Hydroponic
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Hydroponics is a type of
horticulture Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...
and a subset of
hydroculture Hydroponics is a type of horticulture and a subset of hydroculture which involves growing plants, usually crops or medicinal plants, without soil, by using water-based mineral nutrient solutions in aqueous solvents. Terrestrial or aquatic ...
which involves growing
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
s, usually crops or medicinal plants, without
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former te ...
, by using
water Water (chemical formula ) is an inorganic, transparent, tasteless, odorless, and nearly colorless chemical substance, which is the main constituent of Earth's hydrosphere and the fluids of all known living organisms (in which it acts as a ...
-based
mineral In geology and mineralogy, a mineral or mineral species is, broadly speaking, a solid chemical compound with a fairly well-defined chemical composition and a specific crystal structure that occurs naturally in pure form.John P. Rafferty, ed. ( ...
nutrient A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...
solution Solution may refer to: * Solution (chemistry), a mixture where one substance is dissolved in another * Solution (equation), in mathematics ** Numerical solution, in numerical analysis, approximate solutions within specified error bounds * Soluti ...
s in aqueous
solvent A solvent (s) (from the Latin '' solvō'', "loosen, untie, solve") is a substance that dissolves a solute, resulting in a solution. A solvent is usually a liquid but can also be a solid, a gas, or a supercritical fluid. Water is a solvent for ...
s.
Terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
or
aquatic plant Aquatic plants are plants that have adapted to living in aquatic environments (saltwater or freshwater). They are also referred to as hydrophytes or macrophytes to distinguish them from algae and other microphytes. A macrophyte is a plant that ...
s may grow with their
root In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the sur ...
s exposed to the nutritious
liquid A liquid is a nearly incompressible fluid that conforms to the shape of its container but retains a (nearly) constant volume independent of pressure. As such, it is one of the four fundamental states of matter (the others being solid, gas, a ...
or in addition, the roots may be mechanically supported by an inert medium such as
perlite Perlite is an amorphous volcanic glass that has a relatively high water content, typically formed by the hydration of obsidian. It occurs naturally and has the unusual property of greatly expanding when heated sufficiently. It is an industrial ...
,
gravel Gravel is a loose aggregation of rock fragments. Gravel occurs naturally throughout the world as a result of sedimentary and erosive geologic processes; it is also produced in large quantities commercially as crushed stone. Gravel is classifi ...
, or other substrates. Despite inert media, roots can cause changes of the
rhizosphere The rhizosphere is the narrow region of soil or substrate that is directly influenced by root secretions and associated soil microorganisms known as the root microbiome. Soil pores in the rhizosphere can contain many bacteria and other microor ...
pH and
root exudate Plant root exudates are fluids emitted through the roots of plants. These secretions influence the rhizosphere around the roots to inhibit harmful microbes and promote the grow of self and kin plants. Plant root systems can grow to be complex due t ...
s can affect rhizosphere
biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary i ...
and physiological balance of the nutrient solution by
secondary metabolite Secondary metabolites, also called specialised metabolites, toxins, secondary products, or natural products, are organic compounds produced by any lifeform, e.g. bacteria, fungi, animals, or plants, which are not directly involved in the norm ...
s.
Transgenic plants Genetically modified plants have been engineered for scientific research, to create new colours in plants, deliver vaccines, and to create enhanced crops. Plant genomes can be engineered by physical methods or by use of '' Agrobacterium'' for the ...
grown hydroponically allow the release of pharmaceutical proteins as part of the root exudate into the hydroponic medium. The
nutrients A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excre ...
used in hydroponic systems can come from many different organic or
inorganic In chemistry, an inorganic compound is typically a chemical compound that lacks carbon–hydrogen bonds, that is, a compound that is not an organic compound. The study of inorganic compounds is a subfield of chemistry known as '' inorganic chemist ...
sources, including
fish excrement Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...
,
duck Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a form t ...
manure Manure is organic matter that is used as organic fertilizer in agriculture. Most manure consists of animal feces; other sources include compost and green manure. Manures contribute to the Soil fertility, fertility of soil by adding organic ma ...
, purchased
chemical fertilizer A fertilizer (American English) or fertiliser (British English; see spelling differences) is any material of natural or synthetic origin that is applied to soil or to plant tissues to supply plant nutrients. Fertilizers may be distinct from ...
s, or
artificial Artificiality (the state of being artificial or manmade) is the state of being the product of intentional human manufacture, rather than occurring naturally through processes not involving or requiring human activity. Connotations Artificiality ...
nutrient solutions. Plants are commonly grown hydroponically in a
greenhouse A greenhouse (also called a glasshouse, or, if with sufficient heating, a hothouse) is a structure with walls and roof made chiefly of Transparent ceramics, transparent material, such as glass, in which plants requiring regulated climatic condit ...
or contained environment on inert media, adapted to the
controlled-environment agriculture Controlled-environment agriculture (CEA) -- which includes indoor agriculture (IA) and vertical farming -- is a technology-based approach toward food production. The aim of CEA is to provide protection from the outdoor elements and maintain opti ...
(CEA) process. Plants commonly grown hydroponically include
tomatoes The tomato is the edible berry of the plant ''Solanum lycopersicum'', commonly known as the tomato plant. The species originated in western South America, Mexico, and Central America. The Mexican Nahuatl word gave rise to the Spanish word ...
,
peppers Pepper or peppers may refer to: Food and spice * Piperaceae or the pepper family, a large family of flowering plant ** Black pepper * ''Capsicum'' or pepper, a genus of flowering plants in the nightshade family Solanaceae ** Bell pepper ** Chili ...
,
cucumbers Cucumber (''Cucumis sativus'') is a widely-cultivated creeping vine plant in the Cucurbitaceae family that bears usually cylindrical fruits, which are used as culinary vegetables.strawberries The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid species of the genus '' Fragaria'', collectively known as the strawberries, which are cultivated worldwide for their fruit. The fruit is widely ap ...
, lettuces, and
cannabis ''Cannabis'' () is a genus of flowering plants in the family Cannabaceae. The number of species within the genus is disputed. Three species may be recognized: ''Cannabis sativa'', '' C. indica'', and '' C. ruderalis''. Alternatively ...
, usually for commercial use, and ''
Arabidopsis thaliana ''Arabidopsis thaliana'', the thale cress, mouse-ear cress or arabidopsis, is a small flowering plant native to Eurasia and Africa. ''A. thaliana'' is considered a weed; it is found along the shoulders of roads and in disturbed land. A winter a ...
'', which serves as a
model organism A model organism (often shortened to model) is a non-human species that is extensively studied to understand particular biological phenomena, with the expectation that discoveries made in the model organism will provide insight into the workin ...
in
plant science Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
and
genetics Genetics is the study of genes, genetic variation, and heredity in organisms.Hartl D, Jones E (2005) It is an important branch in biology because heredity is vital to organisms' evolution. Gregor Mendel, a Moravian Augustinian friar wor ...
. Hydroponics offers many advantages, notably a decrease in water usage in
agriculture Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people to ...
. To grow of tomatoes using
intensive farming Intensive agriculture, also known as intensive farming (as opposed to extensive farming), conventional, or industrial agriculture, is a type of agriculture, both of crop plants and of animals, with higher levels of input and output per unit of ag ...
methods requires of water; using hydroponics, ; and only using
aeroponics Aeroponics is the process of growing plants in the air or mist environment without soil or an aggregate medium. The word "aeroponic" is derived from the Greek meanings of ''aer'' ("air") and ''ponos'' ("labour"). It is a subset of hydroponic ...
. Hydroponic cultures lead to highest
biomass Biomass is plant-based material used as a fuel for heat or electricity production. It can be in the form of wood, wood residues, energy crops, agricultural residues, and waste from industry, farms, and households. Some people use the terms bi ...
and
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
production compared to other growth substrates, of plants cultivated in the same environmental conditions and supplied with equal amounts of nutrients. Since hydroponics takes much less water and nutrients to grow produce and
climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
threatens
agricultural yields In agriculture, the yield is a measurement of the amount of a crop grown, or product such as wool, meat or milk produced, per unit area of land. The seed ratio is another way of calculating yields. Innovations, such as the use of fertilizer, the c ...
, it could be possible in the future for people in harsh environments with little accessible water to grow their own plant-based
food Food is any substance consumed by an organism for nutritional support. Food is usually of plant, animal, or fungal origin, and contains essential nutrients, such as carbohydrates, fats, proteins, vitamins, or minerals. The substance is inge ...
. Hydroponics is not only used on
earth Earth is the third planet from the Sun and the only astronomical object known to harbor life. While large volumes of water can be found throughout the Solar System, only Earth sustains liquid surface water. About 71% of Earth's surfa ...
, but has also proven itself in plant production experiments in
space Space is the boundless three-dimensional extent in which objects and events have relative position and direction. In classical physics, physical space is often conceived in three linear dimensions, although modern physicists usually consider ...
.


History

The earliest published work on growing terrestrial plants without soil was the 1627 book ''Sylva Sylvarum'' or 'A Natural History' by
Francis Bacon Francis Bacon, 1st Viscount St Alban (; 22 January 1561 – 9 April 1626), also known as Lord Verulam, was an English philosopher and statesman who served as Attorney General and Lord Chancellor of England. Bacon led the advancement of both ...
, printed a year after his death. As a result of his work, water culture became a popular research technique. In 1699,
John Woodward John Woodward or ''variant'', may refer to: Sports * John Woodward (English footballer) (born 1947), former footballer * John Woodward (Scottish footballer) (born 1949), former footballer * Johnny Woodward (1924–2002), English footballer * Jo ...
published his water culture experiments with
spearmint Spearmint, also known as garden mint, common mint, lamb mint and mackerel mint, is a species of mint, ''Mentha spicata'' (, native to Europe and southern temperate Asia, extending from Ireland in the west to southern China in the east. It is nat ...
. He found that plants in less-pure water sources grew better than plants in distilled water. By 1842, a list of nine elements believed to be essential for plant growth had been compiled, and the discoveries of German botanists
Julius von Sachs Julius von Sachs (; 2 October 1832 – 29 May 1897) was a German botanist from Breslau, Prussian Silesia. He is considered the founder of experimental plant physiology and co-founder of modern water culture. Julius von Sachs and Wilhelm Kno ...
and
Wilhelm Knop Wilhelm may refer to: People and fictional characters * William Charles John Pitcher, costume designer known professionally as "Wilhelm" * Wilhelm (name), a list of people and fictional characters with the given name or surname Other uses * Moun ...
, in the years 1859–1875, resulted in a development of the technique of soilless cultivation. To quote von Sachs directly: "In the year 1860, I published the results of experiments which demonstrated that land plants are capable of absorbing their nutritive matters out of watery solutions, without the aid of soil, and that it is possible in this way not only to maintain plants alive and growing for a long time, as had long been known, but also to bring about a vigorous increase of their organic substance, and even the production of seed capable of germination." Growth of terrestrial plants without soil in mineral nutrient solutions was later called "solution culture" in reference to "soil culture". It quickly became a standard research and teaching technique in the 19th and 20th centuries and is still widely used in
plant nutrition Plant nutrition is the study of the chemical elements and compounds necessary for plant growth and reproduction, plant metabolism and their external supply. In its absence the plant is unable to complete a normal life cycle, or that the element i ...
science. Around the 1930s plant nutritionists investigated diseases of certain plants, and thereby, observed symptoms related to existing soil conditions such as
salinity Salinity () is the saltiness or amount of salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensionless and equal ...
. In this context, water culture experiments were undertaken with the hope of delivering similar symptoms under controlled laboratory conditions. This approach forced by
Dennis Robert Hoagland Dennis Robert Hoagland (April 2, 1884 – September 5, 1949) was an American chemist and plant scientist working in the fields of plant nutrition, agricultural chemistry, and physiology. He was Professor of Plant Nutrition at the University o ...
led to innovative model systems (e.g.,
green algae The green algae (singular: green alga) are a group consisting of the Prasinodermophyta and its unnamed sister which contains the Chlorophyta and Charophyta/Streptophyta. The land plants (Embryophytes) have emerged deep in the Charophyte alga as ...
Nitella ''Nitella'' is a genus of charophyte green algae in the family Characeae. Species The species in the genus include: *'' Nitella abyssinica'' A. Braun * '' Nitella acuminata'' A. Braun ex Wallman *'' Nitella aemula'' A. Braun * '' Nitella an ...
) and standardized nutrient recipes playing an increasingly important role in modern plant physiology. In 1929, William Frederick Gericke of the University of California at Berkeley began publicly promoting that the principles of solution culture be used for agricultural crop production. He first termed this cultivation method "aquiculture" created in analogy to "agriculture" but later found that the cognate term
aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. lot ...
was already applied to culture of aquatic organisms. Gericke created a sensation by growing tomato vines high in his back yard in mineral nutrient solutions rather than soil. He then introduced the term ''Hydroponics'', water culture, in 1937, proposed to him by W. A. Setchell, a phycologist with an extensive education in the classics. Hydroponics is derived from
neologism A neologism Greek νέο- ''néo''(="new") and λόγος /''lógos'' meaning "speech, utterance"] is a relatively recent or isolated term, word, or phrase that may be in the process of entering common use, but that has not been fully accepted int ...
υδρωπονικά (derived from Greek ύδωρ=water and πονέω=cultivate), constructed in analogy to γεωπονικά (derived from Greek γαία=earth and πονέω=cultivate),
geoponica The ''Geoponica'' or ''Geoponika'' ( el, Γεωπονικά) is a twenty-book collection of agricultural lore, compiled during the 10th century in Constantinople for the Byzantine emperor Constantine VII Porphyrogenitus. The Greek word ''Geoponic ...
, that which concerns agriculture, replacing, γεω-, earth, with ὑδρο-, water. Despite initial successes, however, Gericke realized that the time was not yet ripe for the general technical application and commercial use of hydroponics for producing crops. He also wanted to make sure all aspects of hydroponic cultivation were researched and tested before making any of the specifics available to the public. Reports of Gericke's work and his claims that hydroponics would revolutionize plant agriculture prompted a huge number of requests for further information. Gericke had been denied use of the university's
greenhouse A greenhouse (also called a glasshouse, or, if with sufficient heating, a hothouse) is a structure with walls and roof made chiefly of Transparent ceramics, transparent material, such as glass, in which plants requiring regulated climatic condit ...
s for his experiments due to the administration's skepticism, and when the university tried to compel him to release his preliminary nutrient recipes developed at home, he requested greenhouse space and time to improve them using appropriate research facilities. While he was eventually provided greenhouse space, the university assigned Hoagland and Arnon to re-evaluate Gericke's claims and show his formula held no benefit over soil grown plant yields, a view held by Hoagland. Because of these irreconcilable conflicts, Gericke left his academic position in 1937 in a climate that was politically unfavorable and continued his research independently in his greenhouse. In 1940, Gericke, whose work is considered to be the basis for all forms of hydroponic growing, published the book, ''Complete Guide to Soilless Gardening''. Therein, for the first time, he published his basic formula involving the macro- and micronutrient salts for hydroponically-grown plants. As a result of research of Gericke's claims by order of the Director of the ''California Agricultural Experiment Station'' of the
University of California The University of California (UC) is a public land-grant research university system in the U.S. state of California. The system is composed of the campuses at Berkeley, Davis, Irvine, Los Angeles, Merced, Riverside, San Diego, San Francisco, ...
, Claude Hutchison, Dennis Hoagland and Daniel Arnon wrote a classic 1938 agricultural bulletin, ''The Water Culture Method for Growing Plants Without Soil'', one of the most important works on solution culture ever, which made the claim that hydroponic
crop yield In agriculture, the yield is a measurement of the amount of a crop grown, or product such as wool, meat or milk produced, per unit area of land. The seed ratio is another way of calculating yields. Innovations, such as the use of fertilizer, the c ...
s were no better than crop yields obtained with good-quality soils. Ultimately, crop yields would be limited by factors other than mineral nutrients, especially light and aeration of the culture medium. However, in the introduction to his landmark book on soilless cultivation, published two years later, Gericke pointed out that the results published by Hoagland and Arnon in comparing the yields of experimental plants in sand, soil and solution cultures, were based on several systemic errors ("...these experimenters have made the mistake of limiting the productive capacity of hydroponics to that of soil. Comparison can be only by growing as great a number of plants in each case as the fertility of the culture medium can support"). For example, the Hoagland and Arnon study did not adequately appreciate that hydroponics has other key benefits compared to soil culture including the fact that the roots of the plant have constant access to
oxygen Oxygen is the chemical element with the symbol O and atomic number 8. It is a member of the chalcogen group in the periodic table, a highly reactive nonmetal, and an oxidizing agent that readily forms oxides with most elements as wel ...
and that the plants have access to as much or as little water and nutrients as they need. This is important as one of the most common errors when cultivating plants is over- and underwatering; and hydroponics prevents this from occurring as large amounts of water, which may drown root systems in soil, can be made available to the plant in hydroponics, and any water not used, drained away, recirculated, or actively aerated, eliminating
anoxic The term anoxia means a total depletion in the level of oxygen, an extreme form of hypoxia or "low oxygen". The terms anoxia and hypoxia are used in various contexts: * Anoxic waters, sea water, fresh water or groundwater that are depleted of diss ...
conditions in the root area. In soil, a grower needs to be very experienced to know exactly with how much water to feed the plant. Too much and the plant will be unable to access oxygen because
air The atmosphere of Earth is the layer of gases, known collectively as air, retained by Earth's gravity that surrounds the planet and forms its planetary atmosphere. The atmosphere of Earth protects life on Earth by creating pressure allowing f ...
in the soil pores is displaced, which can lead to root rot; too little and the plant will undergo
water stress Water scarcity (closely related to water stress or water crisis) is the lack of fresh water resources to meet the standard water demand. There are two types of water scarcity: physical or economic water scarcity. Physical water scarcity is where ...
or lose the ability to absorb nutrients, which are typically moved into the roots while dissolved, leading to nutrient deficiency symptoms such as
chlorosis In botany, chlorosis is a condition in which leaves produce insufficient chlorophyll. As chlorophyll is responsible for the green color of leaves, chlorotic leaves are pale, yellow, or yellow-white. The affected plant has little or no ability to ...
. Eventually, Gericke's advanced ideas led to the implementation of hydroponics into commercial agriculture while Hoagland's views and helpful support by the University prompted Hoagland and his associates to develop several new formulas for mineral nutrient solutions, universally known as
Hoagland solution The Hoagland solution is a hydroponic nutrient solution that was newly developed by Hoagland and Snyder in 1933, modified by Hoagland and Arnon in 1938, and revised by Arnon in 1950. It is one of the most popular artificial solution compositions ...
. One of the earliest successes of hydroponics occurred on
Wake Island Wake Island ( mh, Ānen Kio, translation=island of the kio flower; also known as Wake Atoll) is a coral atoll in the western Pacific Ocean in the northeastern area of the Micronesia subregion, east of Guam, west of Honolulu, southeast of To ...
, a rocky atoll in the Pacific Ocean used as a refueling stop for
Pan American Airlines Pan American World Airways, originally founded as Pan American Airways and commonly known as Pan Am, was an American airline that was the principal and largest international air carrier and unofficial overseas flag carrier of the United States ...
. Hydroponics was used there in the 1930s to grow vegetables for the passengers. Hydroponics was a necessity on Wake Island because there was no soil, and it was prohibitively expensive to airlift in fresh vegetables. From 1943 to 1946, Daniel I. Arnon served as a major in the
United States Army The United States Army (USA) is the land service branch of the United States Armed Forces. It is one of the eight U.S. uniformed services, and is designated as the Army of the United States in the U.S. Constitution.Article II, section 2, cla ...
and used his prior expertise with plant nutrition to feed troops stationed on barren
Ponape Island Pohnpei "upon (''pohn'') a stone altar (''pei'')" (formerly known as Ponape or Ascension, Proto-Chuukic-Pohnpeic: ''*Fawo ni pei)'' is an island of the Senyavin Islands which are part of the larger Caroline Islands group. It belongs to Pohnpei ...
in the western
Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
by growing crops in gravel and nutrient-rich water because there was no
arable land Arable land (from the la, arabilis, "able to be ploughed") is any land capable of being ploughed and used to grow crops.''Oxford English Dictionary'', "arable, ''adj''. and ''n.''" Oxford University Press (Oxford), 2013. Alternatively, for the ...
available.Sullivan, Walter
"Daniel Arnon, 84, Researcher And Expert on Photosynthesis"
''
The New York Times ''The New York Times'' (''the Times'', ''NYT'', or the Gray Lady) is a daily newspaper based in New York City with a worldwide readership reported in 2020 to comprise a declining 840,000 paid print subscribers, and a growing 6 million paid ...
'', December 23, 1994. Accessed April 7, 2020
In the 1960s, Allen Cooper of England developed the
nutrient film technique Nutrient film technique (NFT) is a hydroponic technique where in a very shallow stream of water containing all the dissolved nutrients required for plant growth is re-circulated past the bare roots of plants in a watertight gully, also known as ...
. The Land Pavilion at Walt Disney World's EPCOT Center opened in 1982 and prominently features a variety of hydroponic techniques. In recent decades,
NASA The National Aeronautics and Space Administration (NASA ) is an independent agency of the US federal government responsible for the civil space program, aeronautics research, and space research. NASA was established in 1958, succeeding t ...
has done extensive hydroponic research for its
Controlled Ecological Life Support System Controlled (or closed) ecological life-support systems (acronym CELSS) are a self-supporting life support system for space stations and colonies typically through controlled closed ecological systems, such as the BioHome, BIOS-3, Biosphere 2, Mars ...
(CELSS). Hydroponics research mimicking a Martian environment uses LED lighting to grow in a different color spectrum with much less heat. Ray Wheeler, a plant physiologist at Kennedy Space Center's Space Life Science Lab, believes that hydroponics will create advances within space travel, as a
bioregenerative life support system Bioregenerative life support systems (BLSS) are artificial ecosystems consisting of many complex symbiotic relationships among higher plants, animals, and microorganisms. As the most advanced life support technology, BLSS can provide a habitati ...
. As of 2017, Canada had hundreds of acres of large-scale commercial hydroponic greenhouses, producing tomatoes, peppers and cucumbers. Due to technological advancements within the industry and numerous economic factors, the global hydroponics market is forecast to grow from US$226.45 million in 2016 to US$724.87 million by 2023.


Techniques

There are two main variations for each medium: sub-irrigation and top irrigation. For all techniques, most hydroponic reservoirs are now built of plastic, but other materials have been used, including concrete, glass, metal, vegetable solids, and wood. The containers should exclude light to prevent algae and fungal growth in the nutrient solution.


Static solution culture

In static solution culture, plants are grown in containers of nutrient solution, such as glass Mason jars (typically, in-home applications), Flowerpot, pots, buckets, tubs, or tanks. The solution is usually gently aerated but may be un-aerated. If un-aerated, the solution level is kept low enough that enough roots are above the solution so they get adequate oxygen. A hole is cut (or drilled) in the top of the reservoir for each plant; if it is a jar or tub, it may be its lid, but otherwise, cardboard, foil, paper, wood or metal may be put on top. A single reservoir can be dedicated to a single plant, or to various plants. Reservoir size can be increased as plant size increases. A home-made system can be constructed from food containers or glass canning jars with aeration provided by an aquarium pump, aquarium airline tubing, aquarium valves or even a biofilm of
green algae The green algae (singular: green alga) are a group consisting of the Prasinodermophyta and its unnamed sister which contains the Chlorophyta and Charophyta/Streptophyta. The land plants (Embryophytes) have emerged deep in the Charophyte alga as ...
on the glass, through photosynthesis. Clear containers can also be covered with aluminium foil, butcher paper, black plastic, or other material to eliminate the effects of negative phototropism. The nutrient solution is changed either on a schedule, such as once per week, or when the concentration drops below a certain level as determined with an EC meter, electrical conductivity meter. Whenever the solution is depleted below a certain level, either water or fresh nutrient solution is added. A Mariotte's bottle, or a float valve, can be used to automatically maintain the solution level. In raft solution culture, plants are placed in a sheet of buoyant plastic that is floated on the surface of the nutrient solution. That way, the solution level never drops below the roots.


Continuous-flow solution culture

In continuous-flow solution culture, the nutrient solution constantly flows past the roots. It is much easier to automate than the static solution culture because sampling and adjustments to the temperature, pH, and nutrient concentrations can be made in a large storage tank that has potential to serve thousands of plants. A popular variation is the
nutrient film technique Nutrient film technique (NFT) is a hydroponic technique where in a very shallow stream of water containing all the dissolved nutrients required for plant growth is re-circulated past the bare roots of plants in a watertight gully, also known as ...
or NFT, whereby a very shallow stream of water containing all the dissolved nutrients required for plant growth is recirculated in a thin layer past a bare root mat of plants in a watertight channel, with an upper surface exposed to air. As a consequence, an abundant supply of oxygen is provided to the roots of the plants. A properly designed NFT system is based on using the right channel slope, the right flow rate, and the right channel length. The main advantage of the NFT system over other forms of hydroponics is that the plant roots are exposed to adequate supplies of water, oxygen, and nutrients. In all other forms of production, there is a conflict between the supply of these requirements, since excessive or deficient amounts of one results in an imbalance of one or both of the others. NFT, because of its design, provides a system where all three requirements for healthy plant growth can be met at the same time, provided that the simple concept of NFT is always remembered and practised. The result of these advantages is that higher yields of high-quality produce are obtained over an extended period of cropping. A downside of NFT is that it has very little buffering against interruptions in the flow (e.g., power outages). But, overall, it is probably one of the more productive techniques. The same design characteristics apply to all conventional NFT systems. While slopes along channels of 1:100 have been recommended, in practice it is difficult to build a base for channels that is sufficiently true to enable nutrient films to flow without ponding in locally depressed areas. As a consequence, it is recommended that slopes of 1:30 to 1:40 are used. This allows for minor irregularities in the surface, but, even with these slopes, ponding and waterlogging (agriculture), water logging may occur. The slope may be provided by the floor, benches or racks may hold the channels and provide the required slope. Both methods are used and depend on local requirements, often determined by the site and crop requirements. As a general guide, flow rates for each gully should be one liter per minute. At planting, rates may be half this and the upper limit of 2 L/min appears about the maximum. Flow rates beyond these extremes are often associated with nutritional problems. Depressed growth rates of many crops have been observed when channels exceed 12 meters in length. On rapidly growing crops, tests have indicated that, while oxygen levels remain adequate, nitrogen may be depleted over the length of the gully. As a consequence, channel length should not exceed 10–15 meters. In situations where this is not possible, the reductions in growth can be eliminated by placing another nutrient feed halfway along the gully and halving the flow rates through each outlet.


Aeroponics

Aeroponics is a system wherein roots are continuously or discontinuously kept in an environment saturated with fine drops (a mist or aerosol) of nutrient solution. The method requires no substrate and entails growing plants with their roots suspended in a deep air or growth chamber with the roots periodically wetted with a fine mist of Atomizer nozzle, atomized nutrients. Excellent aeration is the main advantage of aeroponics. Aeroponic techniques have proven to be commercially successful for propagation, seed germination, seed potato production, tomato production, leaf crops, and micro-greens. Since inventor Richard Stoner commercialized aeroponic technology in 1983, aeroponics has been implemented as an alternative to water intensive hydroponic systems worldwide. The limitation of hydroponics is the fact that of water can only hold of air, no matter whether aerators are utilized or not. Another distinct advantage of aeroponics over hydroponics is that any species of plants can be grown in a true aeroponic system because the microenvironment of an aeroponic can be finely controlled. The limitation of hydroponics is that certain species of plants can only survive for so long in water before they become waterlogging (agriculture), waterlogged. The advantage of aeroponics is that suspended aeroponic plants receive 100% of the available oxygen and carbon dioxide to the roots zone, stems, and leaves, thus accelerating biomass growth and reducing rooting times. NASA research has shown that aeroponically grown plants have an 80% increase in dry weight biomass (essential minerals) compared to hydroponically grown plants. Aeroponics used 65% less water than hydroponics. NASA also concluded that aeroponically grown plants require ¼ the nutrient input compared to hydroponics. Unlike hydroponically grown plants, aeroponically grown plants will not suffer transplant shock when transplanted to soil, and offers growers the ability to reduce the spread of disease and pathogens. Aeroponics is also widely used in laboratory studies of plant physiology and plant pathology. Aeroponic techniques have been given special attention from
NASA The National Aeronautics and Space Administration (NASA ) is an independent agency of the US federal government responsible for the civil space program, aeronautics research, and space research. NASA was established in 1958, succeeding t ...
since a mist is easier to handle than a liquid in a zero-gravity environment.


Fogponics

Fogponics is a derivation of aeroponics wherein the nutrient solution is aerosolized by a ultrasonic humidifier, diaphragm vibrating at ultrasonic frequencies. Solution droplets produced by this method tend to be 5–10 µm in diameter, smaller than those produced by forcing a nutrient solution through pressurized nozzles, as in aeroponics. The smaller size of the droplets allows them to diffuse through the air more easily, and deliver nutrients to the roots without limiting their access to oxygen.


Passive sub-irrigation

Passive sub-irrigation, also known as passive hydroponics, semi-hydroponics, or ''hydroculture'', is a method wherein plants are grown in an inert porous medium that moves water and fertilizer to the roots by capillary action from a separate reservoir as necessary, reducing labor and providing a constant supply of water to the roots. In the simplest method, the pot sits in a shallow solution of fertilizer and water or on a capillary mat saturated with nutrient solution. The various hydroponic media available, such as ex-clay, expanded clay and Coir, coconut husk, contain more air space than more traditional potting mixes, delivering increased oxygen to the roots, which is important in epiphyte, epiphytic plants such as Orchidaceae, orchids and Bromeliaceae, bromeliads, whose roots are exposed to the air in nature. Additional advantages of passive hydroponics are the reduction of root rot and the additional ambient humidity provided through evaporations. Hydroculture compared to traditional farming in terms of crops yield per area in a controlled environment was roughly 10 times more efficient than traditional farming, uses 13 times less water in one crop cycle than traditional farming, but on average uses 100 times more kilojoules per kilogram of energy than traditional farming.


Ebb and flow (flood and drain) sub-irrigation

In its simplest form, there is a tray above a reservoir of nutrient solution. Either the tray is filled with growing medium (clay granules being the most common) and then plant directly or place the pot over medium, stand in the tray. At regular intervals, a simple timer causes a pump to fill the upper tray with nutrient solution, after which the solution drains back down into the reservoir. This keeps the medium regularly flushed with nutrients and air. Once the upper tray fills past the drain stop, it begins recirculating the water until the timer turns the pump off, and the water in the upper tray drains back into the reservoirs.


Run-to-waste

In a run-to-waste system, nutrient and water solution is periodically applied to the medium surface. The method was invented in Bengal in 1946; for this reason it is sometimes referred to as "The Bengal System". This method can be set up in various configurations. In its simplest form, a nutrient-and-water solution is manually applied one or more times per day to a container of inert growing media, such as rockwool, perlite, vermiculite, coco fibre, or sand. In a slightly more complex system, it is automated with a delivery pump, a timer and irrigation tubing to deliver nutrient solution with a delivery frequency that is governed by the key parameters of plant size, plant growing stage, climate, substrate, and substrate conductivity, pH, and water content. In a commercial setting, watering frequency is multi-factorial and governed by computers or Programmable logic controller, PLCs. Commercial hydroponics production of large plants like tomatoes, cucumber, and peppers uses one form or another of run-to-waste hydroponics. In environmentally responsible uses, the nutrient-rich waste is collected and processed through an on-site filtration system to be used many times, making the system very productive.


Deep water culture

The hydroponic method of plant production by means of suspending the plant roots in a solution of nutrient-rich, oxygenated water. Traditional methods favor the use of plastic buckets and large containers with the plant contained in a net pot suspended from the centre of the lid and the roots suspended in the nutrient solution. The solution is oxygen saturated by an air pump combined with airstone, porous stones. With this method, the plants grow much faster because of the high amount of oxygen that the roots receive. The Kratky Method is similar to deep water culture, but uses a non-circulating water reservoir.


Top-fed deep water culture

''Top-fed'' deep water culture is a technique involving delivering highly oxygenated nutrient solution direct to the root zone of plants. While deep water culture involves the plant roots hanging down into a reservoir of nutrient solution, in top-fed deep water culture the solution is pumped from the reservoir up to the roots (top feeding). The water is released over the plant's roots and then runs back into the reservoir below in a constantly recirculating system. As with deep water culture, there is an airstone in the reservoir that pumps air into the water via a hose from outside the reservoir. The airstone helps add oxygen to the water. Both the airstone and the water pump run 24 hours a day. The biggest advantage of top-fed deep water culture over standard deep water culture is increased growth during the first few weeks. With deep water culture, there is a time when the roots have not reached the water yet. With top-fed deep water culture, the roots get easy access to water from the beginning and will grow to the reservoir below much more quickly than with a deep water culture system. Once the roots have reached the reservoir below, there is not a huge advantage with top-fed deep water culture over standard deep water culture. However, due to the quicker growth in the beginning, grow time can be reduced by a few weeks.


Rotary

A rotary hydroponic garden is a style of commercial hydroponics created within a circular frame which rotates continuously during the entire growth cycle of whatever plant is being grown. While system specifics vary, systems typically rotate once per hour, giving a plant 24 full turns within the circle each 24-hour period. Within the center of each rotary hydroponic garden can be a high intensity grow light, designed to simulate sunlight, often with the assistance of a mechanized timer. Each day, as the plants rotate, they are periodically watered with a hydroponic growth solution to provide all nutrients necessary for robust growth. Due to the plants continuous fight against gravity, plants typically mature much more quickly than when grown in soil or other traditional hydroponic growing systems. Because rotary hydroponic systems have a small size, they allow for more plant material to be grown per area of floor space than other traditional hydroponic systems. Rotary hydroponic systems should be avoided in most circumstances, mainly because of their experimental nature and their high costs for finding, buying, operating, and maintaining them.


Substrates (growing support materials)

One of the most obvious decisions hydroponic farmers have to make is which medium they should use. Different media are appropriate for different growing techniques.


Rock wool

Rock wool (mineral wool) is the most widely used medium in hydroponics. Rock wool is an inert substrate suitable for both run-to-waste and recirculating systems. Rock wool is made from molten rock, basalt or 'slag' that is spun into bundles of single filament fibres, and bonded into a medium capable of capillary action, and is, in effect, protected from most common microbiological degradation. Rock wool is typically used only for the seedling stage, or with newly cut clones, but can remain with the plant base for its lifetime. Rock wool has many advantages and some disadvantages. The latter being the possible skin irritancy (mechanical) whilst handling (1:1000). Flushing with cold water usually brings relief. Advantages include its proven efficiency and effectiveness as a commercial hydroponic substrate. Most of the rock wool sold to date is a non-hazardous, non-carcinogenic material, falling under Note Q of the European Union Classification Packaging and Labeling Regulation (CLP). Mineral wool products can be engineered to hold large quantities of water and air that aid root growth and nutrient uptake in hydroponics; their fibrous nature also provides a good mechanical structure to hold the plant stable. The naturally high pH of mineral wool makes them initially unsuitable to plant growth and requires "conditioning" to produce a wool with an appropriate, stable pH.


Expanded clay aggregate

Baked clay pellets are suitable for hydroponic systems in which all nutrients are carefully controlled in water solution. The clay pellets are inert, pH-neutral, and do not contain any nutrient value. The clay is formed into round pellets and fired in rotary kilns at . This causes the clay to expand, like popcorn, and become porous. It is light in weight, and does not compact over time. The shape of an individual pellet can be irregular or uniform depending on brand and manufacturing process. The manufacturers consider expanded clay to be an ecologically sustainable and re-usable growing medium because of its ability to be cleaned and sterilized, typically by washing in solutions of white vinegar, chlorine bleach, or hydrogen peroxide (), and rinsing completely. Another view is that clay pebbles are best not re-used even when they are cleaned, due to root growth that may enter the medium. Breaking open a clay pebble after a crop has been shown to reveal this growth.


Growstones

Growstones, made from glass waste, have both more air and water retention space than perlite and peat. This aggregate holds more water than parboiled rice hulls. Growstones by volume consist of 0.5 to 5% calcium carbonate – for a standard 5.1 kg bag of Growstones that corresponds to 25.8 to 258 grams of calcium carbonate. The remainder is soda-lime glass.


Coconut Coir

Regardless of hydroponic demand, coconut coir is a natural byproduct derived from coconut processes. The outer husk of a coconut consists of fibers which are commonly used to make a myriad of items ranging from floor mats to brushes. After the long fibers are used for those applications, the dust and short fibers are merged to create coir. Coconuts absorb high levels of nutrients throughout their life cycle, so the coir must undergo a maturation process before it becomes a viable growth medium. This process removes salt, tannins and phenolic compounds through substantial water washing. Contaminated water is a byproduct of this process, as three hundred to six hundred liters of water per one cubic meter of coir is needed. Additionally, this maturation can take up to six months and one study concluded the working conditions during the maturation process are dangerous and would be illegal in North America and Europe. Despite requiring attention, posing health risks and environmental impacts, coconut coir has impressive material properties. When exposed to water, the brown, dry, chunky and fibrous material expands nearly three or four times its original size. This characteristic combined with coconut coir's water retention capacity and resistance to pests and diseases make it an effective growth medium. Used as an alternative to rock wool, coconut coir, also known as coir peat, offers optimized growing conditions.


Rice husks

Rice husks#Fertilizer and substrate, Parboiled rice husks (PBH) are an agricultural byproduct that would otherwise have little use. They decay over time, and allow drainage, and even retain less water than growstones. A study showed that rice husks did not affect the effects of plant hormones, plant growth regulators.


Perlite

Perlite is a volcanic rock that has been superheated into very lightweight expanded glass pebbles. It is used loose or in plastic sleeves immersed in the water. It is also used in potting soil mixes to decrease soil density. It does contain a high amount of fluorine which could be harmful to some plants. Perlite has similar properties and uses to vermiculite but, in general, holds more air and less water and is buoyant.


Vermiculite

Like perlite, vermiculite is a mineral that has been superheated until it has expanded into light pebbles. Vermiculite holds more water than perlite and has a natural "wicking" property that can draw water and nutrients in a passive hydroponic system. If too much water and not enough air surrounds the plants roots, it is possible to gradually lower the medium's water-retention capability by mixing in increasing quantities of perlite.


Pumice

Like perlite, pumice is a lightweight, mined volcanic rock that finds application in hydroponics.


Sand

Sand is cheap and easily available. However, it is heavy, does not hold water very well, and it must be sterilized between uses.


Gravel

The same type that is used in aquariums, though any small gravel can be used, provided it is washed first. Indeed, plants growing in a typical traditional gravel filter bed, with water circulated using electric powerhead pumps, are in effect being grown using gravel hydroponics, also termed "nutriculture". Gravel is inexpensive, easy to keep clean, drains well and will not become waterlogged. However, it is also heavy, and, if the system does not provide continuous water, the plant roots may dry out.


Wood fiber

Wood fibre, produced from steam friction of wood, is an efficient organic substrate for hydroponics. It has the advantage that it keeps its structure for a very long time. Wood wool (i.e. wood slivers) have been used since the earliest days of the hydroponics research. However, more recent research suggests that wood fibre may have detrimental effects on "plant growth regulators".


Sheep wool

Wool from shearing sheep is a little-used yet promising renewable growing medium. In a study comparing wool with peat slabs, coconut fibre slabs, perlite and rockwool slabs to grow cucumber plants, sheep wool had a greater air capacity of 70%, which decreased with use to a comparable 43%, and water capacity that increased from 23% to 44% with use. Using sheep wool resulted in the greatest yield out of the tested substrates, while application of a biostimulator consisting of humic acid, lactic acid and Bacillus subtilis improved yields in all substrates.


Brick shards

Brick shards have similar properties to gravel. They have the added disadvantages of possibly altering the pH and requiring extra cleaning before reuse.


Polystyrene packing peanuts

Polystyrene packing peanuts are inexpensive, readily available, and have excellent drainage. However, they can be too lightweight for some uses. They are used mainly in closed-tube systems. Note that non-biodegradable polystyrene peanuts must be used; biodegradable packing peanuts will decompose into a sludge. Plants may absorb styrene and pass it to their consumers; this is a possible health risk.


Nutrient solutions


Inorganic hydroponic solutions

The formulation of hydroponic solutions is an application of
plant nutrition Plant nutrition is the study of the chemical elements and compounds necessary for plant growth and reproduction, plant metabolism and their external supply. In its absence the plant is unable to complete a normal life cycle, or that the element i ...
, with nutrient deficiency symptoms mirroring those found in traditional Agricultural soil science, soil based agriculture. However, the underlying chemistry of hydroponic solutions can differ from Soil nutrient, soil chemistry in many significant ways. Important differences include: * Unlike soil, hydroponic nutrient solutions do not have cation-exchange capacity (CEC) from clay particles or organic matter. The absence of CEC and soil pores means the pH, oxygen saturation, and nutrient concentrations can change much more rapidly in hydroponic setups than is possible in soil. * Selective absorption of nutrients by plants often imbalances the amount of counterions in solution. This imbalance can rapidly affect solution pH and the ability of plants to absorb nutrients of similar ionic charge (see article membrane potential). For instance, nitrate anions are often consumed rapidly by plants to form
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
s, leaving an excess of cations in solution. This cation imbalance can lead to deficiency symptoms in other cation based nutrients (e.g. Magnesium, Mg2+) even when an ideal quantity of those nutrients are dissolved in the solution. * Depending on the pH or on the presence of Water pollution, water contaminants, nutrients such as iron can Precipitation (chemistry), precipitate from the solution and become unavailable to plants. Routine adjustments to pH, Buffer solution, buffering the solution, or the use of chelating agents is often necessary. * Unlike soil types, which can vary greatly in their Chemical composition, composition, hydroponic solutions are often standardized and require routine maintenance for plant cultivation. Under controlled conditions hydroponic solutions are periodically pH adjusted to near neutral (pH 6.0) and are aerated with oxygen. Also, water levels must be refilled to account for transpiration losses and nutrient solutions require re-fortification to correct the nutrient imbalances that occur as plants grow and deplete nutrient reserves. Sometimes the regular measurement of nitrate ions is used as a key parameter to estimate the remaining proportions and concentrations of other essential nutrient ions to restore a balanced solution. * Well-known examples of standardized, balanced nutrient solutions are the
Hoagland solution The Hoagland solution is a hydroponic nutrient solution that was newly developed by Hoagland and Snyder in 1933, modified by Hoagland and Arnon in 1938, and revised by Arnon in 1950. It is one of the most popular artificial solution compositions ...
, the Long Ashton Research Station, Long Ashton nutrient solution, or the Wilhelm Knop, Knop solution. As in conventional agriculture, nutrients should be adjusted to satisfy Liebig's law of the minimum for each specific plant Variety (botany), variety. Nevertheless, generally acceptable concentrations for nutrient solutions exist, with minimum and maximum concentration ranges for most plants being somewhat similar. Most nutrient solutions are mixed to have concentrations between 1,000 and 2,500 parts per million, ppm. Acceptable concentrations for the individual nutrient ions, which comprise that total ppm figure, are summarized in the following table. For essential nutrients, concentrations below these ranges often lead to nutrient deficiencies while exceeding these ranges can lead to nutrient toxicity. Optimum nutrition concentrations for plant varieties are found empirically by experience or by plant tissue tests.


Organic hydroponic solutions

Organic fertilizers can be used to supplement or entirely replace the inorganic compounds used in conventional hydroponic solutions. However, using organic fertilizers introduces a number of challenges that are not easily resolved. Examples include: * organic fertilizers are highly variable in their nutritional compositions in terms of Mineral (nutrient), minerals and different chemical species. Even similar materials can differ significantly based on their source (e.g. the quality of
manure Manure is organic matter that is used as organic fertilizer in agriculture. Most manure consists of animal feces; other sources include compost and green manure. Manures contribute to the Soil fertility, fertility of soil by adding organic ma ...
varies based on an animal's diet). * organic fertilizers are often sourced from animal byproducts, making Transmission (medicine), disease transmission a serious concern for plants grown for human consumption or animal forage. * organic fertilizers are often particulate and can clog substrates or other growing equipment. Sieve, Sieving or mill (grinding), milling the organic materials to fine dusts is often necessary. * Biochemistry, biochemical degradation and conversion processes of organic materials can make mineral ingredients Bioavailability (soil), available to plants. * some organic materials (i.e. particularly manures and offal) can further Biodegradation, degrade to emit foul odors under Anaerobic digestion, anaerobic conditions. * many organic molecules (i.e. sugars) demand additional oxygen during aerobic degradation, which is essential for cellular respiration in the plant roots. * organic compounds (i.e. sugars, vitamins, a.o.) are not necessary for normal plant nutrition. Nevertheless, if precautions are taken, organic fertilizers can be used successfully in hydroponics.


Organically sourced macronutrients

Examples of suitable materials, with their average nutritional contents tabulated in terms of percent dried mass, are listed in the following table.


Organically sourced micronutrients

Micronutrients can be sourced from organic fertilizers as well. For example, composted conifer, pine bark is high in manganese and is sometimes used to fulfill that mineral requirement in hydroponic solutions. To satisfy requirements for National Organic Programs, pulverized, unrefined
mineral In geology and mineralogy, a mineral or mineral species is, broadly speaking, a solid chemical compound with a fairly well-defined chemical composition and a specific crystal structure that occurs naturally in pure form.John P. Rafferty, ed. ( ...
s (e.g. Gypsum, Calcite, and glauconite) can also be added to satisfy a plant's nutritional needs.


Additives

Compounds can be added in both organic and conventional hydroponic systems to improve nutrition acquisition and uptake by the plant''.'' Chelating agents and humic acid have been shown to increase nutrient uptake. Additionally, plant growth promoting rhizobacteria (PGPR), which are regularly utilized in field and greenhouse agriculture, have been shown to benefit hydroponic plant growth development and nutrient acquisition. Some PGPR are known to increase nitrogen fixation. While nitrogen is generally abundant in hydroponic systems with properly maintained fertilizer regimens, ''Azospirillum'' and ''Azotobacter'' genera can help maintain mobilized forms of nitrogen in systems with higher microbial growth in the rhizosphere. Traditional fertilizer methods often lead to high accumulated concentrations of nitrate within plant tissue at harvest. ''Rhodopseudo-monas palustris'' has been shown to increase nitrogen use efficiency, increase yield, and decrease nitrate concentration by 88% at harvest compared to traditional hydroponic fertilizer methods in leafy greens. Many ''Bacillus'' spp., ''Pseudomonas'' spp. and ''Streptomyces'' spp. convert forms of phosphorus in the soil that are unavailable to the plant into soluble anions by decreasing soil pH, releasing phosphorus bound in chelated form that is available in a wider pH range, and mineralizing organic phosphorus. Some studies have found that ''Bacillus'' inoculants allow hydroponic leaf lettuce to overcome high salt stress that would otherwise reduce growth. This can be especially beneficial in regions with high electrical conductivity or salt content in their water source. This could potentially avoid costly reverse osmosis filtration systems while maintaining high crop yield.


Tools


Common equipment

Managing nutrient concentrations, oxygen saturation, and pH values within acceptable ranges is essential for successful hydroponic
horticulture Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...
. Common tools used to manage hydroponic solutions include: * Electrical conductivity meters, a tool which estimates nutrient ppm by measuring how well a solution transmits an electric current. * pH meter, a tool that uses an electric current to determine the concentration of hydrogen ions in solution. * Clark electrode, Oxygen electrode, an electrochemical sensor for determining the oxygen concentration in solution. * Litmus paper, disposable pH indicator strips that determine hydrogen ion concentrations by color changing chemical reaction. * Graduated cylinders or measuring spoons to measure out premixed, commercial hydroponic solutions.


Equipment

Chemical equipment can also be used to perform accurate chemical analysis, chemical analyses of nutrient solutions. Examples include: * Weighing scale, Balances for accurately measuring materials. * Laboratory glassware, such as burettes and pipettes, for performing titrations. * Colorimeter (chemistry), Colorimeters for solution tests which apply the Beer–Lambert law. * Spectrophotometry, Spectrophotometer to measure the concentrations of the key parameter nitrate and other nutrients, such as phosphate, sulfate or iron. Using chemical equipment for hydroponic solutions can be beneficial to growers of any background because nutrient solutions are often reusable. Because nutrient solutions are virtually never completely depleted, and should never be due to the unacceptably low osmotic pressure that would result, re-fortification of old solutions with new nutrients can save growers money and can control point source pollution, a common source for the eutrophication of nearby lakes and streams.


Software

Although pre-mixed concentrated nutrient solutions are generally purchased from commercial nutrient manufacturers by hydroponic hobbyists and small commercial growers, several tools exist to help anyone prepare their own solutions without extensive knowledge about chemistry. The free and open source tools HydroBuddy and HydroCal have been created by professional chemists to help any hydroponics grower prepare their own nutrient solutions. The first program is available for Windows, Mac and Linux while the second one can be used through a simple JavaScript interface. Both programs allow for basic nutrient solution preparation although HydroBuddy provides added functionality to use and save custom substances, save formulations and predict electrical conductivity values.


Mixing solutions

Often mixing hydroponic solutions using individual salts is impractical for hobbyists or small-scale commercial growers because commercial products are available at reasonable prices. However, even when buying commercial products, multi-component fertilizers are popular. Often these products are bought as three part formulas which emphasize certain nutritional roles. For example, solutions for vegetative growth (i.e. high in nitrogen), flowering (i.e. high in potassium and phosphorus), and micronutrient solutions (i.e. with trace minerals) are popular. The timing and application of these multi-part fertilizers should coincide with a plant's growth stage. For example, at the end of an annual plant's Biological life cycle, life cycle, a plant should be restricted from high nitrogen fertilizers. In most plants, nitrogen restriction inhibits vegetative growth and helps Flowering transition, induce flowering.


Additional improvements


Growrooms

With pest problems reduced and nutrients constantly fed to the roots, productivity in hydroponics is high; however, growers can further increase yield by manipulating a plant's environment by constructing sophisticated growrooms.


CO2 enrichment

To increase yield further, some sealed greenhouses inject carbon dioxide, CO2 into their environment to help improve growth and plant fertility.


See also

* Aeroponics * Anthroponics * Aquaponics * Fogponics * Folkewall * Grow box * Growroom * Organoponics * Passive hydroponics * Plant factory * Plant nutrition * Plant pathology * Root rot * Vertical farming * Xeriscaping


References

{{Authority control Hydroponics, Hydroculture Aeroponics de:Hydrokultur