Lithornithidae
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Lithornithidae
Lithornithidae is an extinct, possibly paraphyletic (but see below) group of early paleognath birds. They are known from fossils dating to the Upper Paleocene through the Middle Eocene of North America and Europe, with possible Late Cretaceous representatives. All are extinct today; the youngest specimen is the currently unnamed SGPIMH MEV1 specimen from the mid-Eocene Messel Pit site. Lithornithids had long, slender, bills for probing. They closely resembled modern tinamous. They possessed a rhynchokinetic skull with relatively unfused cranial bones, a weakly fused pygostyle and a splenial. The unguals were more curved than in tinamous and probably allowed better perching in trees. The order Lithornithiformes was erected by Dr. Peter Houde in 1988. Initially, only three genera ('' Lithornis'', '' Paracathartes'', and '' Pseudocrypturus'') and eight named species were included. '' Promusophaga'' (Harrison & Walker, 1977) originally considered a stem-turaco The turacos m ...
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Calciavis Grandei
''Calciavis'' is an extinct genus of bird from the Eocene of Wyoming. It is a lithornithid, a member of a lineage of flying palaeognaths, distantly related to modern ratites. Like many other fossils from the Green River Formation, it is exceptionally well preserved, bearing soft-tissue impressions of skin and feathers. Discovery and Description The original specimen of ''Calciavis'', AMNH 30578, was found in the Fossil Butte Member of the Green River Formation, Wyoming, dating to around 51 million years ago. It is an exceptionally well preserved specimen, preserving most of the post-cranial skeleton aside from the femur and associated pelvic region, as well as soft-tissue impressions of feathers, skin, Bird anatomy#Scales, foot scales and claw sheaths. Another specimen, AMNH 30560 , was found in the vicinity, composed of a similarly complete most mostly disarticulate skeleton. It is diagnosed by several features of the skull and pelvis as well as having metatarsals IV and II sube ...
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Lithornis
''Lithornis'' is a genus of extinct paleognathous birds. Although ''Lithornis'' was able to fly well, their closest relatives are the extant tinamous (which are poor flyers) and ratites (which are flightless birds). Fossils of ''Lithornis'' are known with certainty from the Upper Paleocene through the Middle Eocene, but their fossil record may extend to the late Cretaceous. "''Lithornis''" is from ancient Greek for "stone bird", as it is one of the first fossil birds to become widely discussed. Presumably closely related genera are ''Paracathartes'' and '' Pseudocrypturus''. Species Six species have been recognized in modern times; undescribed ones are also likely to exist. The supposed tarsometatarsus piece from which ''"Lithornis" emuinus'' was described is actually a humerus fragment of the giant pseudotooth bird ''Dasornis''.Houde, P. (1988) ''Lithornis celetius'' ''L. celetius'' is from the Bangtail Quarry, Sedan Quadrangle, Park County, Montana, USA, and was describe ...
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Lithornis Plebius
''Lithornis'' is a genus of extinct paleognathous birds. Although ''Lithornis'' was able to fly well, their closest relatives are the extant tinamous (which are poor flyers) and ratites (which are flightless birds). Fossils of ''Lithornis'' are known with certainty from the Upper Paleocene through the Middle Eocene, but their fossil record may extend to the late Cretaceous. "''Lithornis''" is from ancient Greek for "stone bird", as it is one of the first fossil birds to become widely discussed. Presumably closely related genera are ''Paracathartes'' and '' Pseudocrypturus''. Species Six species have been recognized in modern times; undescribed ones are also likely to exist. The supposed tarsometatarsus piece from which ''"Lithornis" emuinus'' was described is actually a humerus fragment of the giant pseudotooth bird ''Dasornis''.Houde, P. (1988) ''Lithornis celetius'' ''L. celetius'' is from the Bangtail Quarry, Sedan Quadrangle, Park County, Montana, USA, and was describe ...
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Fissuravis
Fissuravis ("fissure bird") is a genus of extinct bird from the Paleocene of Germany. A lithornithid, it was closely related to modern ratites, but it was a capable flyer.Gerald Mayr, The Birds from the Paleocene Fissure Filling of Walbeck (Germany), Journal of Vertebrate Paleontology 27(2):394–408, June 2007 Ecology Hailing from the Walbeck Paleocene deposits, it is found amidst a rich avian fauna, which also included the gigantic ''Gastornis'' and the enigmatic ratite ''Remiornis''. Like most lithornithids, it was probably a very competent flyer, its coracoid remnants suggesting powerful flight musculature, and it likely engaged in a similar style of soaring flight to ''Lithornis'' and ''Pseudocrypturus ''Pseudocrypturus'' is a genus of extinct paleognathous bird. One species is known, ''Pseudocrypturus cercanaxius''. It is a relative of such modern birds as ostriches. It lived in the early Eocene. The holotype fossil is in the collection of th ...''. References {{ ...
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Lithornis Celetius
''Lithornis'' is a genus of extinct paleognathous birds. Although ''Lithornis'' was able to fly well, their closest relatives are the extant tinamous (which are poor flyers) and ratites (which are flightless birds). Fossils of ''Lithornis'' are known with certainty from the Upper Paleocene through the Middle Eocene, but their fossil record may extend to the late Cretaceous. "''Lithornis''" is from ancient Greek for "stone bird", as it is one of the first fossil birds to become widely discussed. Presumably closely related genera are ''Paracathartes'' and '' Pseudocrypturus''. Species Six species have been recognized in modern times; undescribed ones are also likely to exist. The supposed tarsometatarsus piece from which ''"Lithornis" emuinus'' was described is actually a humerus fragment of the giant pseudotooth bird ''Dasornis''.Houde, P. (1988) ''Lithornis celetius'' ''L. celetius'' is from the Bangtail Quarry, Sedan Quadrangle, Park County, Montana, USA, and was describe ...
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Paracathartes
''Paracathartes'' is a genus of extinct bird from the Wasachtian horizon of lower Eocene Wyoming. One species, ''Paracathartes howardae'' has been described. It is a paleognathous bird, turkey-like in stature and size, that probably resembled a tinamou quite closely. ''Paracathartes'' was described by Harrison as the earliest known cathartid vulture. Rich criticized this assignment.Rich, P. V. (1983) Houde (1988) included it as a member of the order Lithornithiformes and family Lithornithidae.Houde, Peter W. (1988) The holotype specimen is in the collection of the Royal Ontario Museum. It has catalog number ROM 22658. It is the distal end of a left tibiotarsus. It was collected by G. E. Lindblad and G. Sternberg on 4 August 1949. It was found at the northernmost branch of Elk Creek, near Basin, Wyoming. The horizon is Greybullian, middle Wasatchian (early Eocene), Willwood Formation, Bighorn Basin. Further specimens of ''Paracathartes'' were collected, including almost the ...
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Messel Pit
The Messel pit (german: Grube Messel) is a disused quarry near the village of Messel ( Landkreis Darmstadt-Dieburg, Hesse) about southeast of Frankfurt am Main, Germany. Bituminous shale was mined there. Because of its abundance of well-preserved fossils dating from the middle of the Eocene, it has significant geological and scientific importance. Over 1000 species of plants and animals have been found at the site. After almost becoming a landfill, strong local resistance eventually stopped these plans and the Messel Pit was declared a UNESCO World Heritage Site on 9 December 1995. Significant scientific discoveries about the early evolution of mammals and birds are still being made at the Messel Pit, and the site has increasingly become a tourist site as well. History Brown coal and later oil shale was actively mined from 1859. The pit first became known for its wealth of fossils around 1900, but serious scientific excavation only started around the 1970s, when falling oil ...
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Lithornis Promiscuus
''Lithornis'' is a genus of extinct paleognathous birds. Although ''Lithornis'' was able to fly well, their closest relatives are the extant tinamous (which are poor flyers) and ratites (which are flightless birds). Fossils of ''Lithornis'' are known with certainty from the Upper Paleocene through the Middle Eocene, but their fossil record may extend to the late Cretaceous. "''Lithornis''" is from ancient Greek for "stone bird", as it is one of the first fossil birds to become widely discussed. Presumably closely related genera are ''Paracathartes'' and '' Pseudocrypturus''. Species Six species have been recognized in modern times; undescribed ones are also likely to exist. The supposed tarsometatarsus piece from which ''"Lithornis" emuinus'' was described is actually a humerus fragment of the giant pseudotooth bird ''Dasornis''.Houde, P. (1988) ''Lithornis celetius'' ''L. celetius'' is from the Bangtail Quarry, Sedan Quadrangle, Park County, Montana, USA, and was describe ...
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Lithornis Nasi
''Lithornis'' is a genus of extinct paleognathous birds. Although ''Lithornis'' was able to fly well, their closest relatives are the extant tinamous (which are poor flyers) and ratites (which are flightless birds). Fossils of ''Lithornis'' are known with certainty from the Upper Paleocene through the Middle Eocene, but their fossil record may extend to the late Cretaceous. "''Lithornis''" is from ancient Greek for "stone bird", as it is one of the first fossil birds to become widely discussed. Presumably closely related genera are ''Paracathartes'' and '' Pseudocrypturus''. Species Six species have been recognized in modern times; undescribed ones are also likely to exist. The supposed tarsometatarsus piece from which ''"Lithornis" emuinus'' was described is actually a humerus fragment of the giant pseudotooth bird ''Dasornis''.Houde, P. (1988) ''Lithornis celetius'' ''L. celetius'' is from the Bangtail Quarry, Sedan Quadrangle, Park County, Montana, USA, and was describe ...
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Lithornis Hookeri
''Lithornis'' is a genus of extinct paleognathous birds. Although ''Lithornis'' was able to fly well, their closest relatives are the extant tinamous (which are poor flyers) and ratites (which are flightless birds). Fossils of ''Lithornis'' are known with certainty from the Upper Paleocene through the Middle Eocene, but their fossil record may extend to the late Cretaceous. "''Lithornis''" is from ancient Greek for "stone bird", as it is one of the first fossil birds to become widely discussed. Presumably closely related genera are ''Paracathartes'' and '' Pseudocrypturus''. Species Six species have been recognized in modern times; undescribed ones are also likely to exist. The supposed tarsometatarsus piece from which ''"Lithornis" emuinus'' was described is actually a humerus fragment of the giant pseudotooth bird ''Dasornis''.Houde, P. (1988) ''Lithornis celetius'' ''L. celetius'' is from the Bangtail Quarry, Sedan Quadrangle, Park County, Montana, USA, and was describe ...
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Promusophaga
''Lithornis'' is a genus of extinct paleognathous birds. Although ''Lithornis'' was able to fly well, their closest relatives are the extant tinamous (which are poor flyers) and ratites (which are flightless birds). Fossils of ''Lithornis'' are known with certainty from the Upper Paleocene through the Middle Eocene, but their fossil record may extend to the late Cretaceous. "''Lithornis''" is from ancient Greek for "stone bird", as it is one of the first fossil birds to become widely discussed. Presumably closely related genera are ''Paracathartes'' and '' Pseudocrypturus''. Species Six species have been recognized in modern times; undescribed ones are also likely to exist. The supposed tarsometatarsus piece from which ''"Lithornis" emuinus'' was described is actually a humerus fragment of the giant pseudotooth bird ''Dasornis''.Houde, P. (1988) ''Lithornis celetius'' ''L. celetius'' is from the Bangtail Quarry, Sedan Quadrangle, Park County, Montana, USA, and was describe ...
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Paleognathae
Palaeognathae (; ) is a infraclass of birds, called paleognaths, within the class Aves of the clade Archosauria. It is one of the two extant infraclasses of birds, the other being Neognathae, both of which form Neornithes. Palaeognathae contains five extant branches of flightless lineages (plus two extinct clades), termed ratites, and one flying lineage, the Neotropic tinamous.Wetmore, A. (1960) A Classification for Birds of the World. Smithsonian Miscellaneous Collection (Washington D.C.: Smithsonian Institution) 139: 1–37. There are 47 species of tinamous, five of kiwis (''Apteryx''), three of cassowaries (''Casuarius''), one of emus (''Dromaius'') (another became extinct in historic times), two of rheas (''Rhea'') and two of ostrich (''Struthio'').Clements, J. C. ''et al''. (2010) Recent research has indicated that paleognaths are monophyletic but the traditional taxonomic split between flightless and flighted forms is incorrect; tinamous are within the ratite radiation, me ...
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