Harttiini
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Loricariinae is a subfamily of the family
Loricariidae The Loricariidae is the largest family of catfish (order Siluriformes), with 92 genera and just over 680 species. Loricariids originate from freshwater habitats of Costa Rica, Panama, and tropical and subtropical South America. These fish are not ...
of
catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive ...
(order Siluriformes). This subfamily is divided into two
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confli ...
s and about 30 genera. They are mainly native to freshwater habitats in South America, but there are also several species (in genera '' Crossoloricaria'', '' Dasyloricaria'', '' Fonchiiichthys'', '' Rineloricaria'', '' Spatuloricaria'', ''
Sturisoma ''Sturisoma'' is a genus of armored catfishes native to Central and South America. Taxonomy ''Sturisoma'' has been shown to be sister to ''Farlowella''. Species There are currently 12 recognized species in this genus: * '' Sturisoma barbatum' ...
'' and '' Sturisomatichthys'') in
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Co ...
and a single (''Fonchiiichthys'') in Costa Rica.Angulo; Garita-Alvarado; Bussing; and López (2013).
Annotated checklist of the freshwater fishes of continental and insular Costa Rica: additions and nomenclatural revisions.
'' Check List 9(5): 987–1019.


Taxonomy

Loricariinae was first described in 1831. Later, in 1979, many genera were described and Loricariinae was divided into four subfamilies: Loricariini, Harttiini, Farlowellini, and Acestridiini. Eventually, the genera of Acestridiini was included under
Hypoptopomatinae The Hypoptopomatinae are a subfamily of catfishes (order Siluriformes) of the family Loricariidae, composed of 17 genus, genera and approximately 80 species. This subfamily represents about one-tenth of all loricariid species. It has been divide ...
and genera of Farlowellini was reclassified into Hartiini. This subfamily has been found to be monophyletic. The most recent papers conclude that there are 30 or 31 genera; the status of ''Cteniloricaria'' is dubious; although recognized as a synonym of '' Harttia'' by some, this conclusion was reached without diagnosing the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
of ''Cteniloricaria''.


Description

As loricariids, loricariinae species are characterized by a depressed body covered by bony plates, a unique pair of maxillary barbels, and modification of the mouth into a
suckermouth In fish, a suckermouth is a ventrally-oriented (inferior) mouth adapted for grazing on algae and small organisms that grow on submerged objects. All Loricariidae possess a suckermouth as do the cypriniform algae eaters of the genus ''Gyrinocheilus' ...
. Members of the subfamily Loricariinae show a greater diversity of lip structures and shapes than other loricariids. Loricariines are characterized by a long and depressed
caudal peduncle Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as see ...
(rectangular cross-section) and by the absence of an
adipose fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
. They also show dramatic variation in body shape, lip morphology and dentition. The sexual dimorphism is often pronounced and is expressed through the hypertrophy of odontodes on the
pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
rays, on the snout margin, and sometimes on the pre
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal c ...
area of mature males. Certain genera also show sexual differences in lip and tooth structures. The Harttiini are characterized by numerous and pedunculated teeth, a
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
with more branched rays, the absence of postorbital notches and predorsal keels, a rounded mouth,
papillose This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
lips weakly or not fringed, and short maxillary barbels. The Loricariini are characterized by a more important variation in lips and teeth shape, the frequent presence of postorbital notches and predorsal keels, longer maxillary barbels, and less numerous teeth and branched rays in the caudal fin.


Genera

Genera list based on Covain and Fisch-Muller (2007), except for ''Cteniloricaria''. *Tribe Loricariini **''Loricaria'' group ***'' Brochiloricaria'' ***'' Loricaria'' ***'' Paraloricaria'' ***'' Proloricaria'' ***''
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'' **''Loricariichthys'' group ***'' Furcodontichthys'' ***'' Hemiodontichthys'' ***'' Limatulichthys'' ***'' Loricariichthys'' ***'' Pseudoloricaria'' **''Pseudohemiodon'' group ***'' Apistoloricaria'' ***'' Crossoloricaria'' ***'' Dentectus'' ***'' Planiloricaria'' ***'' Pseudohemiodon'' ***'' Pyxiloricaria'' ***'' Reganella'' ***'' Rhadinoloricaria'' **''Rineloricaria'' group ***'' Dasyloricaria'' ***'' Fonchiiichthys'' ***'' Ixinandria'' ***'' Rineloricaria'' ***'' Spatuloricaria'' *Tribe Harttiini **'' Aposturisoma'' **'' Cteniloricaria'' **''
Farlowella ''Farlowella'' is a genus of fish in the family Loricariidae native to South America. This genus is broadly distributed in Amazon, Orinoco, Paraná and coastal rivers of the Guyana Shield. It is absent from the Pacific slope of the Andes and fro ...
'' **'' Harttia'' **'' Harttiella'' **'' Lamontichthys'' **'' Metaloricaria'' **'' Pterosturisoma'' **''
Sturisoma ''Sturisoma'' is a genus of armored catfishes native to Central and South America. Taxonomy ''Sturisoma'' has been shown to be sister to ''Farlowella''. Species There are currently 12 recognized species in this genus: * '' Sturisoma barbatum' ...
'' **'' Sturisomatichthys''


References

{{Taxonbar, from=Q137875 Loricariidae Fish of South America Fish subfamilies Taxa named by Charles Lucien Bonaparte