† Patagopterygidae
**†'' Alamitornis minutus''
**†'' Patagopteryx deferrariisi''
*† Hongshanornithidae
**?†'' Archaeornithura meemannae''
**†''Hongshanornis longicresta
''Hongshanornis'' is a genus of ornithuromorph birds known from early Cretaceous lake deposits of the Yixian Formation, Inner Mongolia, China. The holotype specimen, recovered in 2005, is currently held by the Institute of Vertebrate Paleontolog ...
'' (Early Cretaceous)
**†'' Longicrusavis houi'' (Early Cretaceous)
**†'' Parahongshanornis chaoyangensis'' (Early Cretaceous)– ornithurine?
**†'' Tianyuornis cheni''
*† Chaoyangiiformes
**† Chaoyangiidae haoyangornithidae ***†'' Chaoyangia beishanensis''
*† Songlingornithiformes berratiodontiformes ; Yanornithiformes ; Yixianornithiformes **†Songlingornithidae
Songlingornithidae is a family of basal euornitheans from the Early Cretaceous of China. All known specimens come from the Jiufotang Formation and the Yixian Formation, dating to the early Barremian and Aptian ages, 125–120 million years ag ...
berratiodontidae ; Chaoyangornithidae ; Yanornithidae ; Yixiaornithidae ***†'' Piscivoravis lii''
***†'' Songlingornis linghensis'' (Early Cretaceous)
***†'' Yixianornis grabaui'' (Early Cretaceous)
***†''Yanornis
''Yanornis'' () is an extinct genus of fish-eating Early Cretaceous birds. Two species have been described, both from Liaoning province, China: ''Yanornis martini'', based on several fossils found in the 120-million-year-old Jiufotang Formati ...
'' 'Archaeoraptor''_;_''Archaeovolans.html" ;"title="Archaeoraptor.html" ;"title="'Archaeoraptor">'Archaeoraptor'' ; ''Archaeovolans">Archaeoraptor.html" ;"title="'Archaeoraptor">'Archaeoraptor'' ; ''Archaeovolans'' ; ''Aberratiodontus'' ] (Early Cretaceous)
Note that Holtz also included the genera ''Eurolimnornis'', ''Holbotia'', ''Palaeocursornis'' and ''Piksi'' as euornitheans, though they have since been re-identified as pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 ...
s.
Basal Ornithurae (extinct)
* '' Limenavis patagonica'' (Late Cretaceous)– paleognath?
* Palintropidae
** ''Palintropus retusus
''Palintropus'' is a prehistoric bird genus from the Late Cretaceous. A single species has been named (''Palintropus retusus'') based on a proximal coracoid from the Lance Formation of Wyoming, dated to the latest Maastrichtian, million years ag ...
'' 'Cimolopteryx_retusa''_Marsh_1892;_''Apatornis_retusus.html" ;"title="Cimolopteryx_retusa.html" ;"title="'Cimolopteryx retusa">'Cimolopteryx retusa'' Marsh 1892; ''Apatornis retusus">Cimolopteryx_retusa.html" ;"title="'Cimolopteryx retusa">'Cimolopteryx retusa'' Marsh 1892; ''Apatornis retusus'' (Marsh 1892) Brodkorb 1962] (Late Cretaceous) (?Galliformes: Quercymegapodiidae)
*†Ambiortiformes
**†Ambiortidae
***†''Ambiortus, Ambiortus dementjevi'' (Late Cretaceous)
*†Apsaraviformes
**† Apsaravidae
***†'' Apsaravis ukhaana'' (Late Cretaceous)
*† Gansuiformes
**† Gansuidae
***†''Gansus
''Gansus'' is a genus of aquatic birds that lived during the Aptian age of the Early Cretaceous (Aptian-Albian) period in what are now Gansu and Liaoning provinces, western China. The rock layers from which their fossils have been recovered are d ...
'' (Early Cretaceous)
*†Ichthyornithiformes
Ichthyornithes is an extinct group of toothed avialans very closely related to the common ancestor of all modern birds. They are known from fossil remains found throughout the late Cretaceous period of North America, though only two genera, ''Ich ...
teropappi **† Apatornithidae
***†'' Apatornis celer'' ' 'Ichthyornis_celer''_Marsh_1873">Ichthyornis_celer.html"_;"title="'Ichthyornis_celer">'Ichthyornis_celer''_Marsh_1873(Late_Cretaceous)
***†''Guildavis.html" ;"title="Ichthyornis_celer''_Marsh_1873.html" ;"title="Ichthyornis_celer.html" ;"title="'Ichthyornis celer">'Ichthyornis celer'' Marsh 1873">Ichthyornis_celer.html" ;"title="'Ichthyornis celer">'Ichthyornis celer'' Marsh 1873(Late Cretaceous)
***†''Guildavis">Guildavis tener'' [''Ichthyornis tener'' Marsh 1880] (Cretaceous of Wallace County, US)
***†''Iaceornis, Iaceornis marshi'' (Late Cretaceous)
**†Ichthyornithidae
*** ''Ichthyornis'' [''Angelinornis'' ; ''Colonosaurus'' ; ''Graculavus
''Graculavus'' is a prehistoric bird genus that was described in the 19th century by American paleontologist O. C. Marsh. Its remains were found in the Late Cretaceous Austin Chalk of Texas, USA, and Lance Formation (late Maastrichtian faunal sta ...
'' ; '' Plegadornis'' ] (Late Cretaceous)
*† Hesperornithiformes (Large, toothed, loon-like diving birds)
**†'' Chupkaornis keraorum''
**†'' Potamornis skutchi'' (Late Cretaceous)
**†''Pasquiaornis
''Pasquiaornis'' is a prehistoric flightless bird genus from the Late Cretaceous. It lived during the late Cenomanian, between 95 and 93 million years ago in North America.Tokaryk, Cumbaa and Storer, 1997. Early Late Cretaceous birds from Saskat ...
'' (Late Cretaceous)
**†Enaliornithidae
''Enaliornis'' is a genus of hesperornithine birds which lived in the early Late Cretaceous, making them the oldest known hesperornithines. Fossils have been found near Cambridge, England. Due to its lack of certain hesperornithid apomorphies, ...
***†''Enaliornis
''Enaliornis'' is a genus of hesperornithine birds which lived in the early Late Cretaceous, making them the oldest known hesperornithines. Fossils have been found near Cambridge, England. Due to its lack of certain hesperornithid apomorphies, ...
'' 'Palaeocolyntus''_;_''Palaeocolymbus''_;_''Pelargonis''_;_''Pelagornis.html" ;"title="Palaeocolymbus.html" ;"title="'Palaeocolyntus'' ; ''Palaeocolymbus">'Palaeocolyntus'' ; ''Palaeocolymbus'' ; ''Pelargonis'' ; ''Pelagornis">Palaeocolymbus.html" ;"title="'Palaeocolyntus'' ; ''Palaeocolymbus">'Palaeocolyntus'' ; ''Palaeocolymbus'' ; ''Pelargonis'' ; ''Pelagornis'' ] (Early Cretaceous)
**†Baptornithidae
***†Baptornis [''Parascaniornis'' ]
**†Brodavidae
***''Brodavis'' (Late Cretaceous)
**†Judinornithidae
***†'' Judinornis nogontsavensis'' (Late Cretaceous)
**† Hesperornithidae
***†'' Fumicollis hoffmani''
***†'' Parahesperornis alexi'' (Late Cretaceous)
***†''Hesperornis
''Hesperornis'' (meaning "western bird") is a genus of cormorant-like bird that spanned the first half of the Campanian age of the Late Cretaceous period (83.5–78 mya). One of the lesser-known discoveries of the paleontologist O. C. Marsh i ...
'' 'Coniornis''_;_''Lestornis.html" ;"title="Coniornis.html" ;"title="'Coniornis">'Coniornis'' ; ''Lestornis">Coniornis.html" ;"title="'Coniornis">'Coniornis'' ; ''Lestornis'' ; ''Hargeria'' ; ''Asiahesperornis'' ; ''Canadaga'' ] (Late Cretaceous)
Neornithes
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
The subclass that contains all modern birds.
Unresolved and basal forms
These modern birds are known from remains that cannot be placed in relation to any one modern group and are neither autapomorph
In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...
ic enough to assign them to own orders. Especially the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
/early Paleogene
The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...
taxa
In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
are probably basal to several modern orders, while later Paleogene
The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...
taxa often represent extinct lineages outside the modern families.
*†'' Australornis lovei'' (late early Paleocene
The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
)
*†'' Gallornis straeleni'' (Late Cretaceous)
*†'' Ceramornis major'' (Late Cretaceous) – charadriiform?
*†"Presbyornithidae" gen. et sp. indet. (Barun Goyot Late Cretaceous of Udan Sayr, Mongolia) - anseriform (presbyornithid)?
*†''Torotix clemensi
''Torotix'' is a Late Cretaceous genus of aquatic birds. They lived along the shores of the Western Interior Seaway, but it is not clear whether they were seabirds or freshwater birds, as the genus is only known from a humerus. Consequently, t ...
'' (Late Cretaceous) – pelecaniform, charadriiform, procellariiform or phoenicopteriform
*Neornithes incerta sedis (Nemegt Late Cretaceous of S Mongolia) - phalacrocoracid?
*Neornithes incerta sedis AMNH FR 25272 (Lance Creek Late Cretaceous of Converse County, US) - phalacrocoracid?
*Neornithes incerta sedis PVPH 237 (Portezuelo Late Cretaceous of Sierra de Portezuelo, Argentina) - galliform?
*Neornithes incerta sedis UCMP 117598 (Hell Creek Late Cretaceous of Bug Creek West, US)
*Neornithes incerta sedis UCMP 117599 (Hell Creek Late Cretaceous of Bug Creek West, US) – anseriform?
*†''"Lonchodytes" pterygius'' (Late Cretaceous/?Early Palaeocene) – charadriiform?
*†'' Novacaesareala hungerfordi'' (Late Cretaceous/Early Palaeocene) – related to ''Torotix''?
*†''"Palaeotringa" vetus'' (Lance Late Cretaceous of Wyoming – Hornerstown Late Cretaceous/?Early Palaeocene of New Jersey, US) - gruiform? anseriform (presbyornithid)?
*†'' Volgavis marina'' (Early Palaeocene of Volgograd, Russia) – charadriiform? phalacrocoraciform?
*†'' Tshulia litorea'' (Late Paleocene of Zhylga, Kazakhstan)
*†'' Eupterornis remensis'' (Paleocene of Cernay, France) – charadriiform (larid?)? gaviiform?
*†'' Gradiornis walbeckensis'' (Paleocene of Walbeck, Germany) – cariamid?
*†''" Messelornis" russelli'' (Paleocene of Cernay, France) – messelornithid?
*†'' Walbeckornis creber'' (Paleocene of Walbeck, Germany) – charadriiform? messelornithid?
*Neornithes incerta sedis (Late Paleocene/Early Eocene of Ouled Abdoun Basin, Morocco) – charadriiform, ciconiiform, gruiform?
*†'' Argillipes'' (London Clay Early Eocene of England) – galliform?
**†'' A. aurorum''
**†'' A. magnus''
**†'' A. paralectoris''
*†'' Coturnipes cooperi'' (Early Eocene of England, and Virginia, US?)– galliform, falconiform?
*†'' Fluviatilavis antunesi'' (Early Eocene of Silveirinha, Portugal)– charadriiform?
*†'' Mopsitta tanta'' (Early Eocene) – threskiornithid (may belong in ''Rhynchaeites''), psittacid?
*†''Neanis schucherti
''Neanis'' is an extinct genus of bird probably related to woodpeckers and toucans. It contains at least one species, ''N. schucherti''; ''N. kistneri'' resembles this, but it probably belongs to a distinct genus and may not be closely related. B ...
'' (Early Eocene)– coraciiform (primobucconid), piciform?
*†'' Paleophasianus meleagroides'' (Willwood Early Eocene of Bighorn County, US)– galliform (tetraonine or cracid) or gruiform (aramid)?
*†'' Precursor parvus'' (Early Eocene)– several species? psittaciform (pseudasturid or psittacid) + charadriiform (glareolid)?
*†'' "Precursor" litorum''
*†'' "Precursor" magnus''
*†'' Procuculus minutus'' (Early Eocene of Bognor Regis, England)– cuculiform (parvicuculid), coraciiform (primobucconid), close to ''Primapus''?
*†'' Pulchrapollia'' (Early Eocene) – psittaciform (pseudasturid or psittacid)?
**†'' P. gracilis''
**†'' P. olsoni'' ' 'Primobucco_olsoni''_">Primobucco_olsoni.html"_;"title="'Primobucco_olsoni">'Primobucco_olsoni''_*Neornithes_incerta_sedis_USNM_496384_(Nanjemoy_Early_Eocene_of_Virginia,_US)–_parvicuculid?_aegithalornithid?
*†''[ ''_.html"_;"title="Primobucco_olsoni.html"_;"title="'Primobucco_olsoni">'Primobucco_olsoni''_">Primobucco_olsoni.html"_;"title="'Primobucco_olsoni">'Primobucco_olsoni''_*Neornithes_incerta_sedis_USNM_496384_(Nanjemoy_Early_Eocene_of_Virginia,_US)–_parvicuculid?_aegithalornithid?
*†''Palaeopsittacus">Palaeopsittacus_georgei''__(Early_–_middle_Eocene_of_NW_Europe)_–_caprimulgiform_(podargid?)_or_quercypsittid?
*†''Amitabha_(bird).html" ;"title="alaeopsittacus.html" ;"title="Primobucco_olsoni''_.html" ;"title="Primobucco_olsoni.html" ;"title="' 'Primobucco_olsoni''_">Primobucco_olsoni.html"_;"title="'Primobucco_olsoni">'Primobucco_olsoni''_*Neornithes_incerta_sedis_USNM_496384_(Nanjemoy_Early_Eocene_of_Virginia,_US)–_parvicuculid?_aegithalornithid?
*†''Palaeopsittacus">Palaeopsittacus_georgei''__(Early_–_middle_Eocene_of_NW_Europe)_–_caprimulgiform_(podargid?)_or_quercypsittid?
*†''Amitabha_(bird)">Amitabha_urbsinterdictensis''__(Bridger_middle_Eocene_of_Forbidden_City,_US)_-_galliform_(phasianid)_or_gruiform_(rallid?)?
*†'' 'Primobucco_olsoni''_">Primobucco_olsoni.html"_;"title="'Primobucco_olsoni">'Primobucco_olsoni''_*Neornithes_incerta_sedis_USNM_496384_(Nanjemoy_Early_Eocene_of_Virginia,_US)–_parvicuculid?_aegithalornithid?
*†''Palaeopsittacus">Palaeopsittacus_georgei''__(Early_–_middle_Eocene_of_NW_Europe)_–_caprimulgiform_(podargid?)_or_quercypsittid?
*†''Amitabha_(bird)">Amitabha_urbsinterdictensis''__(Bridger_middle_Eocene_of_Forbidden_City,_US)_-_galliform_(phasianid)_or_gruiform_(rallid?)?
*†''Eociconia">Eociconia_sangequanensis''__(middle_Eocene_of_China)–_ciconiiform_(ciconiid)?
*†''Protocypselomorphus.html" ;"title="Eociconia.html" ;"title="Primobucco olsoni">' Primobucco_olsoni.html"_;"title="'Primobucco_olsoni">'Primobucco_olsoni''_*Neornithes_incerta_sedis_USNM_496384_(Nanjemoy_Early_Eocene_of_Virginia,_US)–_parvicuculid?_aegithalornithid?
*†''Palaeopsittacus">Palaeopsittacus_georgei''__(Early_–_middle_Eocene_of_NW_Europe)_–_caprimulgiform_(podargid?)_or_quercypsittid?
*†''Amitabha_(bird)">Amitabha_urbsinterdictensis''__(Bridger_middle_Eocene_of_Forbidden_City,_US)_-_galliform_(phasianid)_or_gruiform_(rallid?)?
*†''Eociconia">Eociconia_sangequanensis''__(middle_Eocene_of_China)–_ciconiiform_(ciconiid)?
*†''Protocypselomorphus">Protocypselomorphus_manfredkelleri''__(middle_Eocene_of_Messel,_Germany)–_caprimulgiform,_apodiform_or_ancestral_to_both
*†'' ''_">Primobucco_olsoni.html"_;"title="'Primobucco_olsoni">'Primobucco_olsoni''_*Neornithes_incerta_sedis_USNM_496384_(Nanjemoy_Early_Eocene_of_Virginia,_US)–_parvicuculid?_aegithalornithid?
*†''Palaeopsittacus">Palaeopsittacus_georgei''__(Early_–_middle_Eocene_of_NW_Europe)_–_caprimulgiform_(podargid?)_or_quercypsittid?
*†''Amitabha_(bird)">Amitabha_urbsinterdictensis''__(Bridger_middle_Eocene_of_Forbidden_City,_US)_-_galliform_(phasianid)_or_gruiform_(rallid?)?
*†''Eociconia">Eociconia_sangequanensis''__(middle_Eocene_of_China)–_ciconiiform_(ciconiid)?
*†''Protocypselomorphus">Protocypselomorphus_manfredkelleri''__(middle_Eocene_of_Messel,_Germany)–_caprimulgiform,_apodiform_or_ancestral_to_both
*†''Pumiliornis">Pumiliornis_tessellatus''__(middle_Eocene_of_Messel,_Germany)
*†''Ludiortyx.html" ;"title="Pumiliornis.html" ;"title="Primobucco olsoni'' ">Primobucco_olsoni.html" ;"title="'Primobucco olsoni">'Primobucco olsoni'' *Neornithes incerta sedis USNM 496384 (Nanjemoy Early Eocene of Virginia, US)– parvicuculid? aegithalornithid?
*†''Palaeopsittacus">Palaeopsittacus georgei'' (Early – middle Eocene of NW Europe) – caprimulgiform (podargid?) or quercypsittid?
*†''Amitabha (bird)">Amitabha urbsinterdictensis'' (Bridger middle Eocene of Forbidden City, US) - galliform (phasianid) or gruiform (rallid?)?
*†''Eociconia">Eociconia sangequanensis'' (middle Eocene of China)– ciconiiform (ciconiid)?
*†''Protocypselomorphus">Protocypselomorphus manfredkelleri'' (middle Eocene of Messel, Germany)– caprimulgiform, apodiform or ancestral to both
*†''Pumiliornis">Pumiliornis tessellatus'' (middle Eocene of Messel, Germany)
*†''Ludiortyx">Ludiortyx hoffmanni'' (Late Eocene) – rallid, quercymegapodid? [Includes ''Tringa hoffmanni'' ; ''Palaeortyx blanchardi'' ; ''Palaeortyx hoffmanni'' ; ''Ludiortyx blanchardi'' ; ''Eortyx hoffmanni'' ]
*†''Minggangia, Minggangia changgouensis'' (Late Eocene of China) – rallid, threskiornithid?
*†'' Petropluvialis simplex'' (Late Eocene of England)– may be same as ''Palaeopapia''; anseriform?
*†''"Phasianus" alfhildae'' (Washakie B Late Eocene of Haystack Butte, US)– gruiform, ciconiiform, phoenicopteriform?
*†'' Telecrex grangeri'' (Irdin Manha Late Eocene of Chimney Butte, Mongolia)– meleagrid or gruiform (rallid?)
*Neornithes incerta sedis AMNH FR 2941 (Irdin Manha Late Eocene of Chimney Butte, China) – falconiform (accipitrid)? gruiform (''Eogrus'')?
*†'' Zheroia kurochkini'' (Late Eocene of Kazakhstan) – gruiform? pelagornithid?
*†''"Falco" falconellus'' (or ''falconella''; Eocene of Wyoming, US) – falconiform (falconid)?
*†''Agnopterus
''Agnopterus'' is an extinct genus of stem-flamingo phoenicopteriform with fossil material from France, as well as possibly England, Kazakhstan, and Brazil. The holotype specimen for type species ''A. laurillardi'' is an incomplete distal ...
'' (Late Eocene– Late Oligocene of Europe)– phoenicopteriform or anseriform
**†'' A. laurillardi''
**†'' A. sicki''
**†'' A. turgaiensis'' 'Cygnopterus_lambrechti''_.html" ;"title="Cygnopterus_lambrechti.html" ;"title="'Cygnopterus lambrechti">'Cygnopterus lambrechti'' ">Cygnopterus_lambrechti.html" ;"title="'Cygnopterus lambrechti">'Cygnopterus lambrechti'' *†''Plesiocathartes'' (Late Eocene–? Early Miocene of SW Europe) - cathartid, leptosomid?
**†''Plesiocathartes europaeus, P. europaeus''
**†''Plesiocathartes gaillardi, P. gaillardi''
**†''Plesiocathartes wyomingensis, P. wyomingensis''
**†'' Plesiocathartes major, P. major''
**†'' P. geiselensis''
**†'' P. kelleri''
*†'' Botauroides parvus'' (Eocene of Wyoming, US) – coliiform?
*†'' Aminornis excavatus'' (Deseado Early Oligocene of Rio Deseado, Argentina) – gruiform (aramid)?
*†'' Ciconiopsis antarctica'' (Deseado Early Oligocene of Patagonia, Argentina) – ciconiiform (ciconiid)?
*†'' Climacarthrus incompletus'' (Deseado Early Oligocene of Argentina) – falconiform (accipitrid)? A '' nomen dubium''
*†'' Cruschedula revola'' (Deseado Early Oligocene of Golfo San Jorge, Argentina) Aves ''incertae sedis''; A '' nomen dubium''
*†'' Dolichopterus viator'' 'Dolicopterus''_;_''Camaskelus.html" ;"title="Dolicopterus.html" ;"title="'Dolicopterus">'Dolicopterus'' ; ''Camaskelus">Dolicopterus.html" ;"title="'Dolicopterus">'Dolicopterus'' ; ''Camaskelus'' ; ''Camaskelus'' ; ''Dolicopterus viator'' ; ''Camaskelus palustris'' ; ''Camaskelus palustris'' ] (Early Oligocene of Ronzon, France) – charadriiform (charadriid)? Not ''Dolicopterus'' as sometimes claimed
*†'' Loncornis erectus'' (Deseado Early Oligocene of Rio Deseado, Argentina)– gruiform (aramid)?
*†'' Loxornis clivus'' – anatid? (Deseado Early Oligocene of Argentina)
*†'' Manu antiquus'' (Early Oligocene) – pelagornithid? procellariiform (diomedeid)?
*†'' Palaeocrex rex'' (Early Oligocene of Trigonias Quarry, US) – gruiform (rallid)?
*†'' Palaeopapia eous'' 'Howardia''_;_''Howardia_eous.html" ;"title="Howardia.html" ;"title="'Howardia">'Howardia'' ; ''Howardia eous">Howardia.html" ;"title="'Howardia">'Howardia'' ; ''Howardia eous'' ] (Hampstead Early Oligocene of Isle of Wight, England) – anseriform?
*†''Paracygnopterus, Paracygnopterus scotti'' (Early Oligocene of Belgium and England) – anseriform (anatid)?
*†''"Pararallus" hassenkampi'' (Sieblos Dysodil Early Oligocene of Sieblos, Germany)
*†'' Riacama caliginea'' (Deseado Early Oligocene of Argentina) – gruiform?
*†'' Smiliornis penetrans'' (Deseado Early Oligocene of Argentina)– gruiform?
*†'' Teracus littoralis'' 'leracus''_Aymard_1856_;_''leracus_littoralis.html" ;"title="leracus.html" ;"title="'leracus">'leracus'' Aymard 1856 ; ''leracus littoralis">leracus.html" ;"title="'leracus">'leracus'' Aymard 1856 ; ''leracus littoralis'' ] (Early Oligocene of France)
*†''Teleornis impressus'' (Deseado Early Oligocene of Argentina)– anatid?
*†''Pseudolarus, Pseudolarus guaraniticus'' (Deseado Early Oligocene – Miocene of Argentina) – gruiform?
*Neornithes incerta sedis BMNH PAL 4989 (Hampstead Early Oligocene of Isle of Wight, England) – formerly ''" Ptenornis"'' and included in '' Headonornis''; anseriform?
*†''"Anas" creccoides'' (Early-mid Oligocene of Belgium) – anseriform?
*†''"Charadrius" sheppardianus'' (Florissant middle Oligocene of Florissant, US) – charadriiform (charadriid?)
*†'' Megagallinula harundinea'' (Indricotherium middle Oligocene of Chelkar-Teniz, Kazakhstan)
*†''"Palaeorallus" alienus'' (middle Oligocene of Tatal-Gol, Mongolia)– galliform?
*†''"Vanellus" selysii'' (middle Oligocene of Rupelmonde, Belgium) – charadriiform (charadriid)?
*†'' Anserpica kiliani'' (Late Oligocene of France) – gruiform (gruid?) or anseriform (anseranatid?)?
*†'' Gnotornis'' (Brule Late Oligocene of Shannon County, US) – gruiform (aramid)?
**†'' G. aramiellus''
**†'' G. roardeola''
**†'' G. walkeri''
*†'' Guguschia nailiae'' (Late Oligocene of Pirəkəşkül, Azerbaijan) – anseriform (anserine)? pelagornithid (same as ''Caspiodontornis'')?
*†'' Tiliornis senex'' (Late? Oligocene of Argentina)– phoenicopteriform? A '' nomen dubium''
*Neornithes incerta sedis QM F40203 (Late Oligocene of Riversleigh, Australia)– gruiform (rallid)?
*†'' Gaviella pusilla'' (Oligocene? of Wyoming, US)– gaviiform? plotopterid?
*†''"Anas" skalicensis'' (Early Miocene of "Skalitz", Czech Republic) - anseriform?
*†'' Chenornis graculoides'' (Early Miocene) - Anseriformes (Anatidae) or Pelecaniformes (Phalacrocoracidae)?
*†''"Propelargus" olseni'' (Hawthorne Early Miocene of Tallahassee, US) – ciconiiform?
*Neornithes incerta sedis MNHN SA 1259-1263 (Early/Middle Miocene of Sansan, France) – passeriform?
*†'' Anisolornis excavatus'' (Santa Cruz Middle Miocene of Karaihen, Argentina) – gruiform, galliform, tinamiform?
*†''"Ardea" perplexa'' (Middle Miocene of Sansan, France) – ardeid? strigiform?
*†''"Cygnus herrenthalsi"'' (Middle Miocene of Belgium)
*†''"Anas" risgoviensis'' (Late Miocene of Bavaria, Germany) – anseriform?
*†''"Ardea" aureliensis'' (Late Miocene of France) – ardeid?
*†'' Eoneornis australis'' (Miocene of Argentina) – anatid? A '' nomen dubium''
*†'' Eutelornis patagonica'' (Miocene of Argentina) – anatid?
*†'' Protibis cnemialis'' (Miocene of Argentina) – ciconiiform (threskiornithid)?
*†''"Limnatornis" paludicola'' (Miocene of France) – coliid? phoeniculid?
*†''"Picus" gaudryi'' (Miocene of France) – piciform?
*†''"Ardea" lignitum'' (Late Pliocene of Germany) – ardeid? strigid (genus ''Bubo'')?
*†''Bathoceleus hyphalus
''Bathoceleus hyphalus'' is an extinct bird discovered in the Bahamas. It is believed to be a primitive woodpecker
Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of th ...
'' (Pliocene of New Providence, Bahamas) – picid?
*†''" Homalopus"'' – piciform? Preoccupied by a subgenus
In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus.
In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
of '' Cryptocephalus'' leaf beetle
The insects of the beetle family Chrysomelidae are commonly known as leaf beetles, and include over 37,000 (and probably at least 50,000) species in more than 2,500 genera, making up one of the largest and most commonly encountered of all beetle ...
s described in 1835.
*†''" Liptornis hesternus"'' ' 'Liptornis_cuvierii''">Liptornis_cuvierii.html"_;"title="'Liptornis_cuvierii">'Liptornis_cuvierii''–_pelecaniform_(pelecanid)?_A_''_nomen_dubium''
*†'' ''.html"_;"title="Liptornis_cuvierii.html"_;"title="'Liptornis_cuvierii">'Liptornis_cuvierii''">Liptornis_cuvierii.html"_;"title="'Liptornis_cuvierii">'Liptornis_cuvierii''–_pelecaniform_(pelecanid)?_A_''_nomen_dubium''
*†''Proceriavis">Proceriavis_martini''__–_pelagornithid?
*†'' ''.html"_;"title="Liptornis_cuvierii.html"_;"title="'Liptornis_cuvierii">'Liptornis_cuvierii''">Liptornis_cuvierii.html"_;"title="'Liptornis_cuvierii">'Liptornis_cuvierii''–_pelecaniform_(pelecanid)?_A_''_nomen_dubium''
*†''Proceriavis">Proceriavis_martini''__–_pelagornithid?
*†''Protopelicanus">Protopelicanus_cuvieri''__–_pelecaniform_(pelecanid)?_pelagornithid?
*†''Eurofluvioviridavis.html" ;"title="Protopelicanus.html" ;"title="Proceriavis.html" ;"title="Liptornis_cuvierii''.html" ;"title="Liptornis_cuvierii.html" ;"title="'Liptornis cuvierii">'Liptornis cuvierii''">Liptornis_cuvierii.html" ;"title="'Liptornis cuvierii">'Liptornis cuvierii''– pelecaniform (pelecanid)? A '' nomen dubium''
*†''Proceriavis">Proceriavis martini'' – pelagornithid?
*†''Protopelicanus">Protopelicanus cuvieri'' – pelecaniform (pelecanid)? pelagornithid?
*†''Eurofluvioviridavis">Eurofluvioviridavis robustipes'' (middle Eocene of Messel, Germany)
*†Archaeotrogonidae – basal Cypselomorphae?
**†Archaeotrogonidae gen. et sp. indet. (Early Eocene)
**†''[ Hassiavis laticauda''
**†'' Archaeotrogon'' (Late Eocene/Early Oligocene)
*†Cimolopterygidae
Cimolopterygidae is an extinct family of ornithurines known from the Late Cretaceous epoch. Remains attributed to cimolopterygids have been found in the Frenchman Formation of Saskatchewan, the Lance Formation of Wyoming, the Fox Hills Formation ...
– charadriiform?
**†''Cimolopteryx
''Cimolopteryx'' (meaning "Cretaceous wing"Holtz, Thomas R. Jr. (2008) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages'Winter 2010 Supplementary Information) is a prehistoric bird genus from the Late Creta ...
'' (Late Cretaceous)
***†'' C. rara''
***†'' C. maxima''
**†''Lamarqueavis
''Lamarqueavis'' is an extinct genus of prehistoric birds from the Cimolopterygidae known from Late Cretaceous-aged rocks from Argentina, Canada, and the United States. The type species, ''L. australis'', was named in 2010 and is based on the ho ...
'' (Late Cretaceous)
***†'' L. australis''
***†'' L. minima'' ' 'Cimolopteryx_minima''_">Cimolopteryx_minima.html"_;"title="'Cimolopteryx_minima">'Cimolopteryx_minima''_***†''Lamarqueavis_petra.html" ;"title="Cimolopteryx_minima''_.html" ;"title="Cimolopteryx_minima.html" ;"title="'Cimolopteryx minima">'Cimolopteryx minima'' ">Cimolopteryx_minima.html" ;"title="'Cimolopteryx minima">'Cimolopteryx minima'' ***†''Lamarqueavis petra">L. petra'' [''Cimolopteryx petra'' ]
*†Cladornithidae Wetmore 1930 [Cladornithes ; Cladornidae ] (pelecaniform?
**†''Cladornis, Cladornis pachypus'' (Deseado Early Oligocene of Patagonia, Argentina)
*†Eremopezidae – pelecaniform? ratite?
**†'' Eremopezus, Eremopezus eocaenus'' (Late Eocene) - includes '' Stromeria fajumensis''
*† Gracilitarsidae – close to Sylphornithidae?
**'' Eutreptodactylus itaboraiensis'' (Late Paleocene of Brazil) - a '' nomen dubium''
**'' Gracilitarsus mirabilis'' (middle Eocene of Messel, Germany)
*Halcyornithidae
Halcyornithidae or Pseudasturidae is a family of fossil birds, possibly belonging to the order Psittaciformes (parrots and relatives). Members of this family lived in the Eocene, around 55-48 million years ago. Fossil remains are known from the ...
– psittaciform (= Pseudasturidae?), coraciiform?
**†'' Halcyornis toliapicus'' (London Clay Early Eocene of England)
**†'' Cyrilavis'' (Green River Early Eocene of C US) - psittaciform (halcyornithid or psittacid)?
***†'' C. olsoni'' 'Primobucco_olsoni''_.html"_;"title="Primobucco_olsoni.html"_;"title="'Primobucco_olsoni">'Primobucco_olsoni''_">Primobucco_olsoni.html"_;"title="'Primobucco_olsoni">'Primobucco_olsoni''_***†''Cyrilavis_colburnorum.html" ;"title="Primobucco_olsoni">'Primobucco_olsoni''_.html" ;"title="Primobucco_olsoni.html" ;"title="'Primobucco olsoni">'Primobucco olsoni'' ">Primobucco_olsoni.html" ;"title="'Primobucco olsoni">'Primobucco olsoni'' ***†''Cyrilavis colburnorum">C. colburnorum''
*†Juncitarsidae
**†''Kashinia, Kashinia magnum'' [''Tenuicrus'' ; ''Tenuicrus magnum'' ] – phoenicopteriform?
**†''Juncitarsus'' – phoenicopteriform?
*Laornithidae – charadriiform? gruiform? pelagornithid?
**'' Laornis edvardsianus'' 'Laopteryx'' (sic) (Late Cretaceous?)
*† Lonchodytidae
**†'' Lonchodytes estesi'' (Late Cretaceous/?Early Palaeocene) – gaviiform/pelecaniform? procellariiform?
*† Palaeospizidae – passeriform? coraciiform? coliiform?
**†'' Palaeospiza bella'' (Late Eocene of Florissant Fossil Beds, US)
*† Parvicuculidae – cypselomorph, cuculiform, coraciiform (primobucconid)?
**†'' Parvicuculus minor'' (Early Eocene of NW Europe)
*† Remiornithidae (Paleocene of France) – palaeognath?
**†'' Remiornis heberti'' 'Remiornis_minor''_.html" ;"title="Remiornis_minor.html" ;"title="'Remiornis minor">'Remiornis minor'' ">Remiornis_minor.html" ;"title="'Remiornis minor">'Remiornis minor'' *†Sylphornithidae – cuculiform? coraciiform? close to Gracilitarsidae?
**†''Sylphornis, Sylphornis bretouensis'' (middle Eocene of France)
**†''Oligosylphe, Oligosylphe mourerchauvireae'' (Borgloon Early Oligocene of Hoogbutsel, Belgium)
*† Tytthostonychidae – procellariiform, pelecaniform?
**†'' Tytthostonyx glauconiticus'' (Late Cretaceous/Early Palaeocene)
*† Zygodactylidae rimoscenidae near passerine
Near passerines and higher land-bird assemblage are terms of traditional, pre-cladistic taxonomy that have often been given to tree-dwelling birds or those most often believed to be related to the true passerines (order Passeriformes) owing to mor ...
**†'' Eozygodactylus americanus''
**†'' Zygodactylus'' (Early Oligocene – Middle? Miocene of C Europe)
***†'' Z. ignotus''
***†'' Z. luberonensis''
***†'' Z. grivensis''
**†'' Primoscens minutus''
**†'' Primozygodactylus''
***†'' P. ballmanni''
***†'' P. danielsi''
***†'' P. major''
***†'' P. eunjooae''
*†" Graculavidae" akotornithidae_Erickson_1975;_Scaniornithidae;_Telmatornithidae.html" ;"title="Scaniornithidae.html" ;"title="akotornithidae Erickson 1975; Scaniornithidae">akotornithidae Erickson 1975; Scaniornithidae; Telmatornithidae">Scaniornithidae.html" ;"title="akotornithidae Erickson 1975; Scaniornithidae">akotornithidae Erickson 1975; Scaniornithidae; Telmatornithidae Cracraft 1972] – a paraphyletic form taxon, the "transitional shorebirds"
**†''Limosavis'' [''Graculavus
''Graculavus'' is a prehistoric bird genus that was described in the 19th century by American paleontologist O. C. Marsh. Its remains were found in the Late Cretaceous Austin Chalk of Texas, USA, and Lance Formation (late Maastrichtian faunal sta ...
'' ] (Late Cretaceous –? Early Palaeocene) – charadriiform?
***†'' Limosavis velox, L. velox'' ' 'Graculavus_velox''_">Graculavus_velox.html"_;"title="'Graculavus_velox">'Graculavus_velox''_***†''Limosavis_augustus.html" ;"title="Graculavus_velox''_.html" ;"title="Graculavus_velox.html" ;"title="'Graculavus velox">'Graculavus velox'' ">Graculavus_velox.html" ;"title="'Graculavus velox">'Graculavus velox'' ***†''Limosavis augustus">L. augustus'' [''Graculavus augustus'' ]
**†''Palaeotringa'' (Late Cretaceous/Early Palaeocene) – charadriiform?
**†''Telmatornis, Telmatornis priscus'' [''Telmatornis affinis'' ; ''Graculavus pumilis'' ; ''Palaeotringa vetus'' ] (Late Cretaceous?) – charadriiform? gruiform? podicipediform?
**†'' Zhylgaia, Zhylgaia aestiflua'' (Early Paleocene) – presbyornithid?
**†'' Scaniornis lundgreni'' (Early/Middle Paleocene) – phoenicopteriform?
**†'' Dakotornis cooperi'' (Paleocene of North Dakota, US)
**†''Placement unresolved''
***"Graculavidae" gen. et sp. indet. (Gloucester County, US)
Struthioniformes
Struthioniformes is an order of birds with only a single extant family, Struthionidae, containing the ostriches. Several other extinct families are known, spanning across the Northern Hemisphere, from the Early Eocene to the early Pliocene, includ ...
Ostrich
Ostriches are large flightless birds of the genus ''Struthio'' in the order Struthioniformes, part of the infra-class Palaeognathae, a diverse group of flightless birds also known as ratites that includes the emus, rheas, and kiwis. There ...
and related ratites.
* Struthionidae Struthiolithidae ; †Palaeotididae (ostriches)
**†'' Palaeotis weigelti'' 'Palaeogrus_geiseltalensis''_;_''Ornitocnemus_geiseltalensis.html" ;"title="Palaeogrus_geiseltalensis.html" ;"title="'Palaeogrus geiseltalensis">'Palaeogrus geiseltalensis'' ; ''Ornitocnemus geiseltalensis">Palaeogrus_geiseltalensis.html" ;"title="'Palaeogrus geiseltalensis">'Palaeogrus geiseltalensis'' ; ''Ornitocnemus geiseltalensis'' ] (middle Eocene)
**†''Remiornis heberti''
**†''Orientornis, Orientornis linxiaensis'' [''Struthio linxiaensis'' ]
**''Prehistoric species of extant genera''
***''Struthio'' ''Palaeostruthio''_;_†''Struthiolithus.html" ;"title="Palaeostruthio.html" ;"title="''Palaeostruthio">''Palaeostruthio'' ; †''Struthiolithus">Palaeostruthio.html" ;"title="''Palaeostruthio">''Palaeostruthio'' ; †''Struthiolithus'' ootaxa; ''Struthio'' (''Pachystruthio'') ; ''Megaloscelornis'' ] (Early Miocene – Recent)
Casuariiformes
Cassowaries, emus and related ratites.
*Casuariidae Kaup 1847 [Dromaiidae Huxley 1868; Dramaiinae Gray 1870; Dramiceiidae Richmond 1908; Dramaeidae Newton 1896] (emus and cassowaries)
**†'' Diogenornis, Diogenornis fragilis'' (Early Eocene) – possible casuariiform
**†''Emuarius
''Emuarius'' is an extinct genus of casuariiform flightless bird from Australia that lived during the early Miocene and late Oligocene. It is one of two known genera of emu. There are two known species in the genus, ''Emuarius gidju'' and ' ...
'' (emuwaries) (Late Oligocene – Late Miocene)
**''Prehistoric species of extant genera''
***''Dromaius
''Dromaius'' (from greek δρομαίυς "runner") is a genus of ratite present in Australia. There is one extant species, '' Dromaius novaehollandiae'' commonly known as the emu.
In his original 1816 description of the emu, Louis Jean Pierre ...
'' 'Tachea''_;_''Emou.html"_;"title="Tachea.html"_;"title="'Tachea">'Tachea''_;_''Emou">Tachea.html"_;"title="'Tachea">'Tachea''_;_''Emou''_;_''Peronista.html" ;"title="Tachea">'Tachea''_;_''Emou.html" ;"title="Tachea.html" ;"title="'Tachea">'Tachea'' ; ''Emou">Tachea.html" ;"title="'Tachea">'Tachea'' ; ''Emou'' ; ''Peronista">Tachea">'Tachea''_;_''Emou.html" ;"title="Tachea.html" ;"title="'Tachea">'Tachea'' ; ''Emou">Tachea.html" ;"title="'Tachea">'Tachea'' ; ''Emou'' ; ''Peronista'' ; ''Metapteryx'' ] (emus) (Middle Miocene – Recent)
***''Casuarius'' [''Cela (bird), Cela'' ; ''Cela (bird), Cela'' ; '' Rhea'' ; ''Chelarga'' ; ''Oxyporus'' ; ''Thrasys'' ; '' Cassowara'' ; '' Hippalectryo'' ]
Rheiformes
Rheiformes is an order that contains the family Rheidae (rheas). It is in the infraclass Paleognathae, which contains all ratites. Extant members are found in South America. While the IOC World Bird List and the Clements Checklist categorise Rh ...
Rheas
The rheas ( ), also known as ñandus ( ) or South American ostriches, are large ratites (flightless birds without a keel on their sternum bone) in the order Rheiformes, native to South America, distantly related to the ostrich and emu. Most tax ...
and related ratites.
*† Opisthodactylidae
**†''Opisthodactylus
''Opisthodactylus'' is an extinct genus of rhea from the Early to Middle Miocene Santa Cruz and Chichinales Formations and the Late Miocene ( Montehermosan) Andalhuala Formation of Argentina. Three species are described: the type species, ''O ...
'' (Miocene) – rheid?
*Rheidae
Rheidae is a family of flightless ratite birds which first appeared in the Paleocene. It is today represented by the sole living genus '' Rhea'', but also contains several extinct genera.
Taxonomy
Order Rheiformes (Forbes, 1884) Furbringer, 18 ...
heinae (rheas)
**†'' Heterorhea dabbenei'' (Pliocene)
**†'' Hinasuri nehuensis''
**''Prehistoric species of extant genera''
***'' Rhea'' 'Rhea''_;_''Pterocnemia''_;_'' 'Rhea''_;_''Pterocnemia''_;_''Toujou_(bird)">Toujou''_;_''Tujus.html" ;"title="Toujou_(bird).html" ;"title="Pterocnemia.html" ;"title="'Rhea'' ; ''Pterocnemia">'Rhea'' ; ''Pterocnemia'' ; ''Toujou (bird)">Toujou'' ; ''Tujus">Toujou_(bird).html" ;"title="Pterocnemia.html" ;"title="'Rhea'' ; ''Pterocnemia">'Rhea'' ; ''Pterocnemia'' ; ''Toujou (bird)">Toujou'' ; ''Tujus'' ]
†Dinornithiformes
Moas.
*†Megalapteryx didinus, Megalapterygidae - (Upland moas)
**†''Megalapteryx didinus'' [''Palaeocasuarius'' ; ''Dinornis didinus'' ; '' Megalapteryx benhami'' ; '' Megalapteryx hamiltoni'' ; '' Megalapteryx hectori'' ; '' Megalapteryx tenuipes'' ; '' Anomalopteryx didina'' ; '' Palaeocasuarius velox'' ; '' Palaeocasuarius elegans'' ; '' Palaeocasuarius haasti'' ]
*†Dinornithidae
The giant moa (''Dinornis'') is an extinct genus of birds belonging to the moa family. As with other moa, it was a member of the order Dinornithiformes. It was endemic to New Zealand. Two species of ''Dinornis'' are considered valid, the Nort ...
alapteryginae ; Palapterygidae ; Dinornithnideae (great moas)
**†''Dinornis
The giant moa (''Dinornis'') is an extinct genus of birds belonging to the moa family. As with other moa, it was a member of the order Dinornithiformes. It was endemic to New Zealand. Two species of ''Dinornis'' are considered valid, the North ...
'' ' 'Palapteryx''_;_''Megalornis">Palapteryx.html"_;"title="'Palapteryx">'Palapteryx''_;_''Megalornis''_;_''Dinornis.html" ;"title="Palapteryx''_;_''Megalornis.html" ;"title="Palapteryx.html" ;"title="'Palapteryx">'Palapteryx'' ; ''Megalornis">Palapteryx.html" ;"title="'Palapteryx">'Palapteryx'' ; ''Megalornis'' ; ''Dinornis">Movia'' ; ''Moa'' ; ''Dinornis, Owenia'' ; ''Tylopteryx'' ]
*†Emeidae [Emeinae ; Anomalopterygidae ; Anomalapteryginae ] (lesser moas)
**†''Anomalopteryx didiformis'' (Little bush moas)
**†''Emeus crassus
The eastern moa (''Emeus crassus'') is an extinct species of moa. When the first specimens were originally described by Richard Owen, they were placed within the genus ''Dinornis'' as three different species, but, was later split off into their ...
'' Reichenbach 1853 (Eastern moas)
**†'' Euryapteryx'' (Broad-billed or turkey moas)
**†''Pachyornis
''Pachyornis'' is an extinct genus of ratites from New Zealand which belonged to the moa family. Like all ratites it was a member of the order Struthioniformes. The Struthioniformes are flightless birds with a sternum without a keel. They al ...
'' (Stout moas)
Two unnamed Saint Bathans Fauna
The St Bathans fauna is found in the lower Bannockburn Formation of the Manuherikia Group of Central Otago, in the South Island of New Zealand. It comprises a suite of fossilised prehistoric animals from the late Early Miocene (Altonian) period, ...
species.
Apterygiformes
* Apterygidae pteryginae (Kiwis)
**†'' Proapteryx micromeros''
†
Lithornithiformes
Lithornithidae is an extinct, possibly paraphyletic (but see below) group of early paleognath birds. They are known from fossils dating to the Upper Paleocene through the Middle Eocene of North America and Europe, with possible Late Cretaceous ...
*†Lithornithidae
Lithornithidae is an extinct, possibly paraphyletic (but see below) group of early paleognath birds. They are known from fossils dating to the Upper Paleocene through the Middle Eocene of North America and Europe, with possible Late Cretaceous ...
(False tinamous)
**†'' Fissuravis weigelti''
**†'' Pseudocrypturus cercanaxius'' (Green River Early/middle Eocene of C US)
**†''Paracathartes howardae
''Paracathartes'' is a genus of extinct bird from the Wasachtian horizon of lower Eocene Wyoming. One species, ''Paracathartes howardae'' has been described.
It is a paleognathous bird, turkey-like in stature and size, that probably resembled ...
''
**†''Lithornis
''Lithornis'' is a genus of extinct paleognathous birds. Although ''Lithornis'' was able to fly well, their closest relatives are the extant tinamous (which are poor flyers) and ratites (which are flightless birds).
Fossils of ''Lithornis'' a ...
'' 'Promusophaga''_;_''Pediorallus.html" ;"title="Promusophaga.html" ;"title="'Promusophaga">'Promusophaga'' ; ''Pediorallus">Promusophaga.html" ;"title="'Promusophaga">'Promusophaga'' ; ''Pediorallus'' ; ''Parvigyps'' ] (Paleocene – Early Eocene)
Tinamiformes
*Tinamidae [Crypturidae ; Tinamotidae ; Eudromiidae ; Rhynchotidae ] (tinamous)
**†''Querandiornis, Querandiornis romani'' (Ensenada Early/Middle Pleistocene of Argentina)
**†'' Roveretornis''
**Tinamidae gen. et sp. indet. MACN-SC Fleagle Collection (Early – Middle Miocene of S Argentina) - at least 2 species
**''Prehistoric species of extant genera''
***''Eudromia
''Eudromia'' is a genus of birds in the tinamou family. This genus comprises two crested members of this South American family.
Etymology
''Eudromia'' comes from two Greek words, ''eu'' meaning well or nicely, and ''dromos'' meaning a running e ...
'' on_''Eudromias''_;_''Calodromas.html" ;"title="Eudromias.html" ;"title="on ''Eudromias">on ''Eudromias'' ; ''Calodromas">Eudromias.html" ;"title="on ''Eudromias">on ''Eudromias'' ; ''Calodromas'' ''Calodromus'' ; ''Calopezus'' ; ''Tinamisornis'' ; '' Roveretornis'' ]
***''Nothura'' [†''Cayetanornis'' ]
***''Crypturellus
''Crypturellus'' is a genus of tinamous containing mostly forest species. However, there are the odd few that are grassland or steppe tinamous. The genus contains 21 species.
Taxonomy
The genus ''Crypturellus'' was introduced in 1914 by the Brit ...
'' 'Microcrypturus''_;_''Crypturornis.html" ;"title="Microcrypturus.html" ;"title="'Microcrypturus">'Microcrypturus'' ; ''Crypturornis">Microcrypturus.html" ;"title="'Microcrypturus">'Microcrypturus'' ; ''Crypturornis'' ; ''Orthocrypturus'' ]
Vegaviiformes
*†''Neogaeornis, Neogaeornis wetzeli'' Lambrecht 1929 (Late Cretaceous)
*†''Polarornis, Polarornis gregorii'' Chatterjee 2002 (Late Cretaceous)
*†''Vegavis iaai
''Vegavis'' is a genus of extinct bird that lived during the Late Cretaceous (Maastrichtian stage) of Antarctica, some 68 to 66 mya. Among modern birds, most studies show that ''Vegavis'' is most closely related to ducks and geese (Anatidae), b ...
'' (Late Cretaceous)
*†'' Maaqwi cascadensis'' (Late Cretaceous)
*†'' Australornis lovei'' (Paleocene)
Anseriformes
The group that includes modern duck
Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a form ...
s and geese
A goose ( : geese) is a bird of any of several waterfowl species in the family Anatidae. This group comprises the genera '' Anser'' (the grey geese and white geese) and ''Branta'' (the black geese). Some other birds, mostly related to the she ...
.
*Basal and unresolved forms
**†''Anatalavis
''Anatalavis'' is genus of prehistoric birds related to ducks and geese, perhaps in particular the magpie-goose. The species ''Anatalavis rex'' – formerly placed in ''Telmatornis'' – is known from the Hornerstown Formation (Late Cretaceous o ...
'' ' 'Nettapterornis''_">Nettapterornis.html"_;"title="'Nettapterornis">'Nettapterornis''_(Late_Cretaceous/Early_Paleocene–_Early_Eocene)–_anseranatid_or_basal.
***†''Anatalavis_rex.html" ;"title="Nettapterornis''_.html" ;"title="Nettapterornis.html" ;"title="'Nettapterornis">'Nettapterornis'' ">Nettapterornis.html" ;"title="'Nettapterornis">'Nettapterornis'' (Late Cretaceous/Early Paleocene– Early Eocene)– anseranatid or basal.
***†''Anatalavis rex">A. rex'' [''Telmatornis rex'' ]
***†''Anatalavis oxfordi, A. oxfordi'' [''Nettapterornis oxfordi'' ]
**†''Romainvillia, Romainvillia stehlini'' (Late Eocene/Early Oligocene)– anseranatide or anatide
**†''Saintandrea, Saintandrea chenoides''
**†'' Proherodius, Proherodius oweni'' (Early Eocene) – presbyornithid?
**†'' Paranyroca magna'' (Early Miocene) – anatide or own family?
**†'' Garganornis ballmanni''
* Anhimidae alamedeinae ; Palamedeidae (screamers
**†'' Chaunoides antiquus''
*†Dromornithidae
Dromornithidae, known as mihirungs and informally as thunder birds or demon ducks, were a clade of large, flightless Australian birds of the Oligocene through Pleistocene Epochs. All are now extinct. They were long classified in Struthioniform ...
– Australian mihirungs or "demon ducks"
**†'' Barawertornis tedfordi'' (Late Oligocene – Early Miocene)
**†''Bullockornis planei
''Dromornis'' is a genus of large to enormous prehistoric birds.
The species were flightless, possessing greatly reduced wing structures but with large legs, similar to the modern ostrich or emu. They were likely to have been predominantly, if ...
'' (Demon-Duck of Doom) (Middle Miocene)
**†''Ilbandornis
''Ilbandornis'' was a genus of ostrich-sized dromornithid. It was far more lightly built than other members of the family, indicating a more cursorial lifestyle. The majority of researchers consider Dromornithids to be herbivorous; this is borne ...
'' (Late Miocene)
***†'' I. woodburnei''
***†'' I. lawsoni''
**†''Dromornis
''Dromornis'' is a genus of large to enormous prehistoric birds.
The species were flightless, possessing greatly reduced wing structures but with large legs, similar to the modern ostrich or emu. They were likely to have been predominantly, i ...
'' (Late Miocene – Pliocene)
***†'' D. australis''
***†'' D. stirtoni'' (Stirton's Thunder Birds)
*Anseranatidae
Anseranatidae, the magpie-geese, is a biological family of waterbirds. The only living species, the magpie goose, is a resident breeder in northern Australia and in southern New Guinea.
Systematics and evolution
This family is placed in the o ...
(magpie-geese
**†'' Eoanseranas handae'' (Hand's Dawn Magpie Goose)
**Anseranatidae gen. et sp. nov (Late Oligocene)
*†Presbyornithidae
Presbyornithidae is an extinct group of birds with a global distribution. They had evolved by the late Cretaceous period and became extinct during the early Miocene. Initially, they were believed to present a mix of characters shown by waterbirds ...
elmabatidae **†'' Teviornis gobiensis'' (Late Cretaceous)
**†'' Headonornis hantoniensis'' 'Agnopterus_hantoniensis''_;_''Presbyornis_hantoniensis.html" ;"title="Agnopterus_hantoniensis.html" ;"title="'Agnopterus hantoniensis">'Agnopterus hantoniensis'' ; ''Presbyornis hantoniensis">Agnopterus_hantoniensis.html" ;"title="'Agnopterus hantoniensis">'Agnopterus hantoniensis'' ; ''Presbyornis hantoniensis'' ; '' Ptenornis''] – only BMNH PAL 30325 belongs to this bird, may belong to ''Presbyornis''.
**†''Presbyornis'' [''Nautilornis'' ; ''Coltonia'' ] (Paleocene– Early Oligocene)
***†''Presbyornis, P. isoni''
***†'' Presbyornis, P. mongoliensis''
***†'' P. recurvirostrus'' ' 'Coltonia_recurvirostra''_">Coltonia_recurvirostra.html"_;"title="'Coltonia_recurvirostra">'Coltonia_recurvirostra''_***†''Presbyornis.html" ;"title="Coltonia_recurvirostra''_.html" ;"title="Coltonia_recurvirostra.html" ;"title="'Coltonia recurvirostra">'Coltonia recurvirostra'' ">Coltonia_recurvirostra.html" ;"title="'Coltonia recurvirostra">'Coltonia recurvirostra'' ***†''Presbyornis">P. pervetus'' [''Nautilornis avus'' ; ''Nautilornis proavitus'' ]
**†''Telmabates''
***†''Telmabates antiquus, T. antiquus''
***†''Telmabates howardae, T. howardae''
**†''Wilaru''
***†''Wilaru, W. tedfordi''
***†'' Wilaru, W. prideauxi''
*Anatidae
The Anatidae are the biological family of water birds that includes ducks, geese, and swans. The family has a cosmopolitan distribution, occurring on all the world's continents except Antarctica. These birds are adapted for swimming, floating ...
nemiornithidae ; Cygnidae ; Anseridae ; Cereopsidae ; Fuligulidae ; Merganidae ; Mergidae ; Plectropteridae ; Erismaturidae ; Tadornidae (ducks, geese and swans)
**†'' Allgoviachen tortonica'' (Late Miocene)
**†''Annakacygna
''Annakacygna'' is a genus of flightless marine swan from the Miocene of Japan. Named in 2022, ''Annakacygna'' displays a series of unique adaptations setting it apart from any other known swan, including a filter feeding lifestyle, a highly mob ...
'' (Late Miocene)
***†'' A. hajimei''
***†'' A. yoshiiensis ''
**†'' Bonibernicla ponderosa'' ' 'Branta_woolfendeni''_Bickart_1990">Branta_woolfendeni.html"_;"title="'Branta_woolfendeni">'Branta_woolfendeni''_Bickart_1990**†''Eonessa.html" ;"title="Branta_woolfendeni''_Bickart_1990.html" ;"title="Branta_woolfendeni.html" ;"title="'Branta woolfendeni">'Branta woolfendeni'' Bickart 1990">Branta_woolfendeni.html" ;"title="'Branta woolfendeni">'Branta woolfendeni'' Bickart 1990**†''Eonessa">Eonessa anaticula'' (Middle Eocene)
**†''Cygnavus'' (Early Oligocene – Early Miocene)
***†''Cygnavus senckenbergi, C. senckenbergi'' Lambrecht 1931
***†''Cygnavus formosus, C. formosus'' Korochkin 1968
**†''Cygnopterus'' Lambrecht 1931 (middle Oligocene – Early Miocene)– sometimes included in ''Cygnavus''
***†'' C. neogradiensis'' Kessler & Hir 2009
***†'' C. affinis'' (Beneden 1883) Lambrecht 1931 'Sula_affinis''_Beneden_1883;_''Palaeopapia_hamsteadiensis.html" ;"title="Sula_affinis.html" ;"title="'Sula affinis">'Sula affinis'' Beneden 1883; ''Palaeopapia hamsteadiensis">Sula_affinis.html" ;"title="'Sula affinis">'Sula affinis'' Beneden 1883; ''Palaeopapia hamsteadiensis'' Harrison & Walker 1979b]
**†''Mionetta'' Livezey & Martin 1988 (Late Oligocene – Early Miocene)
***†''Mionetta robusta, M. robusta'' (Milne-Edwards 1868) Mlíkovský 2002 [''Anas robusta'' Milne-Edwards, 1868; '' Anserobranta robusta'' (Milne-Edwards 1868) Cheneval 1987]
***†'' Mionetta arvernensis, M. arvernensis'' (Lydekker 1891a) 'Aythya_arvernensis''_(Lydekker_1891a)_Brodkorb_1964;_''Fuligula_arvernensis.html" ;"title="Aythya_arvernensis.html" ;"title="'Aythya arvernensis">'Aythya arvernensis'' (Lydekker 1891a) Brodkorb 1964; ''Fuligula arvernensis">Aythya_arvernensis.html" ;"title="'Aythya arvernensis">'Aythya arvernensis'' (Lydekker 1891a) Brodkorb 1964; ''Fuligula arvernensis'' Lydekker, 1891a]
***†''Mionetta blanchardi, M. blanchardi'' (Milne-Edwards 1863) Livezey & Martin 1988 [''Anas blanchardi'' Milne-Edwards 1863; ''Palaeortyx phasianoides'' Milne-Edwards 1869; ''Anas macroptera'' Milne-Edwards 1871 Nomen nudum; '' Taoperdix phasianoides'' (Milne-Edwards 1869) Brodkorb 1964; '' Dendrochen blanchardi'' (Milne-Edwards 1863) Cheneval 1983b]
***†'' Mionetta consobrina, M. consobrina'' (Milne-Edwards 1867) Livezey & Martin 1988 'Anas_consobrina''_Milne-Edwards_1867;_''Dendrochen_consobrina.html" ;"title="Anas_consobrina.html" ;"title="'Anas consobrina">'Anas consobrina'' Milne-Edwards 1867; ''Dendrochen consobrina">Anas_consobrina.html" ;"title="'Anas consobrina">'Anas consobrina'' Milne-Edwards 1867; ''Dendrochen consobrina'' (Milne-Edwards 1867) Cheneval 1983b]
***†''Mionetta natator, M. natator'' (Milne-Edwards 1867) Livezey & Martin 1988 [''Anas natator'' Milne-Edwards 1867; ''Querquedula natator'' (Milne-Edwards 1867) Brodkorb 1962; ''Dendrochen natator'' (Milne-Edwards 1867) Cheneval 1983b]
**†'' Notochen bannockburnensis'' (Early Miocene)
**†'' Australotadorna alecwilsoni'' Worthy 2009 (Late Oligocene – Early Miocene)
**†'' Pinpanetta'' Worthy 2009 (Late Oligocene – Early Miocene)
***†'' P. tedfordi'' Worthy 2009
***†'' P. vickersrichae'' Worthy 2009
***†'' P. fromensis'' Worthy 2009
**†'' Dunstanetta johnstoneorum'' Worthy et al. 2007 (Johnstone's ducks) (Early/Middle Miocene)
**†'' Manuherikia'' Worthy et al. 2007 (Early/Middle Miocene)
***†'' M. lacustrina'' Worthy et al. 2007 (Manuherikia duck)
***†'' M. minuta'' Worthy et al. 2007 (Minute Manuherikia duck)
***†'' M. douglasi'' Worthy et al. 2008 (Douglas’ duck)
**†'' Matanas enrighti'' Worthy et al. 2007 (Enright's ducks) (Early/Middle Miocene)
**†''Miotadorna
''Miotadorna'' is a genus of extinction, extinct Tadorninae, tadornine ducks from the Miocene of New Zealand. It contains two species, ''M. sanctibathansi'', and ''M. catrionae'' (Catriona's shelduck).
''M. sanctibathansi''
''M. sanctibathansi'' ...
'' Worthy et al. 2007 (Early/Middle Miocene)
***†'' M. sanctibathansi'' Worthy et al. 2007 (St. Bathans shelducks) (Early/Middle Miocene)
***†'' M. catrionae'' Tennyson et al. 2022 (Catriona's shelduck) (late Early/early Middle Miocene) (synonym of ''M. sanctibathansi'')
**†'' Megalodytes morejohni'' Howard 1992 (Middle Miocene)
**†'' Sinanas diatomas'' Yeh 1980 (Middle Miocene)
**†'' Dendrochen'' Miller 1944 (Late Miocene)
***†'' D. integra'' (Miller 1944) Cheneval 1987 'Querquedula integra'' Miller 1944; ''Anas integra'' (Miller 1944); ''Spatula integra'' (Miller 1944)***†'' D. oligocaena'' (Tugarinov 1940) ' 'Anas_oligocaena''_Tugarinov_1940">Anas_oligocaena.html"_;"title="'Anas_oligocaena">'Anas_oligocaena''_Tugarinov_1940***†'' ''_Tugarinov_1940.html"_;"title="Anas_oligocaena.html"_;"title="'Anas_oligocaena">'Anas_oligocaena''_Tugarinov_1940">Anas_oligocaena.html"_;"title="'Anas_oligocaena">'Anas_oligocaena''_Tugarinov_1940***†''Dendrochen_robusta">D._robusta''_Miller_1944
**†'' ''_Tugarinov_1940.html"_;"title="Anas_oligocaena.html"_;"title="'Anas_oligocaena">'Anas_oligocaena''_Tugarinov_1940">Anas_oligocaena.html"_;"title="'Anas_oligocaena">'Anas_oligocaena''_Tugarinov_1940***†''Dendrochen_robusta">D._robusta''_Miller_1944
**†''Presbychen">Presbychen_abavus''_Wetmore_1930_(Late_Miocene)
**†'' ''_Tugarinov_1940.html"_;"title="Anas_oligocaena.html"_;"title="'Anas_oligocaena">'Anas_oligocaena''_Tugarinov_1940">Anas_oligocaena.html"_;"title="'Anas_oligocaena">'Anas_oligocaena''_Tugarinov_1940***†''Dendrochen_robusta">D._robusta''_Miller_1944
**†''Presbychen">Presbychen_abavus''_Wetmore_1930_(Late_Miocene)
**†''Afrocygnus">Afrocygnus_chauvireae''_Louchart_et_al._2005_(Late_Miocene_–_Early_Pliocene)
**†'' ''_Tugarinov_1940.html"_;"title="Anas_oligocaena.html"_;"title="'Anas_oligocaena">'Anas_oligocaena''_Tugarinov_1940">Anas_oligocaena.html"_;"title="'Anas_oligocaena">'Anas_oligocaena''_Tugarinov_1940***†''Dendrochen_robusta">D._robusta''_Miller_1944
**†''Presbychen">Presbychen_abavus''_Wetmore_1930_(Late_Miocene)
**†''Afrocygnus">Afrocygnus_chauvireae''_Louchart_et_al._2005_(Late_Miocene_–_Early_Pliocene)
**†''Balcanas">Balcanas_pliocaenica''_(Early_Pliocene) –_may_belong_in_''Tadorna''
**†'' ''_Tugarinov_1940.html"_;"title="Anas_oligocaena.html"_;"title="'Anas_oligocaena">'Anas_oligocaena''_Tugarinov_1940">Anas_oligocaena.html"_;"title="'Anas_oligocaena">'Anas_oligocaena''_Tugarinov_1940***†''Dendrochen_robusta">D._robusta''_Miller_1944
**†''Presbychen">Presbychen_abavus''_Wetmore_1930_(Late_Miocene)
**†''Afrocygnus">Afrocygnus_chauvireae''_Louchart_et_al._2005_(Late_Miocene_–_Early_Pliocene)
**†''Balcanas">Balcanas_pliocaenica''_(Early_Pliocene) –_may_belong_in_''Tadorna''
**†''Wasonaka">Wasonaka_yepomerae''_Howard_1966_(Middle_Pliocene)
**†''Paracygnus.html" ;"title="Wasonaka.html" ;"title="Balcanas.html" ;"title="Afrocygnus.html" ;"title="Presbychen.html" ;"title="Dendrochen_robusta.html" ;"title="Anas_oligocaena''_Tugarinov_1940.html" ;"title="Anas_oligocaena.html" ;"title="'Anas oligocaena">'Anas oligocaena'' Tugarinov 1940">Anas_oligocaena.html" ;"title="'Anas oligocaena">'Anas oligocaena'' Tugarinov 1940***†''Dendrochen robusta">D. robusta'' Miller 1944
**†''Presbychen">Presbychen abavus'' Wetmore 1930 (Late Miocene)
**†''Afrocygnus">Afrocygnus chauvireae'' Louchart et al. 2005 (Late Miocene – Early Pliocene)
**†''Balcanas">Balcanas pliocaenica'' (Early Pliocene) – may belong in ''Tadorna''
**†''Wasonaka">Wasonaka yepomerae'' Howard 1966 (Middle Pliocene)
**†''Paracygnus">Paracygnus plattensis'' Short 1969 (Late Pliocene)
**†''Anabernicula'' Ross 1935 (Late Pliocene? – Late Pleistocene)
***†''Anabernicula, A. minuscula'' (Wetmore 1924) [''Branta minuscula'' Wetmore 1924] (Late Pliocene of Benson, Arizona)
***†''Anabernicula, A. oregonensis'' Howard 1964b
***†'' Anabernicula, A. gracilenta'' Ross 1935
**†'' Eremochen russelli'' Brodkorb 1961 (Pliocene)
**†'' Tirarinetta kanunka'' Worthy 2008 (Pliocene)
**†'' Brantadorna'' Howard 1964 (Middle Pleistocene)
***†'' B. downsi'' Howard 1964
***†'' B. robusta'' (Short 1970) Livezey 1997 ' 'Anabernicula_robusta''_Short_1970">Anabernicula_robusta.html"_;"title="'Anabernicula_robusta">'Anabernicula_robusta''_Short_1970**†'' ''_Short_1970.html"_;"title="Anabernicula_robusta.html"_;"title="'Anabernicula_robusta">'Anabernicula_robusta''_Short_1970">Anabernicula_robusta.html"_;"title="'Anabernicula_robusta">'Anabernicula_robusta''_Short_1970**†''Nannonetta">Nannonetta_invisitata''_Campbell_1979_(Late_Pleistocene)
**†'' ''_Short_1970.html"_;"title="Anabernicula_robusta.html"_;"title="'Anabernicula_robusta">'Anabernicula_robusta''_Short_1970">Anabernicula_robusta.html"_;"title="'Anabernicula_robusta">'Anabernicula_robusta''_Short_1970**†''Nannonetta">Nannonetta_invisitata''_Campbell_1979_(Late_Pleistocene)
**†''Aldabranas">Aldabranas_cabri''_Harrison_&_Walker_1978_(Late_Pleistocene)
**†'' ''_Short_1970.html"_;"title="Anabernicula_robusta.html"_;"title="'Anabernicula_robusta">'Anabernicula_robusta''_Short_1970">Anabernicula_robusta.html"_;"title="'Anabernicula_robusta">'Anabernicula_robusta''_Short_1970**†''Nannonetta">Nannonetta_invisitata''_Campbell_1979_(Late_Pleistocene)
**†''Aldabranas">Aldabranas_cabri''_Harrison_&_Walker_1978_(Late_Pleistocene)
**†''Ankonetta">Ankonetta_larriestrai''_Cenizo_&_Agnolín_2010
**†''Cayaoa.html" ;"title="Ankonetta.html" ;"title="Aldabranas.html" ;"title="Nannonetta.html" ;"title="Anabernicula_robusta''_Short_1970.html" ;"title="Anabernicula_robusta.html" ;"title="'Anabernicula robusta">'Anabernicula robusta'' Short 1970">Anabernicula_robusta.html" ;"title="'Anabernicula robusta">'Anabernicula robusta'' Short 1970**†''Nannonetta">Nannonetta invisitata'' Campbell 1979 (Late Pleistocene)
**†''Aldabranas">Aldabranas cabri'' Harrison & Walker 1978 (Late Pleistocene)
**†''Ankonetta">Ankonetta larriestrai'' Cenizo & Agnolín 2010
**†''Cayaoa">Cayaoa bruneti'' Tonni 1979
**†''Chenoanas'' Zelenkov 2012
***†''Chenoanas, C. asiatica'' Zelenkov ''et al.'' 2018
***†''Chenoanas, C. deserta'' Zelenkov 2012
***†''Chenoanas, C. sansaniensis'' (Milne-Edwards 1867) (Middle Miocene of Siberia and Late Miocene of France)
**†'' Helonetta brodkorbi'' Emslie 1992
**†'' Mioquerquedula minutissima'' Zelenkov & Kuročkin 2012 'Anas_velox''_Milne-Edwards_1867;_''Anas_meyerii.html" ;"title="Anas_velox.html" ;"title="'Anas velox">'Anas velox'' Milne-Edwards 1867; ''Anas meyerii">Anas_velox.html" ;"title="'Anas velox">'Anas velox'' Milne-Edwards 1867; ''Anas meyerii'' Milne-Edwards, 1868; ''Nettion velox'' (Milne-Edwards 1868) Brodkorb 1964; ''Aythya meyerii'' (Milne-Edwards 1868) Brodkorb 1964] (Middle – Late? Miocene)
**†''Proanser, Proanser major'' Umanskaya 1979
**†''Shiriyanetta, Shiriyanetta hasegawai'' Watanabe & Matsuoka 2015
**†'' Pleistoanser, Pleistoanser bravardi'' Agnolín 2006
**†'' Lavadytis pyrenae'' Stidham & Hilton 2015
**†'' Asiavis phosphatica'' Nesov 1986
**†'' Nogusunna conflictoides'' Zelenkov 2011
**†'' Sharganetta mongolica'' Zelenkov 2011
**†'' Protomelanitta'' Zelenkov 2011
***†'' P. gracilis'' Zelenkov 2011
***†'' P. shihuibas'' (Hou 1985) Zelenkov 2012 ''Aythya_shihuibas''_Hou_1985">Aythya_shihuibas.html"_;"title="'Aythya_shihuibas">'Aythya_shihuibas''_Hou_1985**†''Bambolinetta.html" "title="Aythya_shihuibas''_Hou_1985.html" ;"title="Aythya_shihuibas.html" ;"title="'Aythya shihuibas">'Aythya shihuibas'' Hou 1985">Aythya_shihuibas.html" ;"title="'Aythya shihuibas">'Aythya shihuibas'' Hou 1985**†''Bambolinetta">Bambolinetta lignitifila
''Bambolinetta lignitifila'' is a fossil species of waterfowl from the Late Miocene of Italy, now classified as the sole member of the genus ''Bambolinetta''. First described in 1884 as a typical dabbling duck, it was not revisited until 2014, ...
'' (Portis 1884) Mayr & Pavia 2014 [''Anas lignitifila'' Portis 1884] (Early Pliocene of Monte Bamboli, Italy)
**†''Heteroanser, Heteroanser vicinus'' (Kuročkin 1976) Zelenkov 2012 [''Heterochen vicinus'' Kuročkin 1976; ''Anser vicinus'' (Kuročkin 1976) Mlíkovský & Švec 1986]
**''Placement unresolved''
***†''"Anas
''Anas'' is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was s ...
" albae'' Jánossy 1979 (Late Miocene) – formerly in ''Mergus''
***†''"Anas
''Anas'' is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was s ...
" amotape'' Campbell 1979
***†''"Anas
''Anas'' is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was s ...
" eppelsheimensis'' Lambrecht 1933 (Early Pliocene)
***†''"Anas
''Anas'' is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was s ...
" isarensis'' Lambrecht 1933 (Late Miocene)
***†''"Anas
''Anas'' is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was s ...
" luederitzensis'' (Early Miocene)
***†''"Anas
''Anas'' is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was s ...
" sanctaehelenae'' Campbell 1979
***†''"Anser
Anser may refer to:
People
* Anser (poet), poet of ancient Rome
* Anser Farooq, Canadian lawyer
Other uses
* ANSER, a security and defense analysis group
* ''Anser'' (bird), a genus of geese
* Anser (putter), a model of golf club made by Pi ...
" scaldii'' 'Anser_scaldii''_Beneden_1872_nomen_nudum;_"Anas"_scaldii.html" ;"title="Anser_scaldii.html" ;"title="'Anser scaldii">'Anser scaldii'' Beneden 1872 nomen nudum; "Anas" scaldii">Anser_scaldii.html" ;"title="'Anser scaldii">'Anser scaldii'' Beneden 1872 nomen nudum; "Anas" scaldii(Late Miocene)
***†''"Aythya" chauvirae'' (Middle Miocene) – 2 species
***†''"Chenopis" nanus'' De Vis 1905 (Pleistocene) – at least 2 taxa, may be living species
***†''"Cygnopterus" alphonsi'' Cheneval 1984
***†''" Oxyura" doksana'' Mlíkovský 2002 (Early Miocene)
***†"cf. '' Megalodytes"'' (Middle Miocene)
***†Anatidae gen. et sp. indet. MNZ S42797 (Early/Middle Miocene)
***†Anatidae gen. et spp. indet. (Middle Miocene)
***†Anatidae gen. et spp. indet. (Late Miocene)
**''Prehistoric species of extant genera''
***''Biziura
''Biziura'' is a genus of stiff-tailed ducks endemic to Australasia and containing one living and one subfossil species.
Species
* †''B. delautouri'' Forbes, 1892 – New Zealand musk duck – previously endemic to New Zealand, and occurri ...
'' Stephens 1824
****†'' B. exhumata'' DeVis 1889
****†'' B. delautouri'' Forbes 1892 'Biziura lobata delautouri'' (Forbes 1892)(New Zealand musk duck)
***'' Dendrocygna''
****†'' D. eversa''
****†'' D. valdipinnis''
****†'' D. soporata'' 'Anas_soporata''_Kuročkin_1976.html" ;"title="Anas_soporata.html" ;"title="'Anas soporata">'Anas soporata'' Kuročkin 1976">Anas_soporata.html" ;"title="'Anas soporata">'Anas soporata'' Kuročkin 1976***''Somateria'' (middle Oligocene – Recent)
***''Bucephala (bird), Bucephala''
****†''Bucephala (bird), B. cereti'' Boeuf & Mourer-Chauviré 1992
***''Clangula''
****†''Clangula, C. matraensis'' Kessler 2009
***''Aix
Aix or AIX may refer to:
Computing
* AIX, a line of IBM computer operating systems
*An Alternate Index, for a Virtual Storage Access Method Key Sequenced Data Set
* Athens Internet Exchange, a European Internet exchange point
Places Belgi ...
''
****†'' A. praeclara'' Zelenkov & Kuročkin 2012
***'' Cygnus''
****†'' C. mariae'' Bickart 1990
****†'' C. csakvarensis'' (Lambrecht 1933) Mlíkovský 1992 'Cygnus_csákvárensis''_Lambrecht_1931a_nomen_nudum;_''Cygnanser_csakvarensis.html" ;"title="Cygnus_csákvárensis.html" ;"title="'Cygnus csákvárensis">'Cygnus csákvárensis'' Lambrecht 1931a nomen nudum; ''Cygnanser csakvarensis">Cygnus_csákvárensis.html" ;"title="'Cygnus csákvárensis">'Cygnus csákvárensis'' Lambrecht 1931a nomen nudum; ''Cygnanser csakvarensis'' (Lambrecht 1933) Kretzoi 1957; ''Olor csakvarensis'' (Lambrecht 1933) Mlíkovský 1992b]
****†''Cygnus (genus), C. liskunae'' (Kuročkin 1976) [''Anser liskunae'' Kuročkin 1976]
****†''Cygnus (genus), C. atavus'' (Fraas 1870) Mlíkovský 1992 'Anas_atava''_Fraas_1870;_''Anas_cygniformis.html" ;"title="Anas_atava.html" ;"title="'Anas atava">'Anas atava'' Fraas 1870; ''Anas cygniformis">Anas_atava.html" ;"title="'Anas atava">'Anas atava'' Fraas 1870; ''Anas cygniformis'' Fraas 1870; ''Palaelodus steinheimensis'' Fraas 1870; ''Anser atavus'' (Fraas 1870) Lambrecht 1933; ''Anser cygniformis'' (Fraas 1870) Lambrecht 1933]
****†''Cygnus paloregonus, C. paloregonus'' Cope 1878 [''Anser condoni'' Schufeldt 1892; ''Cygnus matthewi'' Schufeldt 1913]
****†'' Cygnus (genus), C. hibbardi'' Brodkorb 1958
****†'' C. lacustris'' (De Vis 1905) 'Archaeocygnus_lacustris''_De_Vis_1905.html" ;"title="Archaeocygnus_lacustris.html" ;"title="' 'Archaeocygnus_lacustris''_De_Vis_1905">Archaeocygnus_lacustris.html"_;"title="'Archaeocygnus_lacustris">'Archaeocygnus_lacustris''_De_Vis_1905****†''Cygnus_(genus).html" ;"title="Archaeocygnus lacustris">'Archaeocygnus lacustris'' De Vis 1905">Archaeocygnus_lacustris.html" ;"title="'Archaeocygnus lacustris">'Archaeocygnus lacustris'' De Vis 1905****†''Cygnus (genus)">C. verae'' Boev 2000
***''Chloephaga''
****†''Chloephaga, C. robusta'' Tambussi 1998
***''Histrionicus''
****†''Histrionicus, Histrionicus shotwelli'' (Brodkorb 1961) [''Ocyplonessa shotwelli'' Brodkorb 1961]
***''Mergus''
****†'' Mergus, M. connectens'' Jánossy 1972
****†'' M. minor'' Kessler 2009
****†'' M. miscellus'' Alvarez & Olson 1978
***''Anas
''Anas'' is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was s ...
''
****†'' A. elapsa'' DeVis 1888 'Nettion elapsum'' (DeVis 1888)(Chinchilla Late Pleistocene of Condamine River, Australia) ("Nettion")
****†'' A. gracilipes'' DeVis 1905 'Nettion gracilipes'' (DeVis 1905)(Late Pleistocene of Australia) ("Nettion")
****†'' A. moldovica'' (Late Pliocene of Tchichmiknaia, Moldovia?Georgia?)
****†'' A. strenuum'' (Late Pleistocene of Patteramordu, Australia) ("Nettion")
****†'' A. kisatibiensis'' nser kisatibiensis(Early Pliocene of Kisatibi, Georgia)
****†'' A. lambrechti'' (Spillman 1942) 'Archeoquerquedula lambrechti'' Spillman 1942; ''Archaeoquerquedula'' (sic) ''lambrechti'' Stephens; ''Querquedula lambrechti'' (Spillman 1942)Late? Pleistocene of Santa Elena peninsula, Ecuador)
****†'' A. itchtucknee'' McCoy 1963
****†'' A. kurochkini'' Zelenkov & Panteleyev 2015
****†'' A. pullulans'' (Brodkorb 1961)
****†'' A. schneideri'' Emslie 1985
****†'' A. bunkeri'' (Wetmore 1944) ' 'Nettion_bunkeri''_Wetmore_1944">Nettion_bunkeri.html"_;"title="'Nettion_bunkeri">'Nettion_bunkeri''_Wetmore_1944****†''Anas.html" ;"title="Nettion_bunkeri''_Wetmore_1944.html" ;"title="Nettion_bunkeri.html" ;"title="'Nettion bunkeri">'Nettion bunkeri'' Wetmore 1944">Nettion_bunkeri.html" ;"title="'Nettion bunkeri">'Nettion bunkeri'' Wetmore 1944****†''Anas">A. greeni'' (Brodkorb 1964) [''Nettion greeni'' Brodkorb 1964]
****†''Anas, A. ogallalae'' (Brodkorb 1962) [''Nettion ogallalae'' Brodkorb 1962]
****†''Anas, A. cheuen'' Agnolín 2006
***''Anser (genus), Anser''
****†''Anser (genus), A. arenosus'' Bickart 1990
****†'' Anser (genus), A. arizonae'' Bickart 1990
****†'' A. azerbaidzhanicus'' Serebrovsky 1940
****†'' A. devjatkini'' Kuročkin 1971
****†'' A. djuktaiensis'' Zelenkov & Kuročkin 2014
****†'' A. eldaricus'' Burchak-Abramovich & Gadzyev 1978
****†'' A. oeningensis'' (Meyer 1865) Milne-Edwards 1867b ' 'Anas_oeningensis''_Meyer_1865">Anas_oeningensis.html"_;"title="'Anas_oeningensis">'Anas_oeningensis''_Meyer_1865****†''Anser_(genus).html" ;"title="Anas_oeningensis''_Meyer_1865.html" ;"title="Anas_oeningensis.html" ;"title="'Anas oeningensis">'Anas oeningensis'' Meyer 1865">Anas_oeningensis.html" ;"title="'Anas oeningensis">'Anas oeningensis'' Meyer 1865****†''Anser (genus)">A. pratensis'' (Short 1970) [''Heterochen pratensis'' Short 1970]
****†''Anser (genus), A. tchikoicus'' Kuročkin 1985
****†''Anser (genus), A. thompsoni'' Martin & Mengel 1980
****†''Anser (genus), A. thraceiensis'' Burchak-Abramovich & Nikolov 1984
****†'' A. pressa'' (Brodkorb 1964) ' 'Chen_pressa''_Brodkorb_1964">Chen_pressa.html"_;"title="'Chen_pressa">'Chen_pressa''_Brodkorb_1964(Dwarf_Snow_goose)
****†''Anser_(genus).html" ;"title="Chen_pressa''_Brodkorb_1964.html" ;"title="Chen_pressa.html" ;"title="'Chen pressa">'Chen pressa'' Brodkorb 1964">Chen_pressa.html" ;"title="'Chen pressa">'Chen pressa'' Brodkorb 1964(Dwarf Snow goose)
****†''Anser (genus)">A. udabnensis'' Burchak-Abramovich 1957
***''Aythya''
****†''Aythya, A. chauvirae'' Cheneval 1987
****†''Aythya, A. denesi'' (Kessler 2013) Zelenkov 2016 [''Anas denesi'' Kessler 2013]
****†''Aythya, A. effodiata'' (DeVis 1905) [Nyroca effodiata DeVis 1905]
****†''Aythya, A. magna'' Kuročkin 1985
****†''Aythya, A. molesta'' (Kuročkin 1985) Zelenkov 1985 [''Anas molesta'' Kuročkin 1985]
****†''Aythya, A. reclusa'' (DeVis 1888) [''Nyroca reclusa'' DeVis 1888]
****†''Aythya, A. robusta'' (DeVis 1888) [''Nyroca robusta'' DeVis 1888]
****†''Aythya, A. spatiosa'' Kuročkin 1976
***''Branta''
****†''Branta, B. dickey'' Miller 1924
****†''Branta, B. esmeralda'' Burt 1929
****†''Branta, B. howardae'' Miller 1930
****†''Branta, B. hypsibata'' (Cope 1878) [''Anser hypsibata'' Cope 1878]
****†''Branta, B. propinqua'' Schufeldt 1892
****†''Branta, B. thessaliensis'' Boev & Koufos 2006
***'' Oxyura''
****†''Oxyura, O. bessomi'' Howard 1963
****†''Oxyura, O. hulberti'' Emslie 1992
****†''Oxyura, O. zapatanima'' Alvarez 1977
***''Chendytes''
****†''Chendytes, O. milleri'' Howard 1955 (Early Pleistocene of San Nicolás Island US)
***''Lophodytes'' (Late Pleistocene – Recent)
***''Neochen''
****†''Neochen barbadiana, N. barbadiana'' Brodkorb 1965
****†''Neochen debilis, N. debilis'' (Ameghino 1891) [''Chenalopex debilis'' Ameghino 1891]
****†''Neochen pugil, N. pugil'' (Winge 1888) [''Chenalopex pugil'' Winge 1888]
***''Tadorna''
****†''Tadorna, T. minor'' Kessler & Hir 2012
****†''Tadorna, T. petrina'' Kuročkin 1985
Galliformes
The group that includes domestic chickens and their relatives.
*Placement unresolved
**†''Archaealectrornis, Archaealectrornis sibleyi'' Crowe & Short 1992 (Oligocene) - phasianid?
**†''Archaeophasianus'' Lambrecht 1933 (Oligocene? - Late Miocene) - tetraonid or phasianid
***†''Archaeophasianus mioceanus, A. mioceanus'' (Shufeldt 1915) [''Phasianus mioceanus'' Shufeldt 1915]
***†''Archaeophasianus roberti, A. roberti'' (Stone 1915) [''Phasianus roberti'' Stone 1915]
**†''Austinornis, Austinornis lentus'' (Marsh 1877b) Clarke 2004 [''Graculavus lentus'' Marsh 1877b; ''Ichthyornis lentus'' (Marsh 1877b) Marsh 1880; ''Pedioecetes lentus'' (Marsh 1877b); ''Pedioecetes phasianellus'' (Linnaeus 1758) Shufeldt 1915; ''Tetrao phasianellus'' Linnaeus 1758] - (Late Cretaceous) tentatively placed here;
**†''Chambiortyx, Chambiortyx cristata'' Mourer-Chauviré et al. 2013
**†''"Cyrtonyx tedfordi, Cyrtonyx" tedfordi'' (Miller 1952) (Late Miocene)
**†''Linquornis, Linquornis gigantis'' Yeh 1980 (middle Miocene)
**†''Namaortyx, Namaortyx sperrgebietensis'' Mourer-Chauviré, Pickford & 2011
**†''Palaealectoris, Palaealectoris incertus'' Wetmore 1930 (Early Miocene) - tetraonid?
**†''Palaeoalectoris, Palaeoalectoris songlinensis'' Hou 1987 (middle Miocene)
**†''Palaeonossax, Palaeonossax senectus'' Wetmore 1956 (Late Oligocene) - cracid?
**†''Palaeortyx'' Milne-Edwards 1869 [''Palaeoperdix'' Milne-Edwards 1869; ''Proalector'' Brodkorb 1964] (middle Eocene -? Early Pliocene) - phasianid or odontophorid
***†''Palaeortyx, P. media'' Milne-Edwards 1871 (nomen nudum)
***†''Palaeortyx, P. volans'' Göhlich & Pavia 2008
***†''Palaeortyx, P. gallica'' Milne-Edwards 1869 non Lydekker 1891 [''Rallus dasypus'' Milne-Edwards 1892; ''Quercyrallus dasypus'' (Milne-Edwards) Lambrecht 1933; ''Taoperdix gallica'' (Milne-Edwards) Brodkorb 1964; ?''Palaeortyx intermedia'' Ballmann 1969b; ''Coturnix gallica'' (Milne-Edwards) Mlýkovský 2002a]
***†''Palaeortyx, P. brevipes'' Milne-Edwards 1869 emend. Paris 1912 [''Palaeortyx ocyptera'' Milne-Edwards 1892; ''Palaeortyx grivensis'' Lydekker 1893 non Ballmann 1969a; ''Plioperdix grivensis'' (Lydekker 1893) Brodkorb 1964; ''Palaeortyx cayluxensis'' Lydekker 1891 non Milne-Edwards 1892 emend. Gaillard 1908; ''Taoperdix brevipes'' (Milne-Edwards 1869) Brodkorb 1964]
***†''Palaeortyx, P. prisca'' (Milne-Edwards 1869) [''Alectoris prisca'' (Milne-Edwards 1869) Mlýkovský 2002a; ''Palaeoperdix prisca'' Milne-Edwards 1869 emend. Göhlich & Mourer-Chauviré 2005; ''Palaeoperdix sansaniensis'' Milne-Edwards 1869; ''Palaeocryptonyx grivensis'' Ennouchi 1930]
***†''Palaeortyx, P. phasianoides'' Milne-Edwards 1869 [''Palaeoperdix longipes'' Milne-Edwards 1869; ''Palaeortyx longipes'' (Milne-Edwards 1869) Mlíkovský 2000e; ''Coturnix longipes'' (Milne-Edwards 1869) Mlíkovský 2002; ''Palaeocryptonyx gaillardia'' Ennouchi 1930; ''Proalector gaillardia'' (Ennouchi 1930) Brodkorb 1964; ''Palaeortyx phasianoides grivensis'' Ballmann 1969a non Lydekker 1893b; '' Taoperdix phasianoides'' (Milne-Edwards 1869) Brodkorb 1964]
***†''Palaeortyx, P. joleaudi'' Ennouchi 1930 [''Plioperdix joleaudi'' (Ennouchi 1930) Brodkorb 1964; ''Palaeortyx depereti'' Ennouchi 1930; ''Plioperdix depereti'' (Ennouchi 1930) Brodkorb 1964]
**†''Procrax, Procrax brevipes'' Tordoff & Macdonald 1957 (middle Eocene? - Early Oligocene) - cracid? gallinuloidid?
**†''Shandongornis'' Yeh 1997 (middle Miocene)
***†''Shandongornis, S. shanwanensis'' Yeh 1997
***†''Shandongornis, S. yinansis'' Hou 2003
**†''Sobniogallus, Sobniogallus albinojamrozi'' Tomek et al. 2014
**†''Taoperdix'' Milne-Edwards 1869 (Late Oligocene) - gallinuloidid?
***†''Taoperdix pesseiti, T. pesseiti'' (Gervais 1862) [''Tetrao pessieti'' Gervais 1862]
***†''Taoperdix miocaena, T. miocaena'' (Gaillard 1939) Ballmann 1969 [''Palaeortyx miocaena'' Gaillard 1939]
**Galliformes gen. et sp. indet. MCZ 342506 (Oligocene) - formerly in ''Gallinuloides''; phasianid?
*†Gallinuloididae Lucas 1900
**†''Gallinuloides, Gallinuloides wyomingensis'' Eastman 1900 (Early/middle Eocene)
**†''Paraortygoides'' Mayr 2000 (London Clay Early Eocene of Walton-on-the-Naze, England – middle Eocene of Messel, Germany)
***†''Paraortygoides messelensis, P. messelensis'' Mayr 2000
***†''Paraortygoides radagasti, P. radagasti'' Dyke & Gulas 2002
*†Paraortygidae Mourer-Chauviré 1992
**†''Pirortyx, Pirortyx major'' (Gaillard 1939) Brodkorb 1964 [''Palaeortyx major'' Gaillard 1939]
**†''Scopelortyx, Scopelortyx klinghardtensis'' Mourer-Chauviré, Pickford & Senut 2015
**†''Paraortyx'' Gaillard 1908 sensu Brodkorb 1964
***†''Paraortyx lorteti, P. lorteti'' Gaillard 1908 [''Palaeortyx cayluxensis'' Lydekker 1891; ''Palaeortyx gaillardi'' Lambrecht 1933; ''Ludiortyx gaillardi'' (Lambrecht 1933) Brodkorb 1964]
***†''Paraortyx brancoi, P. brancoi'' Gaillard 1908
*†Quercymegapodiidae Mourer-Chauviré 1992
**†''Taubacrex, Taubacrex granivora'' dAlvarenga 1988 (Late Oligocene/Early Miocene of Brazil)
**†''Ameripodius'' Alvarenga 1995 (Late Oligocene – Early Miocene of Brazil and France)
***†''Ameripodius silvasantosi, A. silvasantosi'' Alvarenga 1995
***†''Ameripodius alexis, A. alexis'' Mourer-Chauviré 2000
**†''Quercymegapodius'' Mourer-Chauviré 1992 (middle Eocene – Early Oligocene)
***†''Quercymegapodius depereti, Q. depereti'' (Gaillard 1908) [''Palaeocryptonyx depereti'' Gaillard 1908]
***†''Quercymegapodius brodkorbi, Q. brodkorbi'' Mourer-Chauviré 1992
*Megapodidae – megapodes
**†''Ngawupodius, Ngawupodius minya'' Boles & Ivison 1999
**''Prehistoric species of extant genera''
***†''Leipoa gallinacea'' – formerly ''Chosornis, Palaeopelargus, Progura''
*Cracidae – guans and curassows
**†''Boreortalis, Boreortalis laesslei'' Brodkorb 1954 (Early Miocene) – may be same as ''Ortalis''
**''Extant genera present in the fossil record''
***''Ortalis'' (Early Miocene – Recent)
****†''Ortalis, O. affinis'' Feduccia & Wilson 1967
****†''Ortalis, O. phengites'' Wetmore 1923
****†''Ortalis, O. pollicaris'' Miller 1944
****†''Ortalis, O. tantala'' Wetmore 1933
*Odontophoridae – New World quails
**†''Nanortyx, Nanortyx inexpectatus'' Weigel 1963 (Cypress Hills Early Oligocene of North Calf Creek, Canada)
**†''Neortyx, Neortyx peninsularis'' Holman 1961 (Early Pleistocene of Reddick, US)
**†''Miortyx'' Miller 1944 (Rosebud Early Miocene of Flint Hill, US)
***†''Miortyx teres, M. teres'' Miller 1944
***†''Miortyx aldeni, M. aldeni'' Howard 1966
**''Placement unresolved''
***Odontophoridae gen. et sp. indet. KUVP 9393 (White River Early/middle Oligocene of Logan County, US)
**''Prehistoric species of extant genera''
***†''Cyrtonyx cooki'' Gutierrez et al. 1981 (Late Miocene? of Upper Sheep Creek, US)
***†''Callipepla shotwelli'' (Brodkorb 1958) [''Lophortyx shotwelli'' Brodkorb 1958] (Middle Pliocene of McKay Reservoir, US)
***†''Colinus hibbardi'' Wetmore 1944 (Rexroad Late Pliocene of Rexroad, US)
***†''Colinus suilium'' Brodkorb 1959 (Early Pleistocene of SE US)
***†''Colinus'' sp. (Late Pliocene of Benson, US)
***†''Dendrortyx''? sp. (Late Pleistocene of San Josecito Cavern, Mexico)
*Phasianidae – pheasants, quails, partridges, grouse and turkeys
**†''Bantamyx, Bantamyx georgicus'' Kuročkin 1982
**†''Diangallus, Diangallus mious'' Hou 1985
**†''Lophogallus, Lophogallus naranbulakensis'' Zelenkov & Kuročkin 2010
**†''Megalocoturnix, Megalocoturnix cordoni'' Sánchez Marco 2009 (Early Pliocene of Layna, Spain)
**†''Miophasianus'' Brodkorb 1952 [''Miophasianus'' Lambrecht 1933 nomen nudum; ''Miogallus'' Lambrecht 1933]
***†''Miophasianus maxima, M. maxima'' (Lydekker 1893) Brodkorb 1964 [''Palaeortyx maxima'' Lydekker 1893]
***†''Miophasianus altus, M. altus'' (Milne-Edwards 1869) Villalta & Crusafont 1950 [''Phasianus altus'' Milne-Edwards 1869; ''Miogallus altus'' (Milne-Edwards 1869) Mlíkovský 2002; ''Phasianus Desnoyersii'' Milne-Edwards 1869; ''Ardea similis'' Fraas 1870; ''Tantalus milne-edwardsii'' Shufeldt 1896; ''Pseudotantalus'' Milne-Edwardsi (Shufeldt 1896) Sharpe; ''Gallus longaevus'' Ammon 1918; ''Phasianus augustus'' Ammon 1918; ''Botaurites similis'' (Fraas 1870) Lambrecht 1933; ''Miophasianus augustus'' (Ammon 1918) Lambrecht 1933; ''Miophasianus desnoyersii'' (Milne-Edwards 1869) Lambrecht 1933; ''Miogallus longaevus'' (Ammon 1918) Lambrecht 1933; ''Proardea similis'' (Ammon 1870) Gaillard 1939; ''Ibis milne-edwardsi'' (Shufeldt 1896) Brodkorb 1963c]
***†''Miophasianus medius, M. medius'' (Milne-Edwards 1869) Lambrecht 1933 [''Phasianus medius'' Milne-Edwards 1869; ''Palaeoperdix medius'' (Milne-Edwards 1869) Cheneval 2000]
**†''Proagriocharis, Proagriocharis kimballensis'' Martin & Tate 1970 (Kimball Late Miocene/Early Pliocene of Lime Creek, US)
**†''Palaeocryptonyx'' Depéret 1892 [''Chauvireria'' Boev 1997; ''Pliogallus'' Tugarinov 1940b non Gaillard 1939; ''Lambrechtia'' Janossy 1974] (Late Pliocene of SW Europe)
***†''Palaeocryptonyx donnezani, P. donnezani'' Depéret 1892 [''Francolinus Čapeki'' Lambrecht 1933; ''Pliogallus coturnoides'' Tugarinov 1940b; ''Plioperdix coturnoides'' (Tugarinov 1940b) Kretzoi 1955b; ''Francolinus capeki wenzensis'' Jánossy 1974a; ''Francolinus capeki villányiensis'' Jánossy 1974a; ''Francolinus (Lambrechtia) capeki'' (Lambrecht 1933) Jánossy 1974a; ''Alectoris baryosefi'' Tchernov 1980; ''Francolinus wezensis'' (Jánossy 1974a) Jánossy 1981; ''Plioperdix capeki'' (Lambrecht 1933) Mlíkovský 1995b; ''Alectoris donnezani'' (Depéret 1892) Mlíkovský 2002; ''Chauvireria balcanica'' Boev 1997a]
***†''Palaeocryptonyx grivensis, P. grivensis'' Ennouchi 1930
***†''Palaeocryptonyx novaki, P. novaki'' Sánchez Marco 2009
***†''Palaeocryptonyx edwardsi, P. edwardsi'' (Depéret 1887) Ballmann 1969a [''Palaeortyx edwardsi'' Depéret 1887; ''Palaeortyx miocaena'' Gaillard 1939; ''Taoperdix miocaena'' (Gaillard 1939) Ballmann 1969; ''Palaeoperdix edwardsi'' (Depéret 1887) Brodkorb 1964; ''Alectoris edwardsi'' (Depéret 1887) Mlíkovský 2002]
***†''Palaeocryptonyx minor, P. minor'' (Jánossy 1974) [''Francolinus minor'' Jánossy 1974; ''Ammoperdix ponticus'' Tugarinov 1940b; ''Pliogallus ponticus'' (Tugarinov 1940); ''Plioperdix ponticus'' (Tugarinov 1940) Kuročkin 1985; ''Francolinus (Lambrechtia) minor'' Jánossy 1974a]
***†''Palaeocryptonyx subfrancolinus, P. subfrancolinus'' (Jánossy 1976b) [''Francolinus subfrancolinus'' Jánossy 1976b; ''Plioperdix subfrancolinus'' (Jánossy 1976b) Mlíkovský 1995b]
**†''Pliogallus'' Běljajeva 1948
***†''Pliogallus, P. coturnoides'' Běljajeva 1948
***†''Pliogallus, P. csarnotanus'' Kessler & Horváth 2022
**†''Plioperdix'' Kretzoi 1955 [''Pliogallus'' Gaillard 1939 non Tugarinov 1940]
***†''Plioperdix crassipes, P. crassipes'' (Gaillard 1939) [''Pliogallus crassipes'' Gaillard 1939]
***†''Plioperdix kormosi, P. kormosi'' (Gaillard 1939) [''Pliogallus kormosi'' Gaillard 1939]
***†''Plioperdix africana, P. africana'' Mourer-Chauviré & Geraads 2010
***†''Plioperdix hungarica, P. hungarica'' (Jánossy 1991) Zelenkov & Panteleyev 2014 [''Palaeocryptonyx hungaricus'' Jánossy 1991]
**†''Rustaviornis, Rustaviornis georgicus'' Burchak-Abramovich & Meladze 1972
**†''Rhegminornis, Rhegminornis calobates'' Wetmore 1943 (Early Miocene of Bell, US)
**†''Schaubortyx, Schaubortyx keltica'' (Eastman 1905) Brodkorb 1964 [''Taoperdix keltica'' Eastman 1905] (middle Eocene – Early Oligocene)
**†''Shandongornis'' Yeh 1997
***†''Shandongornis, S. shanwanensis'' Yeh 1997
***†''Shandongornis, S. yinansis'' Hou 2003
**†''Shanxiornis, Shanxiornis fenyinis'' Wang et al. 2006
**†''Tologuica'' Zelenkov & Kuročkin 2009 (Middle Miocene of Sharga, Mongolia)
***†''Tologuica aurorae, T. aurorae'' Zelenkov & Kuročkin 2009
***†''Tologuica karhui, T. karhui'' Zelenkov & Kuročkin 2009
**''Placement unresolved''
***Tetraoninae gen. et sp. indet. (Sajóvölgyi Middle Miocene of Mátraszõlõs, Hungary)
***Meleagridae gen. et sp. indet. (Late Miocene of Westmoreland County, US)
***''"Tympanuchus stirtoni, Tympanchus" stirtoni'' Miller 1944 (Early Miocene of South Dakota)
***''"Tympanuchus" lulli'' Shuefeldt 1915 (Pleistocene? of New Jersey)
**''Extant genera present in the fossil record''
***''Coturnix'' (Late Oligocene – Recent)
***''Bambusicola'' (Late Miocene – Recent)
****†''Bambusicola, B. dalianensis'' (Hou 1990) [''Tetrastes dalianensis'' Hou 1990]
****†''Bambusicola, B. nini'' Sánchez Marco 2009
****†''Bambusicola, B. praebonasia'' (Jánossy 1974) Bocheński 1991 [''Tetrastes praebonasia'' Jánossy 1974]
***''Phasianus'' (Late Miocene)
****†''Phasianus, P. lufengia'' Hou 1985
****†''Phasianus, P. yanshansis'' Huang & Hou 1984
***''Crossoptilon''
****†''Crossoptilon, C. jiai'' Hou 1982
***''Lophura''
****†''Lophura, L. wayrei'' Harrison & Walker 1982
***''Gallus (biology), Gallus'' (Late Miocene/Early Pliocene – Recent)
****†''Gallus (biology), G. aesculapii'' Jánossy 1976
****†''Gallus (biology), G. beremendensis'' Jánossy 1977
****†''Gallus (biology), G. georgicus''
****†''Gallus (biology), G. imereticus''
****†''Gallus (biology), G. karabachensis'' Baryšnikov & Potapova 1995 (Giant junglefowl)
****†''Gallus (biology), G. kudarensis'' Burčak-Abramovič & Potapova
****†''Gallus (biology), G. meschtscheriensis''
****†''Gallus (biology), G. tamanensis''
***''Lagopus'' (Early? Pliocene – Recent)
****†''Lagopus, L. atavus'' Jánossy 1974
****†''Lagopus, L. balcanicus'' Boev 1995
****†''Lagopus, L. lagopus noaillensis'' Mourer-Chauviré 1975
****†''Lagopus, L. muta correzensis'' Mourer-Chauviré 1975
***''Meleagris'' (Early Pliocene – Recent)
****†''Meleagris, M. celer'' Marsh 1872
****†''Meleagris, M. tridens'' Wetmore 1931
****†''Meleagris, M. progenes'' (Brodkorb 1964) [''Agriocharis progenes'' Brodkorb 1964]
****†''Meleagris, M. leopoldi''
****†''Meleagris, M. anza'' (Howard 1963) [''Agriocharis anza'' Howard 1963]
****†''Meleagris, M. crassipes'' Rea 1980
***''Pavo (genus), Pavo'' (Early Pliocene – Recent)
****†''Pavo (genus), P. bravardi'' (Gervais 1849) [''Gallus moldovicus'' Abramovich, Ganea & Shushpanov 1993]
***''Tetrao'' (Early Pliocene – Recent)
****†''Tetrao, T. conjugens'' Jánossy 1974
****†''Tetrao, T. macropus'' Jánossy 1976
****†''Tetrao, T. partium'' (Kretzoi 1962)
****†''Tetrao, T. praeurogallus'' Jánossy 1969
****†''Tetrao, T. rhodopensis'' Boev 1998
***''Lyrurus''
****†''Lyrurus, Lyrurus tetrix longipes''
***''Francolinus'' (Late Pliocene – Recent)
****†''Francolinus, F. capeki'' Jánossy 1974
****†''Francolinus, F. villanyiensis'' Jánossy 1974
****†''Francolinus, F. wenzensis'' Jánossy 1974
***''Perdix'' (Early Pleistocene – Recent)
****†''Perdix, P. inferna'' (Kurochkin 1985) [''Lophura inferna'' Kuročkin 1985]
****†''Perdix, P. margaritae'' Kuročkin 1985
****†''Perdix, P. palaeoperdix'' Mourer-Chauviré 1975
***''Syrmaticus''
****†''Syrmaticus, S. kozlovae'' Kuročkin 1985
***''Bonasa'' (Early/Middle Pleistocene – Recent)
****†''Bonasa, B. dalianensis'' (Hou 1990) [''Tetrastes dalianensis'' Hou 1990]
****†''Bonasa, B. nini'' Sánchez Marco 2009
****†''Bonasa, B. praebonasia'' (Jánossy 1974) Bocheński 1991 [''Tetrastes praebonasia'' Jánossy 1974]
***''Dendragapus'' (Late Pleistocene – Recent)
****†''Dendragapus, D. gilli'' (Brodkorb 1964) [''Palaeotetrix gilli'' Brodkorb 1964]
****†''Dendragapus, D. lucasi'' Jehl 1969
***''Alectoris''
****†''Alectoris, A. baryosefi'' Tchernov 1980
****†''Alectoris, A. bavarica'' Ballmann 1969
****†''Alectoris, A. peii'' Hou 1982
Charadriiformes
Gulls, auks, shorebirds
*Basal and unresolved taxa
**Charadriiformes gen. et sp. indet. (Late Cretaceous) – burhinid? basal?
**''"Morsoravis"'' (Late Paleocene/Early Eocene) – a ''nomen nudum''?
**''Jiliniornis'' (middle Eocene) – charadriid?
**''Boutersema'' (Early Oligocene) – glareolid?
**''Turnipax'' (Early Oligocene) – turnicid?
**''Elorius'' (Early Miocene)
**''"Larus desnoyersii'' (Early Miocene of SE France) – larid? stercorarid?
**''"Larus pristinus'' (John Day Early Miocene of Willow Creek, US) – larid?
**Charadriiformes gen. et spp. indet. (Early/Middle Miocene) – several species, 1 probably larid
**Charadriiformes gen. et sp. indet. (Middle Miocene)
**''"Totanus" teruelensis'' (Late Miocene of Los Mansuetos, Spain) – scolopacid? larid?
**''"Actitis" balcanica'' (Late Pliocene of Varshets, Bulgaria) – scolopacid? charadriid?
*Scolopacidae – waders and snipes
**''Paractitis'' (Early Oligocene)
**''Mirolia'' (Middle Miocene)
**''Placement unresolved''
***Scolopacidae gen. et sp. indet. (Middle – Late Miocene)
***Scolopacidae gen. et sp. indet. (Early Pliocene)
**''Extant genera present in the fossil record''
***''Limosa'' (Late Eocene? – Recent)
***''Tringa'' (Late Eocene/Early Oligocene? – Recent) – includes ''Totanus''
***''Gallinago'' (Late Miocene/Early Pliocene – Recent)
***''Scolopax'' (Early/Middle Pliocene? – Recent)
***''Phalaropus'' (Middle Pliocene – Recent)
***''Actitis'' (Late Pliocene – Recent)
***''Numenius (bird), Numenius'' (Late Pleistocene – Recent) – includes ''Palnumenius''
*Jacanidae – jacanas
**''Nupharanassa'' (Early Oligocene)
**''Janipes''
**''Prehistoric species of extant genera''
***''Jacana farrandi''
*Laridae – gulls
**Laridae gen. et sp. indet. (Early Oligocene)
**''Laricola'' (Late Oligocene/Early Miocene) – larid? Formerly ''"Larus" elegans'' and ''"L." totanoides''
**''Gaviota (genus), Gaviota'' (Middle/Late Miocene)
**''Extant genera present in the fossil record''
***''Larus'' (Middle Miocene – Recent)
*Alcidae – auks
**''Hydrotherikornis'' (Late Eocene)
**''Pseudocepphus'' (Middle – Late Miocene)
**''Petralca'' (Early –? Late Oligocene)
**''Miocepphus'' (Middle Miocene)
**''Alcodes'' (Late Miocene)
**''Praemancalla'' (Late Miocene – Early Pliocene)
**''Mancalla'' (Late Miocene – Early Pleistocene)
**''Extant genera present in the fossil record''
***''Cepphus'' (Late Miocene – Recent)
***''Rhinoceros auklet, Cerorhinca'' (Late Miocene – Recent)
***''Uria'' (Late Miocene – Recent)
***''Aethia '' (Late Miocene – Recent)
***''Razorbill, Alca'' (Late Miocene/Early Pliocene – Recent)
***''Synthliboramphus'' (Late Miocene/Early Pliocene – Recent)
***''Fratercula'' (Early Pliocene – Recent)
***''Pinguinus'' (Early Pliocene – Recent)
***''Brachyramphus'' (Late Pliocene – Recent)
***''Ptychoramphus'' (Late Pliocene – Recent)
*Stercorariidae – skuas and jaegers
**''Prehistoric species of extant genera''
***''Stercorarius'' sp. (Middle Miocene)
***''Stercorarius shufeldti'' (Fossil Lake Middle Pleistocene of WC US)
***''Prehistoric subspecies of extant species''
****''Stercorarius pomarinus philippi''
*Glareolidae – pratincoles
**''Paractiornis'' (Agate Fossil Beds Early Miocene of Sioux County, US)
**''Mioglareola'' (Early Miocene of Czech Republic) - formerly ''"Larus" dolnicensis''
**''Prehistoric species of extant genera''
***''Glareola neogena''
*Burhinidae – thick-knees
**''Prehistoric species of extant genera''
***''Burhinus lucorum'' (Early Miocene)
***''Burhinus aquilonaris''
***''Burhinus'' sp. (Cuba, West Indies)
***''Burhinus'' sp. (Late Pleistocene of Las Higueruelas, Spain)
***''Prehistoric subspecies of extant species''
****''Burhinus bistriatus nanus'' (Bahamas, West Indies)
*Charadriidae – plovers
**''Limicolavis'' (John Day Early Miocene of Malheur County, US)
**''Viator (bird), Viator'' (Late Pleistocene of Talara, Peru) – may be synonym of ''Vanellus'' (or ''Belanopteryx'' if valid)
**''Extant genera present in the fossil record''
***''Vanellus'' (Middle/Late Pleistocene – Recent) – includes ''Belanopteryx''
**''Additional prehistoric species of extant genera''
***''Oreopholus orcesi''
*Recurvirostridae – avocets
**''Extant genera present in the fossil record''
***''Himantopus'' (Late Miocene – Recent)
**''Additional prehistoric species of extant genera''
***''Recurvirostra sanctaeneboulae''
†Gastornithiformes (extinct)
The diatrymas, a group of huge flightless Paleogene birds of unclear affinities. Traditionally placed within the Gruiformes, they are usually considered a distinct order nowadays and appear closer to the Anseriformes.
*Gastornithidae
**†''Ornitholithes biroi'' Dughi & Sirugue 1969 [ootaxa- Gastornis?]
**†''Ornithoformipes controversus'' [ichnotaxa]
**†''Rivavipes giantess'' [ichnotaxa]
**†Gasthornithidae gen. et sp. indet. (Paleocene) – possibly ''Gastornis''
**†Gasthornithidae gen. et sp. indet. YPM PU 13258 (Early Eocene) – possibly juvenile ''Gastornis giganteus''
**†''"Diatryma" cotei'' Galliard 1936 (middle-Late Eocene)
**†''Gastornis'' Hébert 1855 [''Diatryma'' Cope 1876; ''Barornis'' Marsh 1894; ''Omorhamphus'' Sinclair 1928; ''Palaeornis'' Bonaparte 1856; ''Macrornis'' Seeley 1866; ''Zhongyuanus'' Hou 1980] (Late Paleocene – middle Eocene)
***†''Gastornis parisiensis, G. parisiensis'' Hébert 1855 [''Gastornis edwardsii'' Lemoine 1878; ''Gastornis klaasseni'' Newton 1885]
***†''Gastornis giganteus, G. giganteus'' (Cope 1876) [''Diatryma gigantea'' Cope 1876; ''Barornis regens'' Marsh 1894; ''Diatryma regens'' (Marsh 1894); ''Omorhamphus storchii'' Sinclair 1928 (juvenile); ''Diatryma steini'' Matthew & Granger 1917; ''Diatryma ajax'' Shufeldt 1913; ''Gastornis ajax'' (Shufeldt 1913) Brodkorb 1967]
***†''Gastornis, G. sarasini'' (Schaub 1929) Mlíkovský 2002 [''Diatryma sarasini'' Schaub, 1929b; ''Diatryma geiselensis'' Fischer 1978; ''Gastornis geiselensis'' (Fischer 1978)]
***†''Gastornis, G. russeli'' Martin 1992
***†''Gastornis, G. xichuanensis'' (Hou 1980) Buffetaut 2013 [''Zhongyuanus xichuanensis'' Hou 1980]
Gruiformes
The group that includes modern Rallidae, rails and Crane (bird), cranes. Probably paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
.
*Placement unresolved
**†''Rupelrallus'' (Early Oligocene)– rallid? parvigruid?
***†''Rupelrallus belgicus, R. belgicus''
***†''Rupelrallus saxoniensis, R. saxoniensis''
**†"Gruiformes" gen. et sp. indet. MNZ S42623 (Early/Middle Miocene)– Aptornithidae?
*†Songziidae possibly a ''nomen nudum''
**†''Songzia'' (Eocene) possibly a ''nomen nudum''
***†''Songzia, S. heidangkouensis''
***†''Songzia, S. acutunguis''
*†Eogruidae
**†''Sonogrus, Sonogrus gregalis'' (Ergilin Dzo Late Eocene/Early Oligocene of Khor Dzan, Mongolia)
**†''Eogrus'' [''Progrus'' ] (Irdin Manha Middle/Late Eocene – Tung Gur Late Miocene/Early Pliocene of Mongolia)
***†''Eogrus aeola, E. aeola''
***†''Eogrus crudus, E. crudus''
***†''Eogrus turanicus, E. turanicus'' [''Progrus turanicus'' ]
***†''Eogrus wetmorei, E. wetmorei''
**†Ergilornithinae
**†''Proergilornis, Proergilornis minor'' (Early/middle Oligocene of Ergil-Obo, Mongolia)
**†''Ergilornis'' (Early/middle Oligocene of Ergil-Obo, Mongolia)
***†''Ergilornis rapidus, E. rapidus''
***†''Ergilornis minor, E. minor''
**†''Amphipelargus'' [''Urmiornis'' ]
***†''Amphipelargus brodkorbi, A. brodkorbi'' [''Urmiornis brodkorbi'' ]
***†''Amphipelargus majori, A. majori'' [''Amphipelargus maraghanus'' ; ''Urmiornis maraghanus'' ] [treated as ciconiid by Louchart et al. 2005]
***†''Amphipelargus dzabghanensis, A. dzabghanensis''
***†''Amphipelargus cracrafti, A. cracrafti'' [''Urmiornis cracrafti'' ]
***†''Amphipelargus ukrainus, A. ukrainus'' [''Urmiornis ukrainus'' ]
***†''Amphipelargus orientalis, A. orientalis'' [''Urmiornis orientalis'' ]
*†Geranoididae
**†''Eogeranoides, Eogeranoides campivagus'' (Willwood Early Eocene of Foster Gulch, US)
**†''Geranoides, Geranoides jepseni'' (Willwood Early Eocene of South Elk Creek, US)
**†''Geranodornis, Geranodornis aenigma'' (Bridger middle Eocene of Church Buttes, US)
**†''Galligeranoides, Galligeranoides boriensis''
**†''Palaeophasianus'' (Willwood Early Eocene of Bighorn County, US)
***†''Palaeophasianus meleagroides, P. meleagroides''
***†''Palaeophasianus incompletus, P. incompletus''
**†''Paragrus'' (Early Eocene of WC US)
***†''Paragrus prentici, P. prentici'' [''Gallinuloides prentici'' ; ''Grus prentici'' ]
***†''Paragrus shufeldti, P. shufeldti''
*†Parvigruidae Mayr 2005
**†''Parvigrus, Parvigrus pohli'' (Early Oligocene of Pichovet, France)
*Aramidae
**†''Badistornis, Badistornis aramus'' (middle Oligocene)– aramid?
**''Extant genera present in the fossil record''
***''Aramus paludigrus'' (Middle Miocene of La Venta, Colombia)- Aramid
*Gruidae – cranes
**†''Camusia, Camusia quintanai'' (Late Miocene of Menorca, Mediterranean)
**†''Aramornis'' (Middle Miocene) – gruid? aramid?
***†''Aramornis longurio, A. longurio''
***†''Aramornis robalearica, A. robalearica''
***†''Aramornis crataegensis, A. crataegensis''
**†''Palaeogrus'' (middle Eocene of Germany and Italy – Middle Miocene of France)
***†''Palaeogrus, P. princeps'' [''Ornitocnemus robustus'' ; ''Grus princeps'' ]
***†''Palaeogrus, P. hordwelliensis'' [''Grus hordwellianus'' ; ''Ornitocnemus hordwelliensis'' ]
***†''Palaeogrus, P. mainburgensis''
**†''Geranopsis, Geranopsis hastingsiae'' Hordwell Late Eocene– Early Oligocene of England)– gruiform (gruid?) or anseriform (anseranatid?)?
**†''Eobalearica, Eobalearica tugarinovi'' (Ferghana Late? Eocene of Ferghana, Uzbekistan)
**†''Pliogrus'' (Early Pliocene of Eppelsheim, Germany)
**†''Probalearica, Probalearica mongolica'' (Late Oligocene? – Middle Pliocene) gruid? A '' nomen dubium''?
**Gruidae gen. et sp. indet. – formerly ''Grus conferta'' (Late Miocene/Early Pliocene of Contra Costa County, US)
**''Extant genera present in the fossil record''
***''Balearica'' (Early Miocene – Recent)
****†''Balearica, B. crataegensis'' [''Probalearica crataegensis'' ]
****†''Balearica, B. excelsa'' [''Palaeogrus excelsa'' ; ''Grus excelsa'' ; ''Ornitocnemus excelsus'' ]
****†''Balearica exigua, B. exigua''
****†''Balearica rummeli, B. rummeli'' [''Basityto rummeli'' ] (giant barn-owl)
***''Grus (bird), Grus'' (Middle/Late Miocene – Recent)
****†''Grus (bird), G. bogatshevi''
****†''Grus (bird), G. haydeni''
****†''Grus (bird), G. latipes'' [''Baeopteryx latipes'']
****†''Grus (birds), G. moldavica'' [''Probalearica moldavica'' ]
****†''Grus (birds), G. nannodes''
****†''Grus pagei, G. pagei''
****†''Grus (birds), G. pentelici'' [''Pliogrus pentelici'' ; ''Grus afghana, G. afghana'' ]
****†''Grus (birds), G. primigenia''
****†''Grus (bird), G. melitensis'' (Maltese crane)
*Rallidae – rails
**†''Aletornis'' [''Protogrus'' ] (middle Eocene)
***†''Aletornis marshi, A. marshi'' [''Grus marshi'' ; ''Protogrus marshi'' ]
***†''Aletornis bellus, A. bellus''
***†''Aletornis gracilis, A. gracilis''
***†''Aletornis nobilis, A. nobilis'' [''Grus nobilis'' ; ''Protogrus nobilis'' ]
***†''Aletornis pernix, A. pernix''
**†''Australlus''
***†''Australlus disneyi, A. disneyi'' [''Gallinula disneyi'' ]
***†''Australlus gagensis, A. gagensis''
**†''Baselrallus intermedius''
**†''Belgirallus'' (Early Oligocene)
***†''Belgirallus minutus, B. minutus''
***†''Belgirallus oligocaenus, B. oligocaenus''
**†''Creccoides, Creccoides osbornii'' (Late Pliocene/Early Pleistocene)
**†''Eocrex'' (Early Eocene)
***†''Eocrex primus, E. primus''
***†''Eocrex tagusevae, E. tagusevae''
**†''Euryonotus'' (Pleistocene)– rallid?
***†''Euryonotus uspallatensis, E. uspallatensis''
***†''Euryonotus brachypterus, E. brachypterus''
***†''Euryonotus argentinus, E. argentinus''
**†''Fulicaletornis, Fulicaletornis venustus'' [''Aletornis venustus'' ] (middle Eocene)
**†''Ibidopsis, Ibidopsis hordwelliensis'' (Late Eocene)
**†''Latipons'' (middle Eocene)
***†''Latipons gardneri, L. gardneri''
***†''Latipons robinsoni, L. robinsoni''
**†''Miofulica, Miofulica dejardini'' [''Fulica dejardinii'' ] (Middle Miocene)
**†''Miorallus, Miorallus major'' [''Rallus major'' ] (Middle – Late Miocene)
**†''Palaeoaramides'' [''Pararallus'' ; ''Tertiariaporphyrula'' ] (Late Oligocene/Early Miocene – Late Miocene)
***†''Palaeoaramides lungi, P. lungi'' [''Tertiariaporphyrula lungi'' ; ''Mioporphyrula lungi'' ]
***†''Palaeoaramides minutus, P. minutus''
***†''Palaeoaramides tugarinovi, P. tugarinovi'' [''Tertiariaporphyrula tugarinovi'' ]
***†''Palaeoaramides christyi, P. christyi'' [''Rallus christyi'' ; ''Rallus eximius'' ; ''Palaeoaramides eximius'' ]
***†''Palaeoaramides minor, P. minor'' [''Rallus minor'']
***†''Palaeoaramides beaumontii, P. beaumontii'' [''Rallus beaumontii'' ; ''Pararallus beaumontii'' ; ''Tertiariaporphyrula beaumonti'' ; ''Rallus dispar'' ; ''Pararallus dispar'' ; ''Palaeoaramides dispar'' ]
**†''Palaeorallus'' Wetmore 1931 (Early Eocene)
***†''Palaeorallus troxelli, P. troxelli''
***†''Palaeorallus brodkorbi, P. brodkorbi''
**†''Paraortygometra, Paraortygometra porzanoides'' [''Microrallus'' ; ''Rallus porzanoides'' ; ''Microrallus fejfari'' ] (Late Oligocene/?Early Miocene –? Middle Miocene)
**†''Parvirallus'' (Early – middle Eocene)
***†''Parvirallus gracilis, P. gracilis''
***†''Parvirallus bassetti, P. bassetti''
***†''Parvirallus gassoni, P. gassoni''
***†''Parvirallus medius, P. medius''
**†''Pastushkinia, Pastushkinia zazhigini'' [''Crex zazhigini'' ]
**†''Pleistorallus, Pleistorallus flemingi'' (Fleming's rails)
**†''Quercyrallus'' (Late Eocene –? Late Oligocene)
***†''Quercyrallus, Q. ludianus'' [''Quercyrallus intermedius'' ; ''Rallus adelus'' ; ''Ludiortyx adelus'' ]
***†''Quercyrallus arenarius, Q. arenarius'' [''Rallus arenarius'' ]
***†''Quercyrallus quercy, Q. quercy''
**†''Rallicrex'' (Middle/Late Oligocene)
***†''Rallicrex litkensis, R. litkensis''
***†''Rallicrex polgardiensis, R. polgardiensis''
***†''Rallicrex kolozsvarensis, R. kolozsvarensis''
**†''Rhenanorallus, Rhenanorallus rhenanus''
**†''Wanshuina, Wanshuina lii''
**†''Youngornis'' (Middle Miocene)
***†''Youngornis gracilis, Y. gracilis''
***†''Youngornis qiluensis, Y. qiluensis''
**''Placement unresolved''
***Rallidae gen. et sp. indet. (Late Oligocene)
***Rallidae gen. et spp. indet. (Early/Middle Miocene)– several species
***Rallidae gen. et sp. indet. (Middle Miocene)
***Rallidae gen. et sp. indet. (Late Miocene)
***Rallidae gen. et sp. indet. UMMP V55013/-14; UMMP V55012/V45750/V45746 (Late Pliocene)
***Rallidae gen. et sp. indet. UMMP V29080 (Late Pliocene)
***Rallidae gen. et sp. indet. (Bermuda, West Atlantic)
**''Extant genera present in the fossil record''
***''Rallus'' [†''Epirallus'' ] (Middle Miocene – Recent)
****†''Rallus, R. auffenbergi'' [''Porzana auffenbergi'' ]
****†''Rallus, R. cyanocavi''
****†''Rallus, R. ibycus''
****†''Rallus, R. lacustris''
****†''Rallus, R. minutus''
****†''Rallus, R. natator'' [''Epirallus natator'' ]
****†''Rallus, R. phillipsi''
****†''Rallus, R. prenticei''
****†''Rallus, R. recessus''
****†''Rallus, R. richmondi'' [''Rallus dubius'' ]
***''Porzana'' (Middle? Miocene – Recent)
****†''Porzana, P. botunensis''
****†''Porzana, P. estramosi''
****†''Porzana, P. kretzoii''
****†''Porzana, P. matraensis''
****†''Porzana, P. piercei''
****†''Porzana, P. risillus'' [''Rallus risillus'' ]
***''Gallinula'' (Late Oligocene – Recent)
****†''Gallinula, G. balcanica''
****†''Gallinula brodkorbi, G. brodkorbi''
****†''Gallinula, G. gigantea''
****†''Gallinula, G. kansarum'' [''Fulica americana'' (in partim)]
***''coots, Fulica'' (Early Pliocene – Recent)
****†''F. infelix''
****†''Fulica podagrica, F. podagrica''
****†''F. stekelesi''
***''Laterallus''
****†''Laterallus, L. guti''
****†''Laterallus, L. insignis'' (Rexroad Late Pliocene of Rexroad, US)
****†''Laterallus, L. sp.'' (Late Pliocene of Macasphalt Shell Pit, US)
***''Coturnicops''
****†''Coturnicops, C. avita'' (Glenns Ferry Late Pliocene of Hagerman, US)
***''Porphyrio''
****†''Porphyrio, P. parvus''
Eurypygiformes
*†Aptornithidae
**†''Aptornis proasciarostratus''
*Eurypygidae
**†''Eoeurypyga, Eoeurypyga olsoni''
*†Messelornithidae – Messel pit, Messel-birds
**†''Pellornis, Pellornis mikkelseni''
**†''Itardiornis, Itardiornis hessae''
**†'' Messelornis''
***†''Messelornis, M. russelli'' ["Messelornis" russelli sensu Mayr 2007]
***†''Messelornis, M. nearctica''
***†''Messelornis, M. cristata''
Cariamiformes
*†''Itaboravis, Itaboravis elaphrocnemoides''
*†'' Walbeckornis creber''
*†Qianshanornithidae
**†''Qianshanornis, Qianshanornis rapax''
*†Salmilidae
**†''Salmila, Salmila robusta''
*†Ameghinornithidae
**†''Strigogyps'' [''Aenigmavis'' ; ''Ameghinornis'' ; ''Eocathartes'' ; ''Geiseloceros'' ]
***†''Strigogyps, S. dubius''
***†''Strigogyps, S. robustus'' [''Eocathartes robustus'' ; ''Geiseloceros robustus'' ] (middle Eocene of Germany)
***†''Strigogyps sapea, S. sapea'' [''Aenigmavis sapea'' ; ''Ameghinornis minor'' ]
*†Bathornithidae
**†''Eutreptornis, Eutreptornis uintae'' (Uinta Late Eocene of Ouray Agency, US)
**†''Neocathartes, Neocathartes grallator'' [''Eocathartes'' ; ''Eocathartes grallator'' ; ''Bathornis grallator'' ] (Late Eocene)
**†''Paracrax'' (Early/middle Oligocene of Gerry's Ranch, US – Brule Late Oligocene of South Dakota, US)
***†''Paracrax, P. antiqua'' [''Meleagris antiquus'' ; ''Oligocorax mediterraneus'' ; ''Phalacrocorax mediterraneus'' ]
***†''Paracrax, P. wetmorei''
***†''Paracrax, P. gigantea''
**†Bathornithidae gen. nov. (Early – middle Oligocene of C US) – formerly ''Bathornis celeripes'' and ''B. cursor''
**†''Bathornis'' (Early Oligocene – Early Miocene of C US)
***†''Bathornis, B. minor''
***†''Bathornis veredus, B. veredus'' [''Palaeogyps prodromus'' ] (Early Oligocene of WC North America)
***†''Bathornis, B. fax'' [''Palaeocrex fax'' Wetmore 1927]
***†''Bathornis, B. geographicus''
***†''Bathornis, B. fricki''
*†Idiornithidae [Orthocnemidae]
**†''Dynamopterus'' [''Idiornis'' ; ''Orthocnemus'' ] (middle Eocene of Messel, Germany –? Quercy Phosphorites middle Oligocene of Quercy, France)
***†''Idiornis cursor, I. cursor'' [''Orthocnemus cursor'' ; ''Orthocnemus major'' ; ''Idiornis major'' ]
***†''Idiornis gallicus, I. gallicus'' [''Filholornis gravis'' ; ''Orthocnemus gallicus'' ; ''Agnopterus gallicus'' ]
***†''Idiornis gracilis, I. gracilis'' [''Elaphrocnemus gracilis'' ; ''Agnopterus gracilis'' ]
***†''Idiornis minor, I. minor'' [''Orthocnemus minor'' ; ''Agnopterus minor'' ]
***†''Dynamopterus anthracinus, D. anthracinus'' [''Idiornis anthracinus'' ]
***†''Dynamopterus tuberculata, D. tuberculata'' [''Idiornis tuberculata'' ]
***†''Dynamopterus gaillardi, D. gaillardi'' [''Idiornis gaillardi'' ]
***†''Dynamopterus itardiensis, D. itardiensis'' [''Idiornis itardiensis'' ; ''Agnopterus itardiensis'' ]
**†''Talantatos'' [''Elaphrocnemus'' ; ''Filholornis'' ] (Late Eocene/Early Oligocene of France)
***†''Talantatos bordkorbi, T. bordkorbi'' [''Elaphrocnemus bordkorbi'' ]
***†''Talantatos crex, T. crex'' [''Elaphrocnemus crex'' ]
***†''Talantatos phasianus, T. phasianus'' [''Elaphrocnemus phasianus'' ; ''Filholornis debilis'' ; ''Filholornis paradoxa'' ]
***†''Talantatos fossilis, T. fossilis'' [''Tantalus fossilis'' ; ''Orthocnemus major'' ; ''Idiornis major'' ; ''Telecrex peregrinus'' ]
***†''Talantatos proudlocki, T. proudlocki'' [''Percolinus proudlocki'' ]
**†''Gypsornis'' [''Percolinus'' ] (Montmartre Late Eocene of Montmartre, France)
***†''Gypsornis cuvieri, G. cuvieri''
***†''Gypsornis venablesi, G. venablesi'' [''Percolinus venablesi'' ]
**†''Propelargus'' [''Oblitavis'' ] (Late Eocene/Early Oligocene)
***†''Propelargus edwardsi, P. edwardsi''
***†''Propelargus insolitus, P. insolitus'' [''Oblitavis insolitus'' ]
***†''Propelargus cayluxensis, P. cayluxensis''
**†''Occitaniavis elatus'' [''Geranopsis'' ; ''Geranopsis elatus'' ; ''Propelargus elatus'' ]
**†''Ibidopodia'' (Late Oligocene/Early Miocene of France) – threskiornithid?
***†''Ibidopodia chavrochensis, I. chavrochensis''
***†''Ibidopodia minuta, I. minuta''
***†''Ibidopodia palustris, I. palustris''
*†Phorusrhacidae – Terror birds
**†''Lavocatavis, Lavocatavis africana''
**†''Patagorhacos, Patagorhacos terrificus''
**†''Eleutherornis'' (Late Eocene)
***†''Eleutherornis helveticus, E. helveticus''
***†''Eleutherornis cotei, E. cotei''
**†''Paleopsilopterus, Paleopsilopterus itaboraiensis'' (Middle Paleocene)
**†''Procariama, Procariama simplex'' (Late Miocene – Early Pliocene)
**†''Psilopterus'' [''Pelecyornis'' ; ''Staphylornis'' ] (middle Oligocene – Late Miocene)
***†''Psilopterus, P. bachmanni'' [''Patagornis bachmanni'' ; ''Psilopterus communis'' ; ''Psilopterus intermedius'' ; ''Phororhacos delicatus'' ; ''Pelecyornis puerredonensis'' ]
***†''Psilopterus, P. lemoinei'' [''Patagornis lemoinei'' ; ''Psilopterus australis'' ; ''Pelecyornis tubulatus'' ; ''Phorohacos modicus'' ; ''Staphylornis gallardoi'' ; ''Staphylornis erythacus'' ; ''Pelecyornis tenuirostris'' ]
***†''Psilopterus, P. affinis'' [''Phororhacos affinis'' ]
***†''Psilopterus, P. colzecus''
**†''Llallawavis, Llallawavis scagliai''
**†''Mesembriornis'' [''Paleociconia'' ; ''Prophorohacos'' ; ''Hermosiornis'' ] (Late Miocene – Late Pliocene)
***†''Mesembriornis, M. milneedwardsi'' [''Paleociconia australis'' ; ''Driornis pampeanus'' ; ''Hermosiornis milneedwardsi'' ; ''Hermosiornis rapax'' ; ''Prophororhacos australis'' ]
***†''Mesembriornis, M. incertus'' [''Prophororhacos incertus'' ]
**†''Patagornis, Patagornis marshi'' [''Morenomercatia'' ; ''Tolmodus'' ; ''Tolmodus inflatus'' ; ''Phorusrhacos inflatus'' ; ''Paleociconia cristata'' ; ''Morenomerceraria cristata'' ] (Early – Middle Miocene)
**†''Andrewsornis, Andrewsornis abbotti'' (middle – Late Oligocene)
**†''Andalgalornis, Andalgalornis steulleti'' [''Phororhacos steulleti'' Kraglijevič 1931; ''Phororhacos deautieri'' ; ''Andalgalornis ferox'' ] (Late Miocene – Early Pliocene)
**†''Phorusrhacos, Phorusrhacos longissimus'' [''Stereornis'' ; ''Darwinornis'' ; ''Owenornis'' ; ''Titanornis'' ; ''Callornis'' ''Calornis'' ; ''Eucallornis'' ; ''Liornis'' ; ''Phororhacos longissimus'' ; ''Phororhacos platygnathus'' ; ''Phororhacos sehuensis'' ; ''Stereornis rollieri'' ; ''Stereornis gaundryi'' ; ''Mesembriornis studeri'' ; ''Mesembriornis quatrefragesi'' ; ''Darwinornis copei'' ; ''Darwinornis zittelli'' ; ''Darwinornis socialis'' ; ''Owenornis affinis'' ; ''Owenornis lydekkeri'' ; ''Titanornis mirabilis'' ; ''Callornis giganteus'' ; ''Eucallornis giganteus'' ; ''Liornis floweri'' ; ''Liornis minor'' ] (Early – Middle Miocene)
**†''Devincenzia, Devincenzia pozzi'' [''Onactornis'' Cabrera 1939; ''Phorohacos pozzi'' ; ''Phorohacos longissimus mendocinus'' ; ''Devincenzia gallinali'' ; ''Onactornis depressus'' ; ''Onactornis pozzi'' ; ''Onactornis mendocinus'' ] (Late Miocene – Early Pliocene)
**†''Titanis, Titanis walleri'' (Early – Late Pliocene)
**†''Kelenken, Kelenken guillermoi'' (Early – Late Pliocene)
**†''Physornis, Physornis fortis'' [''Aucornis'' ; ''Aucornis euryrhynchus'' ] (middle – Late Oligocene)
**†''Paraphysornis, Paraphysornis brasiliensis'' [''Physornis brasiliensis'' ] (Late Oligocene/Early Miocene)
**†''Brontornis, Brontornis burmeisteri'' [''Rostrornis'' ; ''Rostrornis floweri'' ; ''Brontornis platyonyx'' ] (Early – Middle Miocene)
*Cariamidae – seriemas
**†'' Riacama caliginea''
**†''Pseudolarus, Pseudolarus guaraniticus'' [''Pseudogavia'' ; ''Ameghinia'' ]
**†'' Smiliornis penetrans''
**†''Noriegavis, Noriegavis santacrucensis'' [''Cariama santacrucensis'' ]
**''Prehistoric species of extant genera''
***''Chunga (bird), Chunga''
****†''Chunga (birds), C. incerta''
Otidiformes
*Otididae – bustards
**†''Gryzaja, Gryzaja odessana'' Zubareva 1939 [''Chlamydotis pliodeserti'' Serebrovskyj 1941; ''Otis gryzaja'' Vojinstvens'kyj 1967]
**†''Ioriotis, Ioriotis gabunii'' Burchak-Abramovich & Vekua 1981
**†''Miootis, Miootis compactus'' Umanskaya 1979
**†''Pleotis, Pleotis liui'' Hou 1982
**''Prehistoric species of extant genera''
***''Chlamydotis'' Brodkorb 1967 [''Otis affinis'' Lydekker 1891a]
****†''Chlamydotis, C. affinis'' (Lydekker)
****†''Chlamydotis, C. mesetaria'' Sánchez Marco 1990
***''Tetrax''
****†''Tetrax, T. paratetrax'' (Bocheński & Kuročkin 1987) [''Otis paratetrax'' Bocheński & Kuročkin 1987]
***''Otis (bird), Otis''
****†''Otis (bird), O. bessarabicus'' Kessler & Gal 1996
****†''Otis (bird), O. hellenica'' Boev, Lazaridis & Tsoukala 2014
****†''Otis (bird), O. khosatzkii'' Bocheński & Kuročkin 1987 (Late Pliocene of Varshets, Bulgaria)
Phoenicopteriformes
*Placement unresolved
**†''Phoeniconotius, Phoeniconotius eyrensis'' Miller 1963 [''Phoenicopterus novaehollandiae'' Miller 1963] (Etadunna Late Oligocene/Early Miocene of Lake Ngapakaldi to Lake Palankarinna Fossil Area, Lake Pitikanta, Australia)
*†Palaelodidae Stejneger 1885 – Swimming-flamingos
**†''Adelalopus, Adelalopus hoogbutseliensis'' Mayr & Smith 2002 (stout-legged flamingos) - (Borgloon Early Oligocene of Hoogbutsel, Belgium)
**†''Palaelodus'' Milne-Edwards 1863 [''Pliogrus'' Lambrecht 1933] (middle Oligocene –? Middle Pleistocene)
***†''Palaelodus ambiguus, P. ambiguus'' Milne-Edwards 1863 [''Grus problematica'' Milne-Edwards 1871; ''Probalearica problematica'' (Milne-Edwards 1871); ''Palaelodus gracilipes'' Milne-Edwards 1863; ''Palaelodus crassipes'' Milne-Edwards 1863; ''Paloelodus minutus'' (sic) Milne-Edwards 1868, ''Grus miocenicus'' Grigorescu & Kessler 1977]
***†''Palaelodus aotearoa, P. aotearoa'' Worthy et al. 2010
***†''Palaelodus germanicus, P. germanicus'' (Lambrecht 1933) [''Pliogrus germanicus'' Lambrecht 1933]
***†''Palaelodus kurochkini, P. kurochkini'' Zelenkov 2013
***†''Palaelodus pledgei, P. pledgei'' Baird & Vickers-Rich 1998
***†''Palaelodus wilsoni, P. wilsoni'' Baird & Vickers-Rich 1998
**†''Megapaloelodus'' Miller 1944 (Late Oligocene – Early Pliocene)
***†''Megapaloelodus, M. conectens'' (Miller 1944)
***†''Megapaloelodus, M. peiranoi'' Agnolin 2009
***†''Megapaloelodus, M. goliath'' (Milne Edwards 1863) Cheneval 1983c [''Paloelodus goliath'' (sic) Milne-Edwards 1868]
***†''Megapaloelodus, M. opsigonus'' Brodkorb 1961
*Phoenicopteridae – flamingos
**†''Harrisonavis, Harrisonavis croizeti'' (Gervais 1852) Kashin 1978 [''Gervaisia'' Harrison & Walker 1976 non Bonaparte 1854 non Waga 1858 non Robineau-Desvoidy 1863; ''Phoenicopterus croizeti'' Gervais 1852; ''Gervaisia croizeti'' (Gervais 1852) Harrison & Walker 1976]
**†''Leakeyornis, Leakeyornis aethiopicus'' (Harrison & Walker 1976) Vickers-Rich & Walker 1983 [''Phoenicopterus aethiopicus'' Harrison & Walker 1976]
**†''Elornis'' Milne-Edwards 1868 [''Elornis'' Aymard 1856 nomen nudum; ''Helornis'' Lydekker 1891; ''Actiornis''] (Middle? Eocene – Early Oligocene)
***†''Elornis anglicus, E. anglicus'' Aymard 1856 nomen nudum [''Helornis anglicus''; ''Actiornis anglicus'' Lydekker 1891]
***†''Elornis grandis, E. grandis'' Milne-Edwards 1868 [''Elornis grandis ''1856 Aymard nomen nudum]
***†''Elornis littoralis, E. littoralis'' Milne-Edwards 1868 [''Elornis littoralis'' Aymard 1856 nomen nudum; ''Elornis antiquus'' Aymard 1856 nomen nudum; ''Elornis antiquus'' Milne-Edwards 1868; ''Helornis littoralis'' (Milne-Edwards 1868) Lydekker 1891]
**Placement unresolved
***Phoenicopteridae gen. et sp. indet. (Middle? – Late Miocene)
**''Extant genera present in the fossil record''
***''Phoenicopterus'' (middle Oligocene – Recent)
****†''Phoenicopterus, P. floridanus'' Brodkorb 1953
****†''Phoenicopterus, P. stocki'' (Miller 1944)
****†''Phoenicopterus, P. siamensis'' Cheneval et al. 1991
****†''Phoenicopterus, P. gracilis'' Miller 1963 (Early Pleistocene of Lake Kanunka, Australia)
Podicipediformes
*Podicipedidae – grebes
**†''Miobaptus'' Švec 1982 (Early Miocene)
***†''Miobaptus huzhiricus, M. huzhiricus'' Zelenkov 2015
***†''Miobaptus walteri, M. walteri'' Švec 1982 [''Podiceps walteri'' (Švec 1984) Mlíkovský 2000]
**†''Miodytes, Miodytes serbicus'' Dimitreijevich, Gál & Kessler 2002
**†''Pliolymbus, Pliolymbus baryosteus'' Murray 1967 (Late Pliocene – Early? Pleistocene)
**†''Thiornis, Thiornis sociata'' Navás 1922 [''Podiceps sociatus'' (Navás 1922) Olson] (Late Miocene –? Early Pliocene)
**''Placement unresolved''
***Podicipedidae gen. et sp. indet. (Late Pliocene) – formerly included in ''Podiceps parvus''
***Podicipedidae gen. et sp. indet. UMMP 49592, 52261, 51848, 52276, KUVP 4484 (Late Pliocene)
***Podicipedidae gen. et sp. indet. (Late Pliocene/Early Pleistocene)
**''Extant genera present in the fossil record''
***''Aechmophorus'' (Late Pliocene – Recent)
****†''Aechmophorus, A. elasson'' Murray 1967
***''Podilymbus'' (Late Pliocene – Recent)
****†''Podilymbus, P. mujusculus'' Murray 1967
****†''Podilymbus, P. wetmorei'' Storer 1976
****†''Podilymbus, P. podiceps magnus''
***''Podiceps'' (Late Oligocene/Early Miocene – Recent)
****†''Podiceps, P. arndti'' Chandler 1990
****†''Podiceps, P. caspicus'' (Habizl 1783) [''Colymbus caspicus'' Habizl 1783]
****†''Podiceps, P. csarnotatus'' Kessler 2009
****†''Podiceps, P. discors'' Murray 1967
****†''Podiceps, P. dixi'' Brodkorp 1963
****†''Podiceps, P. miocenicus'' Kessler 1984
****†''Podiceps, P. oligocaenus'' (Shufeldt)
****†''Podiceps, P. parvus'' (Shufeldt 1913) [''Colymbus parvus'' Schufeldt 1913]
****†''Podiceps, P. solidus'' Kuročkin 1985
****†''Podiceps, P. subparvus'' (Miller & Bowman 1958) [''Colymbus subparvus'' Miller & Bowman 1958]
Phaethontiformes
* †Prophaethontidae
** †''Lithoptila, Lithoptila abdounensis'' (Late Paleocene – Early Eocene of Ouled Abdoun Basin, Morocco)– including ''Abdounornis''
** †''Prophaethon, Prophaethon shrubsolei'' (Late Paleocene ?– Early Eocene)
* Phaethontidae – tropicbirds
**''?†Proplegadis, Proplegadis fisheri'' (London Clay Early Eocene of England) – Threskiornithidae?
** †''Phaethusavis, Phaethusavis pelagicus'' (Early Eocene of Ouled Abdoun Basin, Morocco)
** †''Heliadornis'' (Miocene of North America and Europe)
*** †''Heliadornis ashbyi, H. ashbyi''
*** †''Heliadornis minor, H. minor''
*** †''Heliadornis paratethydicus, H. paratethydicus''
Ciconiiformes
The diverse group that includes storks, herons and New World vultures. Paraphyletic as listed here.
* †Ciconiidae – storks
** †''Ciconiopsis antarctica''
** †''Palaeoephippiorhynchus, Palaeoephippiorhynchus dietrichi'' (Early Oligocene)
** †''Grallavis, Grallavis edwardsi'' [''Propelargus edwardsi'' ; ''Palaeoephippiorhynchus edwardsi'' ] (Early Miocene) – may be same as ''Prociconia''
** †''Pelargosteon, Pelargosteon tothi'' (Early Pleistocene)
** †''Sanshuiornis, Sanshuiornis zhangi''
**''Placement unresolved''
***Ciconiidae gen. et sp. indet. – formerly ''Aquilavus bilinicus'' ; ''Cygnus bilinicus'' (Early Miocene)
***Ciconiidae gen. et sp. indet. (Late Miocene)
***cf. ''Leptoptilos'' gen. et sp. indet. – formerly ''Leptoptilos siwalicensis'' (Late Miocene? - Late Pliocene)
***Ciconiidae gen. et sp. indet. (Late Pleistocene) – ''Ciconia'' or ''Mycteria''?
**''Extant genera present in the fossil record''
***''Ciconia'' (Early Miocene? – Recent) – includes ''Xenorhynchus''
**** †''Ciconia lydekkeri, C. lydekkeri'' (Middle to Late Pleistocene)
**** †''Ciconia, C. gaudryi''
**** †''Ciconia, C. minor''
**** †''Ciconia, C. sarmatica''
**** †''Ciconia, C. kahli''
**** †''Ciconia louisebolesae, C. louisebolesae''
**** †''Ciconia, C. lucida''
**** †''Ciconia maltha, C. maltha'' (Asphalt/La Brea stork)
**** †''Ciconia nana, C. nana'' [''Xenorhynchus nanus'' ] (Australian stork)
**** †''Ciconia, C. stehlini''
***''Mycteria'' (Middle Miocene – Recent)
**** †''Mycteria, M. milleri'' [''Dissourodes milleri'' ]
**** †''Mycteria, M. wetmorei''
***''Ephippiorhynchus'' (Late Miocene – Recent)
**** †''Ephippiorhynchus, E. tchoufour''
**** †''Ephippiorhynchus, E. pakistanensis''
***''Leptoptilos'' (Late Miocene – Recent) – includes ''Cryptociconia''
**** †''Leptoptilos, L. falconeri'' [''Argala falconeri'' ]
**** †''Leptoptilos, L. pliocenicus''
**** †''Leptoptilos, L. inidcus'' [''Cryptociconia indica'' ]
**** †''Leptoptilos lüi, L. lüi''
**** †''Leptoptilos patagonicus, L. patagonicus''
**** †''Leptoptilos robustus, L. robustus''
**** †''Leptoptilos, L. richae''
**** †''Leptoptilos, L. titan''
***''Jabiru'' (Early Pliocene – Recent)
**** †''Jabiru, J. codorensis''
Pelecaniformes
The group that includes modern pelicans and cormorants. As presented here paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
; the tropicbird lineage is not part of this group and relationships with Procellariiformes and Sphenisciformes require more research. Also, as the pelicans are at least as close to the Ciconiiformes as to cormorants, the latter group is being recognized as Phalacrocoraciiformes by some recent authors and the core Pelecaniformes are occasionally merged into the Ciconiiformes.
* Basal and unresolved forms
** †''Piscator, Piscator tenuirostris'' (Early-Late Eocene of England) (Auk98:199)– basal phalacrococacoid?
** †''"Prophalacrocorax, Prophalacrocorax ronzoni'' [''Sula ronzoni'' ; ''Mergus ronzoni'' ] (Early Oligocene of Ronzon, France)
** † "Pelecaniformes" gen. et sp. indet. (Jebel Qatrani Early Oligocene of Fayum, Egypt) basal phalacrococacoid (similar to ''Piscator''?)?
* Fregatidae – frigatebirds
** †''Limnofregata'' (Early Eocene)
*** †''Limnofregata azygosternon, L. azygosternon''
*** †''Limnofregata, L. hutchisoni''
*** †''Limnofregata hasegawai, L. hasegawai''
* Sulidae – gannets and boobies
** †''Bimbisula, Bimbisula melanodactylus''
** †''Eostega, Eostega lebedinskyi'' (Middle/Late Eocene)
** †''Empheresula, Empheresula arvenensis'' [''Sula arvernensis'' ; ''Parasula''] (Late Oligocene – Middle Miocene)
** †''Masillastega, Masillastega rectirostris'' (middle Eocene)
** †''Microsula, Microsula pygmaea'' [''Sula pygmaea''; ''Microsula pygmaea''; ''Sula avita''; ''Enkurosula''; ''Pseudosula''] (Late Oligocene – Middle Miocene)
** †''Miosula, Miosula media'' (Late Miocene)
** †''Palaeosula, Palaeosula stocktoni'' (?Early Pliocene)
** †''Rhamphastosula'' (Early Pliocene)
*** †''Rhamphastosula ramirezi, R. ramirezi''
*** †''Rhamphastosula aguirrei, R. aguirrei''
** †''Sarmatosula, Sarmatosula dobrogensis'' (Middle Miocene)
** ''Placement unresolved''
*** Sulidae gen. et sp. indet. (Late Oligocene)
*** Sulidae gen. et sp. indet. (Late Pliocene)
** ''Extant genera present in the fossil record''
*** ''Morus (bird), Morus'' (Early Miocene – Recent)
**** †''Morus (bird), M. avitus'' [''Sula (Microsula) avita'' ; ''Microsula avitus'' ]
**** †''Morus (birds), M. lompocanus'' [''Sula lompocana'' ]
**** †''Morus (birds), M. loxostylus'' [''Sula loxostyla'' ; ''Morus atlanticus'' ; ''Sula atlantica'' ]
**** †''Morus (bird), M. olsoni''
**** †''Morus (bird), M. peninsularis'' [''Sula peninsularis'' ]
**** †''Morus (bird), M. peruvianus'' [''Sula peruviana'' ]
**** †''Morus (bird), M. recentior''
**** †''Morus (bird), M. reyanus'' (Del Rey gannet)
**** †''Morus (bird), M. vagabundus'' [''Sula vagabundus'' ]
**** †''Morus (bird), M. willetti'' [''Sula willetti'' ]
*** ''Sula (genus), Sula'' (Middle Pliocene – Recent)
**** †''Sula (genus), S. brandi''
**** †''Sula (genus), S. figueroae''
**** †''Sula (genus), S. clarki''
**** †''Sula (genus), S. humeralis'' [''Morus humeralis'' ]
**** †''Sula (genus), S. magna'' [''Morus magnus'' ]
**** †''Sula (genus), S. pohli''
**** †''Sula (genus), S. sulita''
**** †''Sula (genus), S. universitatis''
* Phalacrocoracidae – cormorants and shags
** †''Borvocarbo'' (Late Oligocene of C Europe) – phalacrocoracid? basal phalacrococacoid?
*** †''Borvocarbo guilloti, B. guilloti''
*** †''Borvocarbo tardatus, B. tardatus''
** †''Limicorallus'' (middle Oligocene)
*** †''Limicorallus saiensis, L. saiensis''
*** †''Limicorallus carbunculus, L. carbunculus''
** †''Nambashag''
*** †''Nambashag billerooensis, N. billerooensis''
*** †''Nambashag microglaucus, N. microglaucus''
** †''Nectornis'' (Late Oligocene?/Early Miocene – Middle Miocene)
*** †''Nambashag africanus, N. africanus''
*** †''Nambashag miocaenus, N. miocaenus'' [?''Phalacrocorax miocaenus'' ]
*** †''Nambashag anatolicus, N. anatolicus'' [''Phalacrocorax anatolicus'' ]
** †''Valenticarbo, Valenticarbo praetermissus'' (Late Pliocene/Early Pleistocene) – a '' nomen dubium''
** †''Oligocorax'' Lambrecht 1933
*** †''Oligocorax littoralis, O. littoralis'' [''Phalacrocorax littoralis'' ]
*** †''Oligocorax stoeffelensis, O. stoeffelensis'' [''Oligocorax sp.'' ; ''Borvocarbo stoeffelensis'' ]
** ''Placement unresolved''
*** †Phalacrocoracidae gen. et sp. indet. (Late Eocene ?–? mid-Oligocene)
** ''Extant genera present in the fossil record''
*** ''Phalacrocorax'' (Oligocene –? Recent) – may be several genera. Includes ''Australocorax'', ''Miocorax''
**** †''Phalacrocorax, P. gregorii''
**** †''Phalacrocorax, P. gregorii'' [''Australocorax vetustus'' ]
**** †''Phalacrocorax, P. chapalensis''
**** †''Phalacrocorax, P. destefani'' [''Paracorax destefani'' ]
**** †''Phalacrocorax, P. femoralis'' [''Miocorax femoralis'' ]
**** †''Phalacrocorax, P. filyawi''
**** †''Phalacrocorax, P. goletensis''
**** †''Phalacrocorax, P. ibericus''
**** †''Phalacrocorax, P. idahensis'' [''Graculus idahensis'' ]
**** †''Phalacrocorax, P. intermedius'' [''Phalacrocorax praecarbo'', ''Ardea brunhuberi''; ''Phalacrocorax brunhuberi'' ; ''Botaurites avitus'']
**** †''Phalacrocorax, P. kennelli''
**** †''Phalacrocorax, P. kuehneanus''
**** †''Phalacrocorax, P. longipes'' [''Pliocarbo longipes'' ]
**** †''Phalacrocorax, P. leptopus''
**** †''Phalacrocorax, P. macer''
**** †''Phalacrocorax, P. macropus'' [''Graculus macropus'' ]
**** †''Phalacrocorax, P. marinavis'' [''Oligocorax marinavis'' ]
**** †''Phalacrocorax, P. mongoliensis''
**** †''Phalacrocorax, P. owrei''
**** †''Phalacrocorax, P. reliquus''
**** †''Phalacrocorax, P. rogersi''
**** †''Phalacrocorax, P. serdicensis''
**** †''Phalacrocorax, P. tanzaniae''
**** †''Phalacrocorax, P. wetmorei''
* †Plotopteridae – diving-"boobies"
** †''Copepteryx''
*** †''Copepteryx, C. hexeris''
*** †''Copepteryx, C. titan''
** †''Hokkaidornis, Hokkaidornis abashiriensis''
** †''Phocavis, Phocavis maritimus''
** †''Plotopterum, Plotopterum joaquinensis''
** †''Tonsala''
*** †''Tonsala hildegardae, T. hildegardae''
*** †''Tonsala buchanani, T. buchanani''
* †Protoplotidae
** †''Protoplotus, Protoplotus beauforti'' (Paleocene? – middle Eocene of Sumatra)
* Anhingidae – darters
** †''Meganhinga, Meganhinga chilensis'' (Early Miocene)
** †''Macranhinga'' (giant snake-birds) (Middle/Late Miocene –? Early Pliocene)
*** †''Macranhinga paranensis, M. paranensis'' [''Meganhinga paranensis'' ]
*** †''Macranhinga ranzii, M. ranzii''
** †"Paranavis" (Middle/Late Miocene) – a ''nomen nudum''
** †''Giganhinga, Giganhinga kiyuensis'' (Late Pliocene/Early Pleistocene)
** ''Extant genera present in the fossil record''
*** ''Darter, Anhinga'' (Early Miocene – Recent)
**** †''Anhinga beckeri, A. beckeri''
**** †''Anhinga hadarensis, A. hadarensis''
**** †''Anhinga laticeps, A. laticeps'' [''Plotus laticeps'' ]
**** †''Anhinga malagurala, A. malagurala''
**** †''Anhinga pannonica, A. pannonica''
**** †''Anhinga subvolans, A. subvolans'' [''Phalacrocorax subvolans'' ]
**** †''Anhinga walterbolesi, A. walterbolesi''
**** †''Anhinga fraileyi, A. fraileyi'' [?''Macranhinga fraileyi'' ]
**** †''Anhinga grandis, A. grandis''
**** †''Anhinga minuta, A. minuta''
* Pelecanidae – pelicans
** †''Protopelicanus, Protopelicanus cuvieri''
** †''Miopelecanus'' Cheneval 1984
*** †''Miopelecanus, M. gracilis'' [''Pelecanus gracilis'' ]
*** †''Miopelecanus, M. intermedius'' [''Pelecanus intermedius'' ]
** ''Extant genera present in the fossil record''
*** ''Pelecanus'' (Late Pliocene – Recent)
**** †''Pelecanus aethiopicus, P. aethiopicus''
**** †''Pelecanus cautleyi, P. cautleyi''
**** †''Pelecanus fraasi, P. fraasi''
**** †''Pelecanus halieus, P. halieus''
**** †''Pelecanus odessanus, P. odessanus''
**** †''Pelecanus schreiberi, P. schreiberi''
**** †''Pelecanus sivalensis, P. sivalensis''
**** †''Pelecanus tirarensis, P. tirarensis''
* †Pelagornithidae Fürbringer 1888 – pseudotooth birds
** †''Pseudodontornis'' Lambrecht 1930 [''Neodontornis'' Harrison & Walker 1976; ''Macrodontopteryx'' Harrison & Walker 1976; ''Palaeochenoides'' Schufeldt 1916] (Late Paleocene – London Clay Early Eocene of England)
*** †''Pseudodontornis longidentata, P. longidentata'' Harrison & Walker 1976 [''Odontopteryx oweni'' (Harrison & Walker 1976); ''Macrodontopteryx oweni'' Harrison & Walker 1976]
*** †''Pseudodontornis tenuirostris, P. tenuirostris'' Harrison 1985
*** †''Pseudodontornis tschulensis, P. tschulensis'' (Averianov et al. 1991) [''Odontopteryx tschulensis'']
*** †''Pseudodontornis longirostris, P. longirostris'' (Spulski 1910) Lambrecht 1930 [''Odontopteryx longirostris'' Spulski 1910; ''Palaeochenoides miocaenus'' Schufeldt 1916; ''Pelagornis longirostris'' (Spulski 1910)]
*** †''Pseudodontornis stirtoni, P. stirtoni'' Howard & Warter 1969 [''Neodontornis stirtoni'' (Howard & Warter 1969) Harrison & Walker 1976; ''Pelagornis stirtoni'' (Howard & Warter 1969)]
** †''Dasornis'' Owen 1870 [''Megalornis'' Seeley 1866 non Gray 1841; ''Argillornis'' Owen 1878] (London Clay Early Eocene of England)
*** †''Dasornis, D. abdoun'' Bourdon 2010
*** †''Dasornis emuinus, D. emuinus'' (Bowerbank 1854) Owen 1870 [''Lithornis emuinus'' Bowerbank 1854; ''Megalornis emuianus'' (sic) (Bowerbank 1854) Seeley 1866; ''Neptuniavis miranda'' Harrison & Walker 1977; ''Argillornis longipennis'' Owen 1878; ''Dasornis londinensis'' Owen 1869 nomen nudum; ''Dasornis londinensis'' Owen 1870; ''Argillornis emuinus'' (Bowerbank 1854) Brodkorb 1963; ''Argillornis longipes'' Lambrecht 1933 (lapsus)]
*** †''Dasornis, D. gigas'' [''Odontopteryx gigas'' Bourdon 2006 nomen nudum]
** †''Macrodontopteryx'' (London Clay Early Eocene of England)
** †''Odontopteryx, Odontopteryx toliapica'' Owen 1873 [''Odontornis'' (sic) Owen 1873; ''Neptuniavis minor'' Harrison & Walker 1977; ''Dasornis toliapica'' (Owen 1873)] (London Clay Early Eocene of England)
** †''Gigantornis, Gigantornis eaglesomei'' Andrews 1916 (middle Eocene of Nigeria)
** †''Cyphornis, Cyphornis magnus'' Cope 1894 (Eocene of Vancouver, Canada)
** †''Osteodontornis, Osteodontornis orri'' Howard 1957 [''Pelagornis orri'' Howard 1957] (Early Oligocene – Pliocene)
** †''Pelagornis'' (Middle Miocene of France – Late Pliocene of Morocco)
*** †''Pelagornis miocaenus, P. miocaenus'' Lartet 1857 (Miocene false-tooothed pelican)
*** †''Pelagornis mauretanicus, P. mauretanicus'' Mourer-Chauviré & Geraads 2008
*** †''Pelagornis chilensis, P. chilensis'' Mayr & Rubilar-Rogers 2010
*** †''Pelagornis sandersi, P. sandersi'' Ksepka 2014
** †''Caspiodontornis, Caspiodontornis kobystanicus'' Aslanova & Burčak-Abramovič 1982 [?''Guguschia nailiae'' Aslanova & Burčak-Abramovič 1968]
** †''Tympanoneisiotes, Tympanoneisiotes wetmorei'' Hopson 1964 [''Pelagornis wetmorei'' (Hopson 1964)]
** †''Lutetodontopteryx, Lutetodontopteryx tethyensis'' Mayr & Zvonok 2012
** †''Aequornis, Aequornis traversei'' Bourdon 2006 nomen nudum
** ''Placement unresolved''
*** ?Pelagornithidae gen. et sp. indet. (middle Eocene)
* †Xenerodiopidae (Early Oligocene)
** †''Xenerodiopus, Xenerodiopus mycter''
*Ardeidae – herons
** †''Ardeagrandis, Ardeagrandis arborea''
** †''Calcardea, Calcardea junnei'' (Paleocene)
** †''Matuku (bird), Matuku otagoense''
** †''Nyctisoma, Nyctisoma robusta''
** †''Palaeophoyx, Palaeophoyx columbiana''
** †''Pikaihao, Pikaihao bartlei''
** †''Proardea, Proardea amissa'' [''Ardea amissa'' ; ''Egretta amissa'' ] (Late Eocene –? Late Oligocene)
** †''Proardeola, Proardeola walkeri'' [''Ardea Formosa'' ; ''Ardeola walker'' ] – possibly same as ''Proardea''
** †''Zeltornis, Zeltornis ginsburgi'' (Early Miocene)
**Placement unresolved
*** †''Anas basaltica'' Bayer 1882 (Late Oligocene)
**''Extant genera present in the fossil record''
*** ''Nyctanassa''
**** †''Nyctanassa, N. kobdoena''
*** ''Ardea (genus), Ardea'' (Middle Miocene – Recent)
**** †''Ardea (genus), A. aurelianensis'' [''Proardea aurelianensis'' ]
**** †''Ardea (genus), A. polkensis''
**** †''Ardea (genus), A. howardae''
**** †''Ardea (genus), A. paloccidentalis''
**** †''Ardea (genus), A. sytchevskayae''
*** ''Egretta'' (Late Miocene/Early Pliocene – Recent)
**** †''Egretta, E. polgardiensis''
**** †''Egretta, E. subfluvia''
*** ''Butorides'' (Early Pleistocene – Recent)
**** †''Butorides, B. validipes''
*** ''Botaurus''
**** †''Botaurus, B. hibbardi''
*** ''Syrigma''
**** †''Syrigma, S. sanctimartini''
*Scopidae – hammerkop
**''Prehistoric species of extant genera''
*** †''Scopus xenopus''
*Threskiornithidae – ibises
**Threskiornithidae gen. et sp. indet. NMMP-KU 1301 (Pondaung middle Eocene of Paukkaung, Myanmar) – Threskiornithinae?
** †''Gerandibis, Gerandibis paganus'' [''Milnea Lydekker'' ; ''Ibis pagana'' ; ''Eudocimus paganus'' ; ''Plegadis paganus'' ; ''Actiornis pagana'' ; ''Milnea gracilis'' ; ''Plegadis gracilis'' ]
** †''Rhynchaeites, Rhynchaeites messelensis'' [''Plumumida'' ; ''Plumumida lutetialis'' ] (middle Eocene of Messel, Germany) – may include ''Mopsitta''
** †''Vadaravis brownae''
**''Extant genera present in the fossil record''
***''Geronticus'' (Middle Miocene – Recent)
**** †''Geronticus, G. perplexus'' [''Ardea perplexa'' ; ''Proardea perplexus'' ]
**** †''Geronticus, G. apelex''
**** †''Geronticus, G. olsoni''
***''Platalea''
**** †''Platalea, P. chione'' [''Ajaia chione'' ]
**** †''Platalea, P. tiangangensis'' (Xiacaowan Middle Miocene of Sihong, China)
***''Plegadis''
**** †''Plegadis, P. pharangites'' [''Plegadis gracilis'' ] (Late Pliocene of WC US)
***''Theristicus''
**** †''Theristicus, T. wetmorei'' (Late Pleistocene of Peru)
***''Eudocimus''
**** †''Eudocimus, E. leiseyi'' (Early Pleistocene of Florida)
**** †''Eudocimus, E. peruvianus'' (Late Pleistocene of Peru; may be living species)
**** †''Eudocimus'' sp. (Middle Pliocene of Florida)
*Balaenicipitidae – shoebills
**''Goliathia, Goliathia andrewsi'' (Late Eocene/Early Oligocene of Egypt)
**''Paludavis, Paludiavis richae'' (Late Miocene of Tunisia and Pakistan)
Procellariiformes
The group that includes modern albatrosses, petrels and storm-petrels.
*Placement unresolved
**†''Eopuffinus, Eopuffinus kazachstanensis'' (Late Paleocene of Zhylga, Kazakhstan)
**†''Makahala, Makahala mirae''
**†''Proaestrelata''
*†Marinavidae
**†''Marinavis, Marinavis longirostris''
*† Tytthostonychidae
**†'' Tytthostonyx glauconiticus''
*†Diomedeoididae
**†''Rupelornis'' van Beneden 1871 [''Gaviota (genus), Gaviota'' ; ''Ardeita'' ; ''Diomedeoides'' ; ''Frigidafrons'' ] (Early Oligocene– Early Miocene of C Europe and Iran)
***†''Rupelornis harmati, R. harmati'' [''Diomedeoides harmati'' ]
***†''Rupelornis definitus, R. definitus'' [''Vanellus selysii'' ; ''Anas creccoides'' ; ''Gaviota lipsiensis'' ; ''Diomedeoides minimus'' ; ''Frigidafrons brodkorbi'' ; ''Ardeita gracilis'' ; ''Diomedeoides lipsiensis'' ; ''Diomedeoides brodkorbi'' ]
***†''Rupelornis babaheydariensis, R. babaheydariensis'' [''Frigidafons babaheydariensis'' ; ''Diomedeoides babaheydariensis'' ]
*Oceanitidae
**''Extant genera present in the fossil record''
***''Oceanites''
****†''Oceanites, O. zaloscarthmus''
*Diomedeidae – albatrosses
**†''Murunkus, Murunkus subitus'' (middle Eocene)
**†''Plotornis'' [''Chenornis'' ] (Early – Middle Miocene)
***†''Plotornis arvernensis, P. arvernensis'' [''Puffinus arvernensis'' ; ''Puffinus arvernensis'' ]
***†''Plotornis deltortrii, P. deltortrii''
***†''Plotornis graculoides, P. graculoides'' [''Chenornis graculoides'' ]
**†''Tydea, Tydea septentrionalis'' (Early Oligocene of the North Sea basin)
**†Diomedeidae gen. et sp. indet. (Late Oligocene of South Carolina)
**''Extant genera present in the fossil record''
***''Diomedea'' (Middle Miocene – Recent)
****†''Diomedea, D. anglica''
****†''Diomedea, D. howardae''
****†''Diomedea, D. milleri''
****†''Diomedea, D. rumana''
****†''Diomedea, D. tanakai''
****†''Diomedea, D. thyridata''
***''Phoebastria'' (Middle Miocene – Recent)
****†''Phoebastria, P. rexsularum''
***''Thalassarche'' (Late Miocene – Recent)
****†''Thalassarche, T. thyridata''
*Hydrobatidae – storm-petrels
**†''Primodroma, Primodroma bournei'' (Early Eocene)
**''Prehistoric species of extant genera''
***†''Oceanodroma hubbsi, "Oceanodroma" hubbsi'' (Hubbs' Storm-petrel)(Capistrano Middle/Late Miocene of Orange County, US)
***†''Pelagodroma'' sp. 1
***''Pelagodroma'' sp. 2
*Procellariidae – petrels
**†''Argyrodyptes, Argyrodyptes microtarsus'' (San Julián Late Eocene/Early Oligocene of Chubut, Argentina)
**†''Pterodromoides, Pterodromoides minoricensis''
**''Extant genera present in the fossil record''
***''Calonectris'' (Early Pliocene – Recent)
****†''Calonectris, C. krantzi''
****†''Calonectris, C. kurodai''
****†''Calonectris, C. wingatei''
***''Fulmarus'' (Middle Miocene – Recent)
****†''Fulmarus, F. hammeri''
****†''Fulmarus, F. miocaenus''
***''Pachyptila'' (Late Miocene– Recent)
****†''Pachyptila, P. salax''
***''Pelecanoides'' (Early/Middle Miocene – Recent)
****†''Pelecanoides, P. cymatotrypetes''
****†''Pelecanoides, P. miokuaka'' (Miocene diving petrel)
****†''Pelecanoides, P. urinatrix''
***''"Pterodroma"'' (Pleistocene – Recent)
****†''Pterodroma, P. kurodai''
****†''Pterodroma rupinarum, P. rupinarum''
***''Puffinus'' (Early Oligocene – Recent)
****†''Puffinus aquitanicus, P. aquitanicus'' ''Procellaria aquitanica''
****†''Puffinus antiquus, P. antiquus'' ''Procellaria antiqua''
****†''Puffinus barnesi, P. barnesi''
****†''Puffinus calhouni, P. calhouni''
****†''Puffinus conradi, P. conradi''
****†''Puffinus diatomicus, P. diatomicus''
****†''Puffinus felthami, P. felthami''
****†''Puffinus gilmorei, P. gilmorei''
****†''Puffinus inceptor, P. inceptor''
****†''Puffinus kanakoffi, P. kanakoffi''
****†''Puffinus micraulax, P. micraulax''
****†''Puffinus mitchelli, P. mitchelli''
****†''Puffinus nestori, P. nestori''
****†''Puffinus priscus, P. priscus''
****†''Puffinus raemdonckii, P. raemdonckii'' [Larus Raemdonckii Beneden 1871]
****†''Puffinus tedfordi, P. tedfordi''
Gaviiformes
*†'' Gaviella pusilla'' (Shufeldt 1915) Wetmore 1940
*†''Nasidytes, Nasidytes ypresianus'' (Mayr & Kitchener, 2022)
*Gaviidae – loons
**†''Colymboides'' Milne-Edwards 1867 [Dyspetornis Oberholser 1905; Hydrornis Milne-Edwards 1867 non Blyth 1843; Davisona Mathews 1935; Megagallinula Kuročkin 1968] (Late Eocene – Early Miocene) – paraphyletic?
***†''Colymboides, C. harundinea'' (Kuročkin 1968) ''Megagallinula harundinea'' Kuročkin 1968)
***†''Colymboides, C. metzleri'' Mayr 2004
***†''Colymboides, C. anglicus'' Lydekker 1891a
***†''Colymboides, C. belgicus'' Mayr & Smith 2002
***†''Colymboides, C. minutus'' Milne-Edwards 1867
** Placement unresolved
***''"Gavia" portisi'' (Late Pliocene of Orciano Pisano, Italy) – rentatively placed here. A '' nomen dubium''?
**''Extant genera present in the fossil record''
***''Gavia (bird), Gavia'' (Early Miocene – Recent)
****†''Gavia (bird), G. egeriana'' Švec 1982
****†''Gavia (bird), G. brodkorbi'' Howard 1978
****†''Gavia (bird), G. paradoxa'' Umanskaja 1981
****†''Gavia (bird), G. schultzi'' Mlíkovsky 1998
****†''Gavia (bird), G. howardae'' Brodkorb 1953c
****†''Gavia (bird), G. moldavica'' Kessler 1984
****†''Gavia (bird), G. concinna'' Wetmore 1940 [''Gavia palaeodytes'' Wetmore 1943]
****†''Gavia (bird), G. fortis'' Olson & Rasmussen 2001
Sphenisciformes
*Unresolved and basal forms
**†''Waimanu'' Jones, Ando & Fordyce 2006 (Early – Late Palaeocene)
***†''Waimanu manneringi, W. manneringi'' Jones, Ando & Fordyce 2006 (Mannering's penguin)
***†''Waimanu, W. tuatahi'' Jones, Ando & Fordyce 2006 (Waipara penguin)
**†''Perudyptes, Perudyptes devriesi'' Clarke et al. 2007 – basal? (middle Eocene)
**Sphenisciformes gen. et sp. indet. CADIC P 21 (middle Eocene)
*Spheniscidae – penguins
**†''Kaiika, Kaiika maxwelli'' Fordyce & Tomas 2011
**†''Palaeoapterodytes, Palaeoapterodytes ictus'' (Ameghino 1891) Ameghino 1905 (nomen dubium) [''Apterodytes'' Ameghino 1891 non Hermann 1783; ''Apterodytes ictus'' Ameghino 1891]
**†''Tereingaornis, Tereingaornis moisleyi'' Scarlett 1983 (Moisley's penguin)
**†''Crossvallia, Crossvallia unienwillia'' Tambussi et al. 2005 (Late Paleocene)
**†''Anthropornis'' Wiman 1905 (mid-Eocene –? Early Oligocene)
***†''Anthropornis, A. nordenskjoeldii'' Wiman 1905 (Nordenskjoeld's giant penguin)
***†''Anthropornis, A. grandis'' (Wiman 1905) [''Arthrodytes grandis'' (Ameghino 1901) Ameghino 1905]
**†''Archaeospheniscus'' Marples 1952 (mid-/Late Eocene – Late Oligocene)
***†''Archaeospheniscus lowei, A. lowei'' Marples 1952 (Lowe's penguin)
***†''Archaeospheniscus lopdelli, A. lopdelli'' Marples 1952 (Lopdell's penguin)
**†''Delphinornis'' Wiman 1905 [''Notodyptes'' Marples 1953] (mid-/Late Eocene –? Early Oligocene)
***†''Delphinornis gracilis, D. gracilis'' Myrcha et al. 2002
***†''Delphinornis larseni, D. larseni'' Wiman 1905
***†''Delphinornis arctowskii, D. arctowskii'' Myrcha et al. 2002
***†''Delphinornis wimani, D. wimani'' (Marples 1953) Ksepka & Clarke 2010 [''Notodyptes wimani'' Marples 1953; ''Archaeospheniscus wimani'' (Marples 1953) Simpson 1971a]
**†''Palaeeudyptes'' Huxley 1859 [''Eosphaeniscus'' Wiman 1905] (mid-/Late Eocene – Late Oligocene)
***†''Palaeeudyptes klekowskii, P. klekowskii'' Myrcha, Tatur & del Valle 1990
***†''Palaeeudyptes gunnari, P. gunnari'' (Wiman 1905) [''Eosphaeniscus gunnari'' Wiman 1905]
**†''Icadyptes, Icadyptes salasi'' Clarke et al. 2007 (Late Eocene)
**†''Inkayacu, Inkayacu paracasensis'' Clarke et al. 2010 (Late Eocene)
**†''Pachydyptes, Pachydyptes ponderosus'' Oliver 1930 (Late Eocene)
**†''Marambiornis, Marambiornis exilis'' Myrcha et al. 2002 (Late Eocene–? Early Oligocene)
**†''Mesetaornis, Mesetaornis polaris'' Myrcha et al. 2002 (Late Eocene –? Early Oligocene)
**†''Tonniornis'' Tambussi et al. 2006 (Late Eocene –? Early Oligocene)
***†''Tonniornis mesetaensis, T. mesetaensis'' Tambussi et al. 2006
***†''Tonniornis minimum, T. minimum'' Tambussi et al. 2006
**†''Wimanornis, Wimanornis seymourensis'' Simpson 1971 (Late Eocene –? Early Oligocene)
**†''Arthrodytes, Arthrodytes andrewsi'' Ameghino 1901 (Late Eocene/Early Oligocene – Early Miocene)
**†''Duntroonornis, Duntroonornis parvus'' Marples 1953 (Duntroon penguin) (Late Oligocene)
**†''Kairuku'' Ksepka et al. 2012 (Late Oligocene)
***†''Kairuku grebneffi, K. grebneffi'' Ksepka et al. 2012
***†''Kairuku waitaki, K. waitaki'' Ksepka et al. 2012
**†''Korora (bird), Korora oliveri'' Marples 1952 (Oliver's penguin) (Late Oligocene)
**†''Platydyptes'' Marples 1952 (Late Oligocene)
***†''Platydyptes marplesi, P. marplesi'' (Brodkorb 1963) Simpson 1971 [''Palaeeudyptes marplesi'' Brodkorb 1963] (Simpson's/Marples' penguin)
***†''Platydyptes novaezealandiae, P. novaezealandiae''(Oliver 1930) Marples 1952 (wide-flippered penguin)
***†''Platydyptes amiesi, P. amiesi'' Marples 1952 (Amies’ penguin)
**†''Eretiscus, Eretiscus tonnii'' (Simpson 1981) Olson 1986 [''Microdytes'' Simpson 1981 non Balfour-Browne 1949; ''Microdytes tonnii'' Simpson 1981] (Patagonia Early Miocene)
**†''Palaeospheniscus'' Moreno & Mercerat 1891 [''Neculus'' Ameghino 1905; ''Paraspheniscus'' Ameghino 1905 nec Hendel 1927; ''Perispheniscus'' Ameghino 1905; ''Pseudospheniscus'' Ameghino 1905; ''Chubutodyptes'' Simpson 1970] (Early? – Late Miocene/Early Pliocene)
***†''Palaeospheniscus, P. huxleyorum'' Simpson 1973
***†''Palaeospheniscus wimani, P. wimani'' (Ameghino 1905) [''Perispheniscus wimani'' Ameghino 1905; ''Palaeospheniscus robustus'' (partim) Simpson 1946]
***†''Palaeospheniscus gracilis, P. gracilis'' Ameghino 1899 [?''Delphinornis gracilis''; ''Palaeospheniscus nereius'' Ameghino 1901; ''Neculus rothi'' Ameghino 1905; ''Palaeospheniscus rothi'' Ameghino 1905; ''Paraspheniscus nereius'' (Ameghino 1901) Ameghino 1905; ''Palaeospheniscus medianus'' Ameghino 1905]
***†''Palaeospheniscus patagonicus, P. patagonicus'' Moreno & Mercerat 1891 [''Palaeospheniscus menzbieri'' Moreno & Mercerat 1891; ''Palaeospheniscus interruptus'' Ameghino 1905; ''Palaeospheniscus intermedius'' Ameghino 1905; ''Palaeospheniscus affinis'' Ameghino 1905]
***†''Palaeospheniscus bergi, P. bergi'' Moreno & Mercerat 1891 [''Paraspheniscus bergi'' (Moreno & Mercerat 1891) Ameghino 1905; ''Palaeospheniscus planus'' Ameghino 1905; ''Palaeospheniscus interplanus'' Ameghino 1905; ''Pseudospheniscus interplanus'' Ameghino 1905; ''Pseudospheniscus planus'' Ameghino 1905 nom rej.; ''Pseudospheniscus concavus'' Ameghino 1905; ''Pseudospheniscus convexus'' Ameghino 1905 nom rej.]
***†''Palaeospheniscus, P. biloculata'' (Simpson 1970) Acosta Hospitaleche 2007 [''Chubutodyptes biloculata'' Simpson 1970]
**†''Paraptenodytes'' Ameghino 1891 [''Isotremornis'' Ameghino 1905; ''Metancylornis'' Ameghino 1905; ''Treleudytes'' Ameghino 1905] (Early – Late Miocene/Early Pliocene)
***†''Paraptenodytes brodkorbi, P. brodkorbi'' Simpson 1972 [''Isotremornis nordenskjöld''i Ameghino 1905 (partim)]
***†''Paraptenodytes robustus, P. robustus'' (Ameghino 1895) [''Palaeospheniscus robustus'' Ameghino 1895; ''Perispheniscus robustus'' (Ameghino 1895) Brodkorb 1963; ''Paraptenodytes curtus'' Ameghino 1901; ''Metancylornis curtus'' (Ameghino 1901) Ameghino 1905; ''Paraptenodytes grandis'' Ameghino 1901; ''Arthrodytes grandis'' (Ameghino 1901) Ameghino 1905; ''Paraptenodytes andrewsi'' Ameghino 1901; ''Arthrodytes andrewsi'' (Ameghino 1901) Ameghino 1905; ''Treleudytes crassus'' Ameghino 1905]
***†''Paraptenodytes antarcticus, P. antarcticus'' (Huxley 1859) Ameghino 1891 [''Palaeeudyptes antarcticus'' Huxley 1859 non Hector 1873; ''Palaeospheniscus antarcticus'' (Huxley 1859) Moreno & Mercerat 1891] (narrow-flippered penguin)
**†''Anthropodyptes, Anthropodyptes gilli'' Simpson 1959 (Middle Miocene)
**†''Madrynornis, Madrynornis mirandus'' Hospitaleche et al. 2007 (Late Miocene)
**†''Pseudaptenodytes'' Simpson 1970 (Late Miocene/Early Pliocene)
***†''Pseudaptenodytes, P. minor'' Simpson 1970
***†''Pseudaptenodytes, P. macraei'' Simpson 1970
**†''Dege (bird), Dege hendeyi'' Simpson 1979 (Early Pliocene)
**†''Marplesornis, Marplesornis novaezealandiae'' (Marples 1960) Simpson 1972 [''Paleospheniscus novaezealandiae'' Marples 1960] (Harris penguins) (Early Pliocene)
**†''Nucleornis, Nucleornis insolitus'' Simpson 1979 (Early Pliocene)
**†''Inguza, Inguza predemersus'' (Simpson 1971) Simpson 1975 [''Spheniscus predemersus'' Simpson 1971] (Late Pliocene)
**''Placement unresolved''
***Spheniscidae gen. et sp. indet (Late Oligocene/Early Miocene of Hakataramea, New Zealand)
**''Extant genera present in the fossil record''
***''Pygoscelis'' (Middle/Late Miocene – Recent)
****†''Pygoscelis, P. calderensis'' Walsh & Suárez 2006
****†''Pygoscelis tyreei, P. tyreei'' Simpson 1972 (Tyree's penguin)
****†''Pygoscelis, P. grandis'' Walsh & Suárez 2006
***''Spheniscus'' (Middle/Late Miocene – Recent)
****†''Spheniscus, S. muizoni'' Göhlich 2007
****†''Spheniscus, S. chilensis'' Emslie & Correa 2003
****†''Spheniscus, S. megaramphus'' Stucchi, Urbina & Giraldo 2003
****†''Spheniscus urbinai, S. urbinai'' Stucchi 2002
***''Aptenodytes'' (Early? Pliocene – Recent)
****†''Aptenodytes ridgeni, A. ridgeni'' Simpson 1972 [''Arthrodytes ridgeni'' (Simpson 1972)] (Ridgen's Penguin)
***''Eudyptes''
****†''Eudyptes, E. calauina'' Hoffmeister et al. 2014
Pterocliformes
*Pteroclidae – sandgrouse
**†''Gerandia, Gerandia calcaria'' (Milne-Edwards 1869) Lambrecht 1933 [''Columba calcaria'' Milne-Edwards 1869] (Early Miocene)
**†''Leptoganga, Leptoganga sepultus'' (Milne-Edwards 1869) Mourer-Chauviré 1993 [''Pterocles sepultus'' Milne-Edwards 1869]
**†''Archaeoganga'' Mourer-Chauviré 1992
***†''Archaeoganga validus, A. validus''(Milne-Edwards 1892) [''Pterocles validus'' Milne-Edwards 1892]
***†''Archaeoganga pinguis, A. pinguis'' Mourer-Chauviré 1992
***†''Archaeoganga larvatus, A. larvatus'' (Milne-Edwards 1892) [''Pterocles larvatus'' Milne-Edwards 1892]
**''Extant genera present in the fossil record''
***''Syrrhaptes''
****†''Syrrhaptes, S. kashini'' Kuročkin 1985
Columbiformes
*Columbidae – doves and pigeons
**''Arenicolumba'' (Early Miocene) – doubtfully distinct from ''Patagioenas''
**''Rupephaps'' (Early Miocene)
**Placement unresolved
***Columbidae gen. et sp. indet. (Early/Middle Miocene)
**''Extant genera present in the fossil record''
***''Columba (genus), Columba'' (Early Pliocene – Recent)
***''Patagioenas'' (Early Pliocene – Recent)
Psittaciformes
Unresolved and basal fossil parrots:
*''Psittacopes'' (Early/middle Eocene)
*''Serudaptus'' – pseudasturid or psittacid?
*Pseudasturidae (= Halcyornithidae
Halcyornithidae or Pseudasturidae is a family of fossil birds, possibly belonging to the order Psittaciformes (parrots and relatives). Members of this family lived in the Eocene, around 55-48 million years ago. Fossil remains are known from the ...
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*Messelasturidae
**''Tynskya, Tynskya eocaena'' (Early Eocene of N America and England)
**''Messelastur, Messelastur gratulator'' (middle Eocene of Messel, Germany)
**Pseudasturidae FU 125 gen. et sp. indet. (Early Eocene)
**''Pseudasturides'' – formerly ''Pseudastur''
*Vastanavidae
**''Vastanavis'' (Early Eocene of Vastan, India)
*Quercypsittidae
**''Quercypsitta'' (Late Eocene)
*Cacatuidae
**''Extant genera present in the fossil record''
***''Cacatua'' (Early Miocene – Recent)
*Psittacidae – parrots, parakeets and lories
**''Archaeopsittacus'' (Late Oligocene/Early Miocene)
**''Xenopsitta'' (Early Miocene)
**Psittacidae gen. et spp. indet. (Early/Middle Miocene) – several species
**''Bavaripsitta'' (Middle Miocene)
**Psittacidae gen. et sp. indet. (Middle Miocene) – erroneously placed in ''Pararallus dispar'', includes ''"Psittacus lartetianus''
**''Extant and recently extinct genera present in the fossil record''
***''Conuropsis'' (Early? Miocene – Holocene)
***''Nandayus'' (Late Pliocene – Recent)
***''Cyanoliseus'' (Middle Pleistocene – Recent)
***''Aratinga'' (Late Pleistocene – Recent)
***''Rhynchopsitta'' (Late Pleistocene – Recent)
*Strigopidae – New Zealand parrots, kakapo
**''Nelepsittacus'' (Middle Miocene)
**''Heracles'' (Middle Miocene)
Opisthocomiformes
Hoatzins
*Opisthocomidae [Foratidae ; Onychopterygidae ]
**†''Foro (bird), Foro panarium'' (Early Eocene) - cuculiform?
**†''Hoatzi, Hoatzi panarium''
**†''Onychopteryx, Onychopteryx simpsoni'' (Early Eocene of Argentina) – falconid? A '' nomen dubium''
**†''Protoazin, Protoazin parisiensis''
**†''Namibiavis, Namibiavis senutae'' (Late Oligocene of Namibia)
**†''Hoazinavis, Hoazinavis lacustris'' (Miocene of Brazil)
**†''Hoazinoides, Hoazinoides magdalenae'' (Late Miocene of Colombia)
Musophagiformes
*Musophagidae – turacos
**''Placement unresolved''
***Musophagidae gen. et sp. indet. (Late Oligocene – Middle Miocene of WC Europe)
***Musophagidae gen. et sp. indet. (Egypt)
**†''Veflintornis'' [''Apopempsis'' ] (Middle Miocene)
***†''Veflintornis meini, V. meini'' [''Musophaga meini'' ]
***†''Veflintornis africanus, V. africanus'' [''Musophaga africanus'' ; ''Apopempsis africanus'' ] (Early Miocene)
Cuculiformes
Cuckoos, turacos and allies.
*Placement unresolved
**Cuculiformes gen. et sp. indet. (Early Eocene)
*Cuculidae – cuckoos
**''Eocuculus'' (Late Eocene)
**''Dynamopterus'' (Late Eocene/Early Oligocene)
**''Neococcyx'' (Early Oligocene)
**''Cursoricoccyx'' (Early Miocene)
**''Placement unresolved''
***Cuculidae gen. et sp. indet. (Early Pliocene)
Accipitriformes
*†Teratornithidae – teratorns
**†''Argentavis, Argentavis magnificens'' (Late Miocene)
**†''Aiolornis, Aiolornis incredibilis'' [''Teratornis incredibilis'' ] (Early Pliocene – Late Pleistocene)
**†''Oscaravis, Oscaravis olsoni'' [''Teratornis olsoni'' ] (Cuban teratorn)(Pleistocene)
**†''Cathartornis, Cathartornis gracilis''
**†''Taubatornis, Taubatornis campbelli''
**†''Teratornis'' (Early Pleistocene – Late Pleistocene)
***†''Teratornis woodburnensis, T. woodburnensis''
***†''Teratornis merriami, T. merriami'' (Merriam's teratorn)
*Cathartidae – New World vultures
**†''Brasilogyps, Brasilogyps faustoi'' (Late Oligocene – Early Miocene)
**†''Kuntur, Kuntur cardenasi''
**†''Pleistovultur, Pleistovultur nevesi''
**†''Tapinopus, Tapinopus ellioti''
**†''Diatropornis, Diatropornis ellioti'' (European vulture) (Late Eocene/Early Oligocene –? middle Oligocene)
**†''Phasmagyps, Phasmagyps patritus'' (Early Oligocene)
**†''Hadrogyps, Hadrogyps aigialeus'' (American dwarf vulture) (Middle Miocene)
**†''Pliogyps'' (Miocene vulture) (Late Miocene – Late Pliocene)
***†''Pliogyps, P. charon''
***†''Pliogyps, P. fisheri''
**†''Perugyps, Perugyps diazi'' (Peruvian vulture) (Late Miocene/Early Pliocene)
**†''Dryornis'' (Argentinean vulture) (Early Miocene – Late Pliocene) – similar to the extant genus ''Vultur''
***†''Dryornis, D. pampeanus''
***†''Dryornis, D. hatcheri''
**†''Aizenogyps, Aizenogyps toomeyae'' (South American vulture) (Late Pliocene)
**†''Breagyps, Breagyps clarki'' (long-legged vulture) (Late Pleistocene)
**†''Geronogyps, Geronogyps reliquus'' (Late Pleistocene)
**†''Wingegyps, Wingegyps cartellei'' (Amazonian vulture) (Late Pleistocene)
**†''Parasarcoramphus, Parasarcoramphus milneedwardsi''
**''Placement unresolved''
***Cathartidae gen. et sp. indet. (Late Oligocene of Mongolia)
***Cathartidae gen. et sp. indet. (Late Miocene/Early Pliocene of Lee Creek Mine, US)
***Cathartidae gen. et sp. indet. (Middle Pliocene of Argentina)
***Cathartidae gen. et sp. indet. (Cuba)
**''Extant genera present in the fossil record''
***''Sarcoramphus'' (Middle Pliocene –? Recent)
****†''Sarcoramphus kernense, S. kernense (Kern vulture)
****†''Sarcoramphus, S. fischeri''
***''Gymnogyps'' (Early Pleistocene – Recent)
****†''Gymnogyps, G. howardae
****†''Gymnogyps, G. kofordi
***''Vultur'' (Pliocene – Recent)– distinctiveness disputed
****†''Vultur fossilis, V. fossilis'' [''Cathartes fossilis''; ''Sarcoramphus fossilis'']
*†Horusornithidae
**†''Horusornis, Horusornis vianeyliaudae'' (Late Eocene)
*Pandionidae – ospreys
**''Extant genera present in the fossil record''
***''Pandion (bird), Pandion'' (Early Oligocene – Recent)
****†''Pandion lovensis, P. lovensis''
****†''Pandion homalopteron, P. homalopteron''
*Sagittariidae – secretarybirds
**†''Amanuensis, Amanuensis pickfordi''
**†''Pelargopappus'' (Late Eocene/Early Oligocene – Late Oligocene/Early Miocene of France) – formerly ''Amphiserpentarius/Amynoptilon/Pelargopsis''
***†''Pelargopappus schlosseri, P. schlosseri''
***†''Pelargopappus magnus, P. magnus''
*Accipitridae – hawks, eagles and Old World vultures
**''Milvoides'' (Late Eocene)
**''Aquilavus'' (Late Eocene/Early Oligocene – Early Miocene)
**''Palaeocircus'' (Late Eocene/Early Oligocene)
**''Palaeastur'' (Early Miocene)
**''Pengana'' (Early Miocene)
**''Promilio'' (Early Miocene)
**''Proictinia'' (Early – Late Miocene/Early Pliocene)
**''Neophrontops'' (Early/middle Miocene – Late Pleistocene) – formerly in ''Neophron''
**''Mioaegypius'' (middle Miocene)
**''Apatosagittarius'' (Late Miocene)
**''Gansugyps'' (Late Miocene)
**''Palaeoborus'' (Miocene)
**''Qiluornis'' (Miocene)
**''Thegornis'' (Miocene)
**''Garganoaetus'' (Early Pliocene)
**''Amplibuteo'' (Late Pliocene of Peru – Late Pleistocene) – Belongs to the extant genus ''Buteogallus''
**''Cryptogyps'' (Middle – Late Pleistocene)
**''Neogyps'' (Late Pleistocene)
**''Palaeohierax'' – includes ''"Aquila" gervaisii''
**''Placement unresolved''
***Accipitridae gen. et sp. indet. AMNH FR 7434 (Early Eocene)
***Accipitridae gen. et sp. indet. (Early Oligocene)
***Accipitridae gen. et sp. indet. (Early/Middle Miocene)
***Accipitridae gen. et sp. indet. MPEF-PV-2523 (Late Miocene)
***Accipitridae gen. et sp. indet. (Early/Middle Pliocene) – ''Parabuteo''?
***Accipitridae gen. et sp. indet. (Late Pliocene/Early Pleistocene) – ''Buteo''?
***Accipitridae gen. et sp. indet. (Egypt)
***''"Aquila" danana'' (Late Miocene/Early Pliocene) – formerly also ''Geranoaetus'' or ''Buteo''
**''Extant genera present in the fossil record''
***''Haliaeetus'' (Early Oligocene – Recent)
***''Buteo'' (middle Oligocene – Recent)
***''Aquila (genus), Aquila'' (Middle Miocene – Recent)
***''Buteogallus'' (Middle Miocene – Recent) – might include ''Harpyhaliaetus''
***''"Hieraaetus"'' (Middle Miocene – Recent) – doubtfully distinct from ''Aquila''
***''Milvus'' (Early Pleistocene – Recent)
***''Gyps'' (Middle Pleistocene – Recent)
***''Aegypius'' (Middle Pleistocene – Recent)
**''Additional prehistoric species of extant genera''
***''Spizaetus grinnelli'' (Rancho La Brea Late Pleistocene of California, US) – formerly ''Geranoaetus'' or ''Buteo''
***''Spizaetus pliogryps'' – formerly ''Aquila''
***''Bearded vulture, Gypaetus georgii'' (Late Miocene)
***''Egyptian vulture, Neophron lolis'' (Late Miocene)
Falconiformes
*†Masillaraptoridae
**†''Masillaraptor, Masillaraptor parvunguis'' (middle Eocene of Messel, Germany)
**†''Danielsraptor, Danielsraptor phorusrhacoides'' (early Eocene of Essex, England)
*Falconidae – falcons
**†''Lagopterus, Lagopterus minutus''
**†''Parvulivenator, Parvulivenator watteli'' (Early Eocene)
**†''Stintonornis, Stintonornis mitchelli'' (Early Eocene)
**†''Badiostes, Badiostes patagonicus'' (Early Miocene)
**†''Pediohierax, Pediohierax ramenta'' [''Falco ramenta'' ] - (Middle Miocene)
**†''Thegornis, Thegornis musculosus'' [''Thegornis debilis'' ; ''Buteo musculosus'' ]
***†''Petrosushkinia, Petrosushkinia pliocaena'' [''Sushkinia'' ; ''Sushkinia pliocaena'' ] (Early Pliocene)
***†Falconidae gen. et sp. indet. (Late Miocene)
**''Extant genera present in the fossil record''
***''Milvago'' (Late Pleistocene -–Recent)
****†''Milvago, M. alexandri''
****†''Milvago, M. brodkorbi''
***''Falcon, Falco'' (Late Miocene – Recent)
****†''Milvago antiquus, F. antiquus''
****†''Milvago bakalovi, F. bakalovi''
****†''Milvago bulgaricus, F. bulgaricus''
****†''Milvago chowi, F. chowi''
****†''Milvago hezhengensis, F. hezhengensis''
****†''Milvago medius, F. medius''
****†''Milvago oregonus, F. oregonus''
****†''Milvago pisanus, F. pisanus''
****†''Milvago umanskajae, F. umanskajae''
Steatornithiformes
*''Paraprefica'' (Early Eocene?)
**†''Paraprefica, P. kelleri''
**†''Paraprefica, P. major''
*Steatornithidae – oilbirds
**†''Prefica, Prefica nivea''
**''Prehistoric species of extant genera''
***''Steatornis'' sp.
*Nyctibiidae – potoos
**†''Euronyctibius, Euronyctibius kurochkini''
Podargiformes
*†Fluvioviridavidae
**†''Fluvioviridavis, Fluvioviridavis platyrhamphus'' (Green River Early Eocene of N America)
*Podargidae – frogmouths
**†''Masillapodargus, Masillapodargus longipes''
**†''Quercypodargus, Quercypodargus olsoni''
Caprimulgiformes
*Caprimulgidae – nightjars
**†''Ventivorus, Ventivorus ragei''
**''Prehistoric species of extant genera''
***''Caprimulgus''
****†''Caprimulgus, C. piurensis''
Aegotheliformes
Owlet-nightjars
*Aegothelidae
**†''Quipollornis, Quipollornis koniberi'' (Early/Middle Miocene)
**''Extant genera present in the fossil record''
***''Aegotheles'' (Early/Middle Miocene – Recent)
****†''Aegotheles savesi, A. savesi'' (New Caledonian owlet-nightjar)
****†''Aegotheles, A. zealandivetus''
Apodiformes
Swifts and hummingbirds.
*†''Wyomingcypselus, Wyomingcypselus pohli''
*†''Procypseloides, Procypseloides mourerchauvireae'' (Late Eocene/Early Oligocene – Early Miocene)
*†''Laputavis, Laputavis robusta'' [''Laputa'' ; ''Laputa robusta'' ]
*†''Palescyvus, Palescyvus escampensis''
*†''Scaniacypselus'' (Early – middle Eocene)
**†''Scaniacypselus wardi, S. wardi''
**†''Scaniacypselus szarskii, S. szarskii'' [''Aegialornis szarskii'' ; ''Primapus szarskii'' ]
*†Aegialornithidae
**†''Primapus, Primapus lacki'' [?''Procuculus minutus'' ] (Early Eocene) ?apodid
**†''Aegialornis'' [''Tachyornis'' ; ''Belornis'' ; ''Mesogiornis'' ](Early? – Late Eocene)
***†''Aegialornis, A. leenhardti''
***†''Aegialornis, A. germanicus''
***†''Aegialornis, A. gallicus'' [''Tachyornis hirundo'' ]
***†''Aegialornis, A. wetmorei'' [''Mesogiornis wetmorei'' ]
***†''Aegialornis, A. broweri'' [''Mesogiornis broweri'' ]
*†Eocypselidae – apodiform (hemiprocnid?)? caprimulgiform? basal Cypselomorphae?
**†''Eocypselus'' (Late Paleocene ?- Early Eocene of NC Europe)
***†''Eocypselus vincenti, E. vincenti''
***†''Eocypselus rowei, E. rowei''
*†Cypselavidae
**†''Parargornis, Parargornis messelensis'' (middle Eocene) jungornithid, trochilid, basal as ''Argornis''?
**†''Argornis, Argornis caucasicus'' (Late Eocene) ?Trochilidae
**†''Cypselavus, Cypselavus gallicus'' (Late Eocene Early Oligocene) aegialornithid or hemiprocnid
*†Jungornithidae
**†''Jungornis'' (Early Oligocene of N Caucasus, Russia)
***†''Jungornis tesselatus, J. tesselatus''
***†''Jungornis geraldmayri, J. geraldmayri''
*Trochilidae hummingbirds
**†''Eurotrochilus'' (Early Oligocene)
***†''Eurotrochilus inexpectatus, E. inexpectatus''
***†''Eurotrochilus, E. noniewiczi''
**''Placement unresolved''
***Trochilidae sp. et gen. indet. (Bahamas, West Indies)
***Trochilidae sp. et gen. indet. (Brazil)
*Apodidae – swifts
**''Extant genera present in the fossil record''
***''Collocalia'' (Early Miocene – Recent)
***''Apus (genus), Apus'' (Middle/Late Miocene – Recent)
***''Chaetura'' (Late Miocene – Recent)
***''Tachornis'' (Late Pleistocene – Recent)
Coliiformes
Mousebirds and relatives
*Unresolved and basal forms
** †'' Botauroides parvus'' (Eocene of Wyoming, US)
** †''Eobucco, Eobucco brodkorbi'' - sandcoleid?
** †''Eocolius, Eocolius walkeri'' (London Clay Early Eocene of Walton-on-the-Naze, England) - sandcoleid or coliid
**†''Limnatornis'' (Early Miocene of Saint-Gérand-le-Puy, France) - coliid? (''Urocolius''?)
** †Coliiformes gen. et sp. indet. (Late Miocene of Kohfidisch, Austria)[Similar to ''Urocolius'' and ''Limnatornis'' (if distinct): Mlíkovský (2002)]
**†''Uintornis'' - sandcoleid?
* Family †Chascacocoliidae
** Genus †''Chascacocolius'' (Late Paleocene ?- Early Eocene) - basal? sandcoleid?
* Family †Selmeidae
**†''Selmes, Selmes absurdipes'' (Middle Eocene ?-Late Oligocene of C Europe) - coliid? (synonym of ''Primocolius''?)
* Family †Sandcoleidae
**†''Sandcoleus, Sandcoleus copiosus'' (Paleocene)
**†''Anneavis, Anneavis anneae''
**†''Eoglaucidium, Eoglaucidium pallas''
* Family Coliidae
**†''Primocolius'' (Late Eocene/Oligocene of Quercy, France)
**†''Oligocolius'' (Early Oligocene of Frauenweiler, Germany)
**†''Masillacolius, Masillacolius brevidactylus'' (middle Eocene of Messel, Germany)
**''Extant genera present in the fossil record''
*** ''Colius'' [''Necrornis'' ]
†Zealandornithidae
* †''Zealandornis, Zealandornis relictus'' (Early Miocene of Otago, New Zealand)
Strigiformes
Owls and barn owls
*Unresolved and basal forms
**''Berruornis'' (Late Paleocene) – basal? Sophornithidae?
**Strigiformes gen. et sp. indet. (Late Paleocene)
**''Palaeoglaux'' (middle – Late Eocene) – own family Palaeoglaucidae or Strigidae?
**''Palaeobyas'' (Late Eocene/Early Oligocene) – Tytonidae? Sophiornithidae?
**''Palaeotyto'' (Late Eocene/Early Oligocene) – Tytonidae?
**Strigiformes gen. et spp. indet. (Early Oligocene)
*Ogygoptyngidae
**''Ogygoptynx'' (Middle/Late Paleocene)
*Protostrigidae
**''Eostrix'' (Early – middle Eocene)
**''Minerva (bird), Minerva'' (middle – Late Eocene) – formerly ''Protostrix'', includes ''"Aquila" ferox'', ''"Aquila" lydekkeri'', and ''"Bubo" leptosteus''
** ''Oligostrix'' (middle Oligocene)
*Sophiornithidae
**''Sophiornis''
*Strigidae – typical owls
**''Mioglaux'' (Late Oligocene? – Early Miocene) – includes ''"Bubo" poirreiri''
**''Intulula'' (Early/Middle –? Late Miocene) – includes ''"Strix/Ninox" brevis''
**''Alasio'' (Middle Miocene) – includes ''"Strix" collongensis''
**''Oraristrix'' (Late Pleistocene)
**''Miosurnia'' (Late Miocene)
**''Placement unresolved''
***''Otus wintershofensis, "Otus" wintershofensis'' (Early/Middle Miocene) – may be close to extant genus ''Ninox''
***''Strix edwardsi, "Strix" edwardsi'' (Middle/Late? Miocene)
***''Asio pygmaeus, "Asio" pygmaeus'' (Early Pliocene)
***Strigidae ''gen. et sp. indet.'' UMMP V31030 (Late Pliocene) – ''Strix/Bubo''?
**''Extant genera present in the fossil record''
***''Strix (bird), Strix'' (Early Miocene – Recent)
***''Bubo (bird), Bubo'' (Late Miocene? – Recent)
***''Asio'' (Late Pliocene – Recent)
***''Athene (bird), Athene'' (Late Pliocene – Recent)
***''Glaucidium (bird), Glaucidium'' (Late Pliocene – Recent)
***''Surnia'' (Late Pliocene – Recent)
***''Pulsatrix'' (Late Pleistocene – Recent)
*Tytonidae – barn owls
**''Nocturnavis'' (Late Eocene/Early Oligocene)
**''Selenornis'' (Late Eocene/Early Oligocene)
**''Necrobyas'' (Late Eocene/Early Oligocene – Early Miocene)
**''Prosybris'' (Early Oligocene? – Early Miocene)
**''Placement unresolved''
***Tytonidae ''gen. et sp. indet.'' TMT 164 (Middle Miocene)
**''Extant genera present in the fossil record''
***''Tyto'' (Late Miocene – Recent)
Coraciiformes
Rollers and allies. Probably paraphyletic.
*Basal and unresolved forms
**†''Quasisyndactylus, Quasisyndactylus longibrachis'' (middle Eocene) – alcediniform, basal?
**†''Paracoracias, Paracoracias occidentalis''
**†''Cryptornis, Cryptornis antiquus'' (Late Eocene) – bucerotid? geranopterid?
**†''Protornis, Protornis glarniensis'' (Oligocene) – basal to motmotids and meropids?
**† Coraciiformes gen. et spp. indet. PQ 1216, QU 15640 (Late Eocene) – 2 species
*†Geranopteridae
**†''Geranopterus'' (Late Eocene – Early Miocene)
***†''Geranopterus, G. alatus''
***†''Geranopterus, G. bohemicus'' [''Nupharanassa bohemica'' ]
***†''Geranopterus, G. milneendwardsi''
*†Eocoraciidae
**†''Eocoracias, Eocoracias brachyptera'' (middle Eocene)
*†Primobucconidae
**†''Primobucco'' [''Neanis'' ] (Early – middle Eocene)
***†''Primobucco, P. kestneri'' [''Neanis kestneri'' ]
***†''Primobucco, P. frugilegus''
***†''Primobucco, P. mcgrewi''
***†''Primobucco, P. perneri''
*Todidae – todies
**†''Palaeotodus'' Olson 1976 (Late Eocene of France – Early Oligocene of WC Europe and Wyoming)
***†''Palaeotodus emryi, P. emryi''
***†''Palaeotodus escampsiensis, P. escampsiensis''
***†''Palaeotodus itardiensis, P. itardiensis''
*Motmotidae – motmots
**Placement unresolved
***Momotidae gen. et sp. indet. (Late Miocene)
*Coraciidae
**†''Miocoracias, Miocoracias chenevali''
**''Extant genera present in the fossil record''
***''Eurystomus''
****†''Eurystomus, E. beremedensis''
Bucerotiformes
*†Messelirrisoridae
**†''Messelirrisor'' Mayr 1998 (middle Eocene)
***†''Messelirrisor, M. halcyorostris''
***†''Messelirrisor, M. parvus''
***†''Messelirrisor, M. grandis''
*Phoeniculidae (woodhoopoes)
**†''Phirriculus, Phirriculus pinicola''
*Bucerotidae – hornbills
**†''Euroceros, Euroceros bulgaricus'' (Late Miocene of Hadzhidimovo, Bulgaria)
**''Extant genera present in the fossil record''
***''Bucorvus''
****†''Bucorvus brailloni, B. brailloni''
Trogoniformes
*Trogonidae – trogons
**†''Foshanornis, Foshanornis songi''
**†''Septentrogon, Septentrogon madseni'' Kristoffersen 2002 (Fur Late Paleocene/Early Eocene of Ejerslev, Denmark)
**†''Paratrogon, Paratrogon gallicus'' [''Trogon gallicus'' ] (Early Miocene of France)
**†''Primotrogon'' [''Masillatrogon'' ] (middle Eocene of Messel, Germany? - Early Oligocene of France)
***†''Primotrogon wintersteini, P. wintersteini''
***†''Primotrogon pumilio, P. pumilio'' [''Masillatrogon pumilio'' ]
**''Placement unresolved''
***Trogonidae gen. et sp. indet. 1 (NW Europe)
***Trogonidae gen. et sp. indet. 2 (NW Europe)
Piciformes
*Placement unresolved
**Piciformes gen. et sp. indet. IRScNB Av 65 (Early Oligocene)
**''Rupelramphastoides'' (Early Oligocene) – ramphastid?
**Piciformes gen. et sp. indet. SMF Av 429 (Late Oligocene)
**''Capitonides'' (Early – Middle Miocene) – ramphastid? "capitonid" (Lybiidae, Megalaimidae)? own family Capitonididae?
**Pici gen. et sp. indet. (Middle Miocene) – "capitonid" (Lybiidae, Megalaimidae?)
*Miopiconidae
**''Miopico''
*Lybiidae – African barbets
**Lybiidae gen. et sp. indet. (Late Miocene) – extant genus ''Pogoniulus''?
*Galbulidae
**''Extant genera present in the fossil record''
***''Galbula hylochoreutes'' (Middle Miocene of La Venta, Colombia)
*Picavidae
**''Picavus''
*Picidae – woodpeckers
**''Palaeopicus'' (Late Oligocene)
**''Palaeonerpes'' (Early Pliocene)
**''Pliopicus'' (Early Pliocene)
**''Placement unresolved''
***Picidae gen. et sp. indet. (Middle Miocene)
***Picidae gen. et sp. indet. (Late Miocene)
***cf. ''Colaptes'' DMNH 1262 (Early Pliocene of Ainsworth, US)
**''Extant genera present in the fossil record''
***''Campephilus'' (Late Pleistocene – Recent)
***''Colaptes''
***''Dendrocopos''
***''Additional prehistoric subspecies of extant species''
****''West Indian woodpecker, Melanerpes superciliaris'' ssp. (Little Exuma, Bahamas)
****''West Indian woodpecker, Melanerpes superciliaris'' ssp. (New Providence, Bahamas)
Passeriformes
*Placement unresolved
**Passeriformes gen. et spp. indet. (Early Eocene) – several species, oscine?
**''Resoviaornis'' (Early Oligocene)
**''Wieslochia'' (Early Oligocene)
**Passeriformes gen. et spp. indet. (Late Oligocene) – several suboscine and oscine species
**''Certhiops'' (Early Miocene of Germany) – basal Certhioidea
**Passeriformes gen. et sp. indet. (Early/Middle Miocene) – suboscine?
**Passeriformes gen. et spp. indet. (Early/Middle Miocene) – several species, oscine?
**Passeriformes gen. et spp. indet. (Middle Miocene) – several species, basal?
**Passeriformes gen. et spp. indet. (Middle Miocene) – several species, oscine?
**Passeriformes gen. et spp. indet. (Late Miocene) – Sylvioidea
**''"Palaeostruthus" eurius'' (Pliocene)
*Eurylaimidae – broadbills
**''Placement unresolved''
***Eurylaimidae gen. et sp. indet. (Early Miocene)
*Palaeoscinidae
**''Palaeoscinis'' (Late Miocene)
*Furnariidae – ovenbirds
**†''Pseudoseisuropsis''
***†''Pseudoseisuropsis, P. nehuen'' (Early Pleistocene of Argentina)
***†''Pseudoseisuropsis, P. cuelloi'' (Late Pleistocene of Uruguay)
***†''Pseudoseisuropsis, P. wintu'' (Early Pleistocene of Argentina)
**''Prehistoric species of extant genera''
***''Pseudoseisura cursor'' (Ensenada Early/Middle Pleistocene of Anchorena, Argentina)
***''Cinclodes major'' (Middle Pleistocene of Buenos Aires Province, Argentina)
*Menuridae – lyretails
**''Extant genera present in the fossil record''
***''Menura'' (Early Miocene – Recent)
*Meliphagidae – honeyeaters
**''Placement unresolved''
***Meliphagidae gen. et spp. indet. (Middle/Late Miocene – Pliocene of Riversleigh, Australia) – at least 7 spp., some may be from extant genera
*Orthonychidae – logrunners
**''Extant genera present in the fossil record''
***''Orthonyx'' (Middle/Late Miocene – Recent)
*Oriolidae – Old World orioles
**''Longmornis'' (Early Miocene of Riversleigh, Australia)
*Artamidae – woodswallows, butcherbirds, currawongs and Australian magpie
**''Placement unresolved''
***Artamidae gen. et sp. indet. (Early/Middle Miocene) – cracticine
*Corvidae – crows, ravens, jays and magpies
**''Miocorvus'' (Middle Miocene)
**''Miopica'' (Middle Miocene)
**''Miocitta'' (Late Miocene)
**''Protocitta'' (Early Pleistocene)
**''Henocitta'' (Middle Pleistocene)
**''Extant genera present in the fossil record''
***''Corvus (genus), Corvus'' (Late Miocene — Recent)
***''Pica (genus), Pica'' (Late Pliocene/Early Pleistocene – Recent)
***''Pyrrhocorax''
**''Placement unresolved''
***Corvidae gen. et sp. indet. (Early Pliocene)
***Corvidae gen. et sp. indet. (Early/Middle Pleistocene) – probably belongs in extant genus
*Laniidae – shrikes
**''Extant genera present in the fossil record''
***''Lanius'' (Early Miocene – Recent)
*Regulidae – kinglets
**''Extant genera present in the fossil record''
***''Regulus (bird), Regulus'' (Late Pliocene – Recent)
*Hirundinidae – swallows and martins
**''Placement unresolved''
***Hirundinidae gen. et spp. indet. (Early Pliocene of Langebaanweg, South Africa) – 2 species
*Megaluridae – grass-warblers and allies
**''Extant genera present in the fossil record''
***?''Locustella'' (Late Miocene – Recent)
*Acrocephalidae – marsh- and tree-warblers
**''Extant genera present in the fossil record''
***?''Acrocephalus (bird), Acrocephalus'' (Late Miocene – Recent)
*Muscicapidae – Old World flycatchers and chats
**''Extant genera present in the fossil record''
***''Luscinia'' (Late Miocene – Recent)
*Turdidae – thrushes
**''Extant genera present in the fossil record''
***?''Turdus'' (Middle? Miocene – Recent)
*Alaudidae – larks
**''Eremarida'' (Late Miocene of Hrabarsko, Bulgaria)
*Motacillidae – wagtails
**''Extant genera present in the fossil record''
***''Motacilla''
*Fringillidae – finches
**''Extant genera present in the fossil record''
***''Loxia'' (Late Pliocene – Recent)
**''Additional prehistoric species of extant genera''
***''Coccothraustes simeonovi'' (Late Pliocene of Varshets, Bulgaria)
***''Coccothraustes balcanicus''
*Icteridae – grackles and New World orioles
**''Pandanaris'' (Pleistocene)
**''Pyelorhamphus'' (Pleistocene)
**''Extant genera present in the fossil record''
***''Euphagus'' (Late Pleistocene – Recent)
*Cardinalidae – cardinals
**''Placement unresolved''
***''Passerina'' sp. (Early Pliocene of Yepómera, Mexico)
*Emberizidae – buntings and New World sparrows
**''Pampaemberiza'' (Middle Pleistocene of Necochea, Argentina)
**''Extant genera present in the fossil record''
***''Ammodramus'' (Late Miocene – Recent) – including ''Palaeostruthus''
**''Additional prehistoric species of extant genera''
***''Pipilo angelensis'' (Pleistocene of Rancho La Brea, US)
Avialans ''incertae sedis''
These fossil taxa cannot be assigned to any major group with reasonable certainty. The "proto-birds" above are of some indeterminate basal position in the entire avialan (and paravian) radiation, but known from such diagnostic material that their relationships at the family (biology), family level are known. In contrast, the taxa here have a hypodigm that is usually just sufficient for giving them a valid scientific name, but not for phylogenetic purposes beyond classing them as pygostylians or more modern birds. Some, however, are known from such fragmentary remains that the possibility that they are non-avian "reptiles" such as dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s cannot be ruled out at present.
*†''"Ichthyornis" minusculus'' Nesov 1990 (Bissekty Late Cretaceous of Kyzyl Kum, Uzbekistan) – enantiornithine?
*†''Qinornis, Qinornis paleocenica'' Xue 1995 (Early/Middle Paleocene) – enantiornithine? neornithine?
See also
*Bird ichnology
*Dominant group (extinction)
*Feathered dinosaurs
*List of recently extinct birds
*Late Quaternary prehistoric birds
*List of paleognaths
*List of Galliformes
*List of Columbidae species
*Flightless birds
*Origin of birds
*Prehistoric life
Footnotes
References
* Chiappe, Luis M. (2001): The rise of birds. ''In'': Derek Briggs, Briggs, Derek E.G. & Crowther, P.R. (eds.): ''Palaeobiology II: A Synthesis'': 102-106. Cambridge University Press, Cambridge, UK.
* Chiappe, Luis M. (2002): Basal bird phylogeny: problems and solutions. ''In:'' Chiappe, L.M. and Witmer, L.M. (eds.): ''Mesozoic Birds: Above the Heads of Dinosaurs'': 448-472. University of California Press, Berkeley, US.
*
Electronic Appendix
* Jacques Gauthier, Gauthier, Jacques A. & de Queiroz, Kevin (2001): Feathered dinosaurs, flying dinosaurs, crown dinosaurs, and the name "Aves". ''In:'' Gauthier, Jacques & Gall, L.F. (eds.): ''New Perspectives on the Origin and Early Evolution of Birds: Proceedings of the International Symposium in Honor of John H. Ostrom'': 7-41. Peabody Museum of Natural History, Yale University, New Haven, Connecticut.
* Mortimer, Michael (2004): The Theropod Database
Phylogeny of taxa
Retrieved 2013-MAR-02.
* Storrs Olson, Olson, Storrs L. (1985): The fossil record of birds. ''In:'' Farner, D.S.; King, J.R. & Parkes, Kenneth C. (eds.): ''Avian Biology'' 8: 79-238. Academic Press, New York. Not in copyright
PDF fulltext
* Paul Sereno, Sereno, Paul Callistus (2005):
TaxonSearch: Stem Archosauria
'. Version 1.0, November 7, 2005. Retrieved 2007-APR-30.
External links
by Ben Creisler. Version of 2003-JUL-7. Retrieved 2007-SEP-1.
*
*
{{DEFAULTSORT:Fossil Birds
Birds by classification, Fossil
Prehistoric bird genera,
Lists of prehistoric birds
Taxonomic lists (genera, taxonomic)
Ornithology lists