Elephants are the

largest Large means of great size. Large may also refer to: Mathematics * Arbitrarily large, a phrase in mathematics * Large cardinal, a property of certain transfinite numbers * Large category, a category with a proper class of objects and morphisms (o ...

existing land animals. Three living

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

are currently recognised: the African bush elephant, the

African forest elephant The African forest elephant (''Loxodonta cyclotis'') is one of the two living African elephant species. It is native to humid forests in West Africa and the Congo Basin. It is the smallest of the three living elephant species, reaching a shoulde ...

, and the

Asian elephant The Asian elephant (''Elephas maximus''), also known as the Asiatic elephant, is the only living species of the genus ''Elephas'' and is distributed throughout the Indian subcontinent and Southeast Asia, from India in the west, Nepal in the no ...

. They are the only surviving members of the

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

Elephantidae Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are terrestrial large mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the ...

and the order Proboscidea. The order was formerly much more diverse during the

Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...

, but most species became extinct during the Late Pleistocene epoch. Distinctive features of elephants include a long

proboscis A proboscis () is an elongated appendage from the head of an animal, either a vertebrate or an invertebrate. In invertebrates, the term usually refers to tubular mouthparts used for feeding and sucking. In vertebrates, a proboscis is an elong ...

called a trunk,

tusk Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine teeth, as with pigs and walruses, or, in the case of elephants, elongated incisors. Tusks share ...

s, large ear flaps, pillar-like legs, and tough but sensitive skin. The trunk is used for breathing, bringing food and water to the mouth, and grasping objects. Tusks, which are derived from the incisor teeth, serve both as weapons and as tools for moving objects and digging. The large ear flaps assist in maintaining a constant body temperature as well as in communication. African elephants have larger ears and concave backs, whereas Asian elephants have smaller ears, and convex or level backs. Elephants are scattered throughout sub-Saharan Africa,

South Asia South Asia is the southern subregion of Asia, which is defined in both geographical Geography (from Greek: , ''geographia''. Combination of Greek words ‘Geo’ (The Earth) and ‘Graphien’ (to describe), literally "earth descr ...

, and

Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainlan ...

and are found in different habitats, including

savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...

hs, forests, deserts, and

marsh A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found a ...

es. They are

herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpar ...

, and they stay near water when it is accessible. They are considered to be keystone species, due to their impact on their environments. Elephants have a

fission–fusion society In ethology, a fission–fusion society is one in which the size and composition of the social group change as time passes and animals move throughout the environment; animals merge into a group (fusion)—e.g. sleeping in one place—or split (fis ...

, in which multiple family groups come together to socialise. Females (cows) tend to live in family groups, which can consist of one female with her calves or several related females with offspring. The groups, which do not include bulls, are usually led by the oldest cow, known as the

matriarch Matriarchy is a social system in which women hold the primary power positions in roles of authority. In a broader sense it can also extend to moral authority, social privilege and control of property. While those definitions apply in general En ...

. Males (bulls) leave their family groups when they reach puberty and may live alone or with other males. Adult bulls mostly interact with family groups when looking for a mate. They enter a state of increased

testosterone Testosterone is the primary sex hormone and anabolic steroid in males. In humans, testosterone plays a key role in the development of male reproductive tissues such as testes and prostate, as well as promoting secondary sexual characteristi ...

and aggression known as

musth Musth or must (from Persian, )''The Oxford Dictionary and Thesaurus: American edition'', published 1996 by Oxford University Press; p. 984 is a periodic condition in bull (male) elephants characterized by aggressive behavior and accompanied ...

, which helps them gain dominance over other males as well as reproductive success. Calves are the centre of attention in their family groups and rely on their mothers for as long as three years. Elephants can live up to 70 years in the wild. They communicate by touch, sight, smell, and sound; elephants use

infrasound Infrasound, sometimes referred to as low status sound, describes sound waves with a frequency below the lower limit of human audibility (generally 20 Hz). Hearing becomes gradually less sensitive as frequency decreases, so for humans to perce ...

and

seismic communication Seismic or vibrational communication is a process of conveying information through mechanical (seismic) vibrations of the substrate. The substrate may be the earth, a plant stem or leaf, the surface of a body of water, a spider's web, a honeycomb, ...

over long distances. Elephant intelligence has been compared with that of

primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians ( monkeys and apes, the latter including ...

s and cetaceans. They appear to have self-awareness, and appear to show

empathy Empathy is the capacity to understand or feel what another person is experiencing from within their frame of reference, that is, the capacity to place oneself in another's position. Definitions of empathy encompass a broad range of social, co ...

for dying and dead family members. African bush elephants and Asian elephants are listed as

endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...

and African forest elephants as Critically Endangered by the

International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of nat ...

(IUCN). One of the biggest threats to elephant populations is the ivory trade, as the animals are poached for their ivory tusks. Other threats to wild elephants include

habitat destruction Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...

and conflicts with local people. Elephants are used as

working animal A working animal is an animal, usually domesticated, that is kept by humans and trained to perform tasks instead of being slaughtered to harvest animal products. Some are used for their physical strength (e.g. oxen and draft horses) or for ...

s in Asia. In the past, they were used in war; today, they are often controversially put on display in zoos, or exploited for entertainment in

circus A circus is a company of performers who put on diverse entertainment shows that may include clowns, acrobats, trained animals, trapeze acts, musicians, dancers, hoopers, tightrope walkers, jugglers, magicians, ventriloquists, and unicyclis ...

es. Elephants are highly recognisable and have been featured in art, folklore, religion, literature, and

popular culture Popular culture (also called mass culture or pop culture) is generally recognized by members of a society as a set of practices, beliefs, artistic output (also known as, popular art or mass art) and objects that are dominant or prevalent in a ...

Etymology

The word "elephant" is based on the

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

''elephas'' ( genitive ''elephantis'') ("elephant"), which is the Latinised form of the

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

ἐλέφας (''elephas'') (genitive ἐλέφαντος (''elephantos''), probably from a non-

Indo-European The Indo-European languages are a language family native to the overwhelming majority of Europe, the Iranian plateau, and the northern Indian subcontinent. Some European languages of this family, English, French, Portuguese, Russian, Dutc ...

language, likely Phoenician. It is attested in

Mycenaean Greek Mycenaean Greek is the most ancient attested form of the Greek language, on the Greek mainland and Crete in Mycenaean Greece (16th to 12th centuries BC), before the hypothesised Dorian invasion, often cited as the '' terminus ad quem'' for th ...

as ''e-re-pa'' (genitive ''e-re-pa-to'') in Linear B syllabic script. As in Mycenaean Greek,

Homer Homer (; grc, Ὅμηρος , ''Hómēros'') (born ) was a Greek poet who is credited as the author of the ''Iliad'' and the ''Odyssey'', two epic poems that are foundational works of ancient Greek literature. Homer is considered one of the ...

used the Greek word to mean

ivory Ivory is a hard, white material from the tusks (traditionally from elephants) and teeth of animals, that consists mainly of dentine, one of the physical structures of teeth and tusks. The chemical structure of the teeth and tusks of mammals i ...

, but after the time of

Herodotus Herodotus ( ; grc, , }; BC) was an ancient Greek historian and geographer A geographer is a physical scientist, social scientist or humanist whose area of study is geography, the study of Earth's natural environment and human society ...

, it also referred to the animal. The word "elephant" appears in

Middle English Middle English (abbreviated to ME) is a form of the English language that was spoken after the Norman conquest of 1066, until the late 15th century. The English language underwent distinct variations and developments following the Old English ...

as ''olyfaunt'' (c.1300) and was borrowed from

Old French Old French (, , ; Modern French: ) was the language spoken in most of the northern half of France from approximately the 8th to the 14th centuries. Rather than a unified language, Old French was a linkage of Romance dialects, mutually intellig ...

''oliphant'' (12th century).

Taxonomy

Alongside their closest relatives belong to the family

, the sole remaining family within the order Proboscidea. Their closest extant relatives are the

sirenia The Sirenia (), commonly referred to as sea-cows or sirenians, are an order of fully aquatic, herbivorous mammals that inhabit swamps, rivers, estuaries, marine wetlands, and coastal marine waters. The Sirenia currently comprise two distinct ...

ns (

dugong The dugong (; ''Dugong dugon'') is a marine mammal. It is one of four living species of the order Sirenia, which also includes three species of manatees. It is the only living representative of the once-diverse family Dugongidae; its closest m ...

s and

manatee Manatees (family Trichechidae, genus ''Trichechus'') are large, fully aquatic, mostly herbivorous marine mammals sometimes known as sea cows. There are three accepted living species of Trichechidae, representing three of the four living speci ...

s) and the hyraxes, with which they share the clade

Paenungulata Paenungulata (from Latin ''paene'' "almost" + ''ungulātus'' "having hoofs") is a clade of "sub-ungulates", which groups three extant mammal orders: Proboscidea (including elephants), Sirenia (sea cows, including dugongs and manatees), and H ...

within the superorder

Afrotheria Afrotheria ( from Latin ''Afro-'' "of Africa" + ''theria'' "wild beast") is a clade of mammals, the living members of which belong to groups that are either currently living in Africa or of African origin: golden moles, elephant shrews (also k ...

. Elephants and sirenians are further grouped in the clade

Tethytheria Tethytheria is a clade of mammals that includes the sirenians and proboscideans, as well as the extinct order Embrithopoda. Though there is strong anatomical and molecular support for the monophyly of Tethytheria, the interrelationships between ...

. Three species of elephants are recognised; the African bush elephant ('' Loxodonta africana'') and

forest elephant The African forest elephant (''Loxodonta cyclotis'') is one of the two living African elephant species. It is native to humid forests in West Africa and the Congo Basin. It is the smallest of the three living elephant species, reaching a shoulde ...

(''Loxodonta cyclotis'') of sub-Saharan Africa, and the

(''

Elephas ''Elephas'' is one of two surviving genera in the family of elephants, Elephantidae, with one surviving species, the Asian elephant, ''Elephas maximus''. Several extinct species have been identified as belonging to the genus, extending back to ...

maximus'') of South and

. African elephants have larger ears, a concave back, more wrinkled skin, a sloping abdomen, and two finger-like extensions at the tip of the trunk. Asian elephants have smaller ears, a convex or level back, smoother skin, a horizontal abdomen that occasionally sags in the middle and one extension at the tip of the trunk. The looped ridges on the

molars The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...

are narrower in the Asian elephant while those of the African are more diamond-shaped. The Asian elephant also has

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal c ...

bumps on its head and some patches of

depigmentation Depigmentation is the lightening of the skin or loss of pigment. Depigmentation of the skin can be caused by a number of local and systemic conditions. The pigment loss can be partial (injury to the skin) or complete (caused by vitiligo). It can be ...

on its skin.Shoshani, pp. 38–41. Among African elephants, forest elephants have smaller and more rounded ears and thinner and straighter tusks than bush elephants and are limited in range to the forested areas of

western Western may refer to: Places *Western, Nebraska, a village in the US *Western, New York, a town in the US *Western Creek, Tasmania, a locality in Australia *Western Junction, Tasmania, a locality in Australia *Western world, countries that id ...

and

Central Africa Central Africa is a subregion of the African continent comprising various countries according to different definitions. Angola, Burundi, the Central African Republic, Chad, the Democratic Republic of the Congo, the Republic of the Congo ...

. Both were traditionally considered a single species, ''Loxodonta africana'', but molecular studies have affirmed their status as separate species. In 2017, DNA sequence analysis showed that ''L. cyclotis'' is more closely related to the extinct '' Palaeoloxodon antiquus'', than it is to ''L. africana,'' possibly undermining the genus ''Loxodonta'' as a whole.

Evolution and extinct relatives

Over 180 extinct members and three major evolutionary radiations of the order Proboscidea have been recorded. The earliest proboscids, the African '' Eritherium'' and '' Phosphatherium'' of the late

Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pal ...

, heralded the first radiation. The

Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...

included ''

Numidotherium ''Numidotherium'' ("Numidia beast") is an extinct genus of early proboscideans, discovered in 1984, that lived during the middle Eocene of North Africa some 46 million years ago. It was about 90-100 cm tall at the shoulder and weighed about 250- ...

'', ''

Moeritherium ''Moeritherium'' ("the beast from Lake Moeris") is an extinct genus of primitive proboscideans. These prehistoric mammals are related to the elephant and, more distantly, sea cows and hyraxes. They lived during the Eocene epoch. Description ''M ...

,'' and ''

Barytherium ''Barytherium'' (meaning "heavy beast") is a genus of an extinct family (Barytheriidae) of primitive proboscideans that lived during the late Eocene and early Oligocene in North Africa. The type species is ''Barytherium grave'', found at the ...

'' from Africa. These animals were relatively small and aquatic. Later on, genera such as ''

Phiomia ''Phiomia'' is an extinct genus of basal proboscid that lived in what is now Northern Africa during the Late Eocene to Early Oligocene some 37–30 million years ago.Strauss, BPrehistoric Elephant Pictures and Profiles/ref> "''Phiomia serridens''" ...

'' and ''

Palaeomastodon ''Palaeomastodon'' an extinct genus of Proboscidea. ''Palaeomastodon'' fossils have been found in Africa, where they lived some 36-35 million years ago. They are believed to be the ancestors of elephants or mastodons. ''Palaeomastodon'' lived in ...

'' arose; the latter likely inhabited forests and open woodlands. Proboscidean diversity declined during the Oligocene. One notable species of this epoch was '' Eritreum melakeghebrekristosi'' of the Horn of Africa, which may have been an ancestor to several later species. The beginning of the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

saw the second diversification, with the appearance of the deinotheres and the mammutids. The former were related to ''Barytherium'' and lived in Africa and Eurasia,Sukumar, pp. 16–19. while the latter may have descended from ''Eritreum'' and spread to North America. The second radiation was represented by the emergence of the gomphotheres in the Miocene, which likely evolved from ''Eritreum'' and originated in Africa, spreading to every continent except Australia and Antarctica. Members of this group included ''

Gomphotherium ''Gomphotherium'' (; "welded beast") is an extinct genus of proboscids from the Neogene and early Pleistocene of Eurasia, Africa, North America and Asia. As of 2021, two species, ''G. annectens'' and possibly ''G. subtapiroideum'', are also k ...

'' and '' Platybelodon''. The third radiation started in the late Miocene and led to the arrival of the elephantids, which descended from, and slowly replaced, the gomphotheres. The African ''

Primelephas ''Primelephas'' is a genus of Elephantinae that existed during the Miocene and Pliocene epochs. The name of the genus suggests 'first elephant'. These primitive elephantids are thought to be the common ancestor of ''Mammuthus'', the mammoths ...

gomphotheroides'' gave rise to ''Loxodonta'', ''Mammuthus,'' and ''Elephas''. ''Loxodonta'' branched off earliest around the Miocene and

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58stegodontids, another proboscidean group descended from gomphotheres, spread throughout Asia, including the Indian subcontinent, China, southeast Asia, and Japan. Mammutids continued to evolve into new species, such as the

American mastodon American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, p ...

. At the beginning of the

, elephantids experienced a high rate of speciation. The Pleistocene also saw the arrival of ''

Palaeoloxodon namadicus ''Palaeoloxodon namadicus'' or the Asian straight-tusked elephant, is an extinct species of prehistoric elephant known from the early Middle to Late Pleistocene of the Indian subcontinent, and possibly also elsewhere in Asia. Some authorities r ...

'', the largest terrestrial mammal of all time. '' Loxodonta atlantica'' became the most common species in northern and southern Africa but was replaced by ''Elephas iolensis'' later in the Pleistocene. Only when ''Elephas'' disappeared from Africa did ''Loxodonta'' become dominant once again, this time in the form of the modern species. ''Elephas'' diversified into new species in Asia, such as ''E. hysudricus'' and ''E. platycephus''; the latter the likely ancestor of the modern Asian elephant.Sukumar, p. 44. ''Mammuthus'' evolved into several species, including the well-known

woolly mammoth The woolly mammoth (''Mammuthus primigenius'') is an extinct species of mammoth that lived during the Pleistocene until its extinction in the Holocene epoch. It was one of the last in a line of mammoth species, beginning with '' Mammuthus s ...

.Sukumar, pp. 28–31. Interbreeding appears to have been common among elephantid species, which in some cases led to species with three ancestral genetic components, such as the ''Palaeoloxodon antiquus''. In the Late Pleistocene, most proboscidean species vanished during the

Quaternary glaciation The Quaternary glaciation, also known as the Pleistocene glaciation, is an alternating series of glacial and interglacial periods during the Quaternary period that began 2.58 Ma (million years ago) and is ongoing. Although geologists describ ...

which

killed off The killing off of a character is a device in fiction, whereby a character dies, but the story continues. The term, frequently applied to television, film, video game, anime, manga and chronological series, often denotes an untimely or unexpected ...

50% of genera weighing over worldwide. Proboscideans experienced several evolutionary trends, such as an increase in size, which led to many giant species that stood up to tall. As with other

megaherbivore Megaherbivores (Greek μέγας megas "large" and Latin ''herbivora'' "herbivore") are large terrestrial herbivores that can exceed in weight. This polyphyletic group of megafauna includes elephants, rhinos, hippos, and giraffes. The largest ...

s, including the extinct sauropod

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

s, the large size of elephants likely developed to allow them to survive on vegetation with low nutritional value. Their limbs grew longer and the feet shorter and broader. The feet were originally plantigrade and developed into a digitigrade stance with cushion pads and the sesamoid bone providing support. Early proboscideans developed longer

mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...

s and smaller craniums while more derived ones developed shorter mandibles, which shifted the head's

centre of gravity In physics, the center of mass of a distribution of mass in space (sometimes referred to as the balance point) is the unique point where the weighted relative position of the distributed mass sums to zero. This is the point to which a force ma ...

. The skull grew larger, especially the cranium, while the neck shortened to provide better support for the skull. The increase in size led to the development and elongation of the mobile trunk to provide reach. The number of premolars, incisors and

canines Canine may refer to: Zoology and anatomy * a dog-like Canid animal in the subfamily Caninae ** ''Canis'', a genus including dogs, wolves, coyotes, and jackals ** Dog, the domestic dog * Canine tooth, in mammalian oral anatomy People with the surn ...

decreased. The cheek teeth (molars and premolars) of proboscideans became larger and more specialized, especially after elephants started to switch from C3-plants to C4-grasses, which caused their teeth to undergo a three-fold increase in teeth height as well as substantial multiplication of lamellae after about five million years ago. Only in the last million years or so did they return to a diet mainly consisting of C3 trees and shrubs. The upper second incisors grew into tusks, which varied in shape from straight, to curved (either upward or downward), to spiralled, depending on the species. Some proboscideans developed tusks from their lower incisors. Elephants retain certain features from their aquatic ancestry, such as their

middle ear The middle ear is the portion of the ear medial to the eardrum, and distal to the oval window of the cochlea (of the inner ear). The mammalian middle ear contains three ossicles, which transfer the vibrations of the eardrum into waves in the ...

anatomy. Several species of proboscideans lived on islands and experienced

insular dwarfism Insular dwarfism, a form of phyletic dwarfism, is the process and condition of large animals evolving or having a reduced body size when their population's range is limited to a small environment, primarily islands. This natural process is disti ...

. This occurred primarily during the Pleistocene when some elephant populations became isolated by fluctuating sea levels, although

dwarf elephant Dwarf elephants are prehistoric members of the order Proboscidea which, through the process of allopatric speciation on islands, evolved much smaller body sizes (around ) in comparison with their immediate ancestors. Dwarf elephants are an example ...

s did exist earlier in the Pliocene. These elephants likely grew smaller on islands due to a lack of large or viable predator populations and limited resources. By contrast, small mammals such as

rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...

s develop gigantism in these conditions. Dwarf elephants are known to have lived in

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...

, the

Channel Islands of California The Channel Islands () are an eight-island archipelago located within the Southern California Bight in the Pacific Ocean, off the coast of California. The four Northern Channel Islands are part of the Transverse Ranges geologic province, ...

, and several islands of the

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western Europe, Western and Southern Europe and Anatolia, on the south by North Africa ...

.Sukumar, pp. 31–33.

Anatomy

Size

Elephants are the largest living terrestrial animals. African bush elephants are the largest species, with males being tall at the shoulder with a body mass of and females standing tall at the shoulder with a body mass of . Male Asian elephants are usually about tall at the shoulder and whereas females are tall at the shoulder and . African forest elephants are the smallest species, with males usually being around tall at the shoulder and . Male African bush elephants are typically 23% taller than females, whereas male Asian elephants are only around 15% taller than females.

Bones

The skeleton of the elephant is made up of 326–351 bones. The vertebrae are connected by tight joints, which limit the backbone's flexibility. African elephants have 21 pairs of ribs, while Asian elephants have 19 or 20 pairs.

Head

An elephant's skull is resilient enough to withstand the forces generated by the leverage of the tusks and head-to-head collisions. The back of the skull is flattened and spread out, creating arches that protect the

brain A brain is an organ that serves as the center of the nervous system in all vertebrate and most invertebrate animals. It is located in the head, usually close to the sensory organs for senses such as vision. It is the most complex organ in a ve ...

in every direction. The skull contains air cavities (

sinuses Paranasal sinuses are a group of four paired air-filled spaces that surround the nasal cavity. The maxillary sinuses are located under the eyes; the frontal sinuses are above the eyes; the ethmoidal sinuses are between the eyes and the sphenoid ...

) that reduce the weight of the skull while maintaining overall strength. These cavities give the inside of the skull a

honeycomb A honeycomb is a mass of hexagonal prismatic wax cells built by honey bees in their nests to contain their larvae and stores of honey and pollen. Beekeepers may remove the entire honeycomb to harvest honey. Honey bees consume about of honey ...

-like appearance. The cranium is particularly large and provides enough room for the attachment of muscles to support the entire head. The lower jaw is solid and heavy.Shoshani, pp. 68–70. Because of the size of the head, the neck is relatively short to provide better support. Lacking a

lacrimal apparatus The lacrimal apparatus is the physiological system containing the orbital structures for tear production and drainage.Cassin, B. and Solomon, S. ''Dictionary of Eye Terminology''. Gainesville, Florida: Triad Publishing Company, 1990. It consist ...

, the eye relies on the

harderian gland The Harderian gland is a gland found within the eye's orbit that occurs in tetrapods (reptiles, amphibians, birds and mammals) that possess a nictitating membrane. The gland can be compound tubular or compound tubuloalveolar, and the fluid it s ...

to keep it moist. A durable nictitating membrane protects the eye globe. The animal's

field of vision The visual field is the "spatial array of visual sensations available to observation in introspectionist psychological experiments". Or simply, visual field can be defined as the entire area that can be seen when an eye is fixed straight at a point ...

is compromised by the location and limited mobility of the eyes. Elephants are considered dichromats and they can see well in dim light but not in bright light.

Ears

Elephant ears have thick bases with thin tips. The ear flaps, or

pinnae The auricle or auricula is the visible part of the ear that is outside the head. It is also called the pinna (Latin for "wing" or " fin", plural pinnae), a term that is used more in zoology. Structure The diagram shows the shape and location ...

, contain numerous blood vessels called

capillaries A capillary is a small blood vessel from 5 to 10 micrometres (μm) in diameter. Capillaries are composed of only the tunica intima, consisting of a thin wall of simple squamous endothelial cells. They are the smallest blood vessels in the body: ...

. Warm blood flows into the capillaries, helping to release excess body heat into the environment. This occurs when the pinnae are still, and the animal can enhance the effect by flapping them. Larger ear surfaces contain more capillaries, and more heat can be released. Of all the elephants, African bush elephants live in the hottest climates, and have the largest ear flaps. Elephants are capable of hearing at low frequencies and are most sensitive at 1 kHz (in close proximity to the Soprano C).

Trunk

The trunk, or

, is a fusion of the nose and upper lip, although in early

fetal A fetus or foetus (; plural fetuses, feti, foetuses, or foeti) is the unborn offspring that develops from an animal embryo. Following embryonic development the fetal stage of development takes place. In human prenatal development, fetal develo ...

life, the upper lip and trunk are separated. The trunk is elongated and specialised to become the elephant's most important and versatile appendage. It contains up to 150,000 separate

muscle fascicle A muscle fascicle is a bundle of skeletal muscle fibers surrounded by perimysium, a type of connective tissue. Structure Muscle cells are grouped into muscle fascicles by enveloping perimysium connective tissue. Fascicles are bundled togethe ...

s, with no bone and little fat. These paired muscles consist of two major types: superficial (surface) and internal. The former are divided into dorsals, ventrals, and laterals while the latter are divided into transverse and

radiating In physics, radiation is the emission or transmission of energy in the form of waves or particles through space or through a material medium. This includes: * ''electromagnetic radiation'', such as radio waves, microwaves, infrared, visib ...

muscles. The muscles of the trunk connect to a bony opening in the skull. The

nasal septum The nasal septum () separates the left and right airways of the nasal cavity, dividing the two nostrils. It is depressed by the depressor septi nasi muscle. Structure The fleshy external end of the nasal septum is called the columella or co ...

is composed of tiny muscle units that stretch horizontally between the nostrils. Cartilage divides the nostrils at the base.Shoshani, pp. 74–77. As a

muscular hydrostat A muscular hydrostat is a biological structure found in animals. It is used to manipulate items (including food) or to move its host about and consists mainly of muscles with no skeletal support. It performs its hydraulic movement without fluid in ...

, the trunk moves by precisely coordinated muscle contractions. The muscles work both with and against each other. A unique proboscis nerve – formed by the maxillary and

facial nerve The facial nerve, also known as the seventh cranial nerve, cranial nerve VII, or simply CN VII, is a cranial nerve that emerges from the pons of the brainstem, controls the muscles of facial expression, and functions in the conveyance of taste ...

s – runs along both sides of the trunk. Elephant trunks have multiple functions, including breathing, olfaction, touching, grasping, and sound production. The animal's sense of smell may be four times as sensitive as that of a

bloodhound The bloodhound is a large scent hound, originally bred for hunting deer, wild boar and, since the Middle Ages, for tracking people. Believed to be descended from hounds once kept at the Abbey of Saint-Hubert, Belgium, in French it is called, ...

. The trunk's ability to make powerful twisting and coiling movements allows it to collect food, wrestle with other elephants,Kingdon, p. 9. and lift up to . It can be used for delicate tasks, such as wiping an eye and checking an orifice, and is capable of cracking a peanut shell without breaking the seed. With its trunk, an elephant can reach items at heights of up to and dig for water under mud or sand. Individuals may show lateral preference when grasping with their trunks: some prefer to twist them to the left, others to the right. Elephants are capable of dilating their nostrils at a radius of nearly 30%, increasing the nasal volume by 64%, and can inhale at over which is around 30 times the speed of a human sneeze. Elephants can suck up food and water both to spray in the mouth and, in the case of the latter, to sprinkle on their bodies. An adult Asian elephant is capable of holding of water in its trunk. They will also spray dust or grass on themselves. When underwater, the elephant uses its trunk as a snorkel. The African elephant has two finger-like extensions at the tip of the trunk that allow it to grasp and bring food to its mouth. The Asian elephant has only one and relies more on wrapping around a food item and squeezing it into its mouth. Asian elephants have more muscle coordination and can perform more complex tasks. Losing the trunk would be detrimental to an elephant's survival, although in rare cases, individuals have survived with shortened ones. One elephant has been observed to graze by kneeling on its front legs, raising on its hind legs and taking in grass with its lips.

Floppy trunk syndrome Floppy trunk syndrome (abbreviated FTS, also known as flaccid trunk paralysis) is a condition that causes trunk paralysis in African bush elephants. Initially observed in 1989, the syndrome primarily affected bull elephants in several select region ...

is a condition of trunk paralysis in African bush elephants caused by the degradation of the

peripheral nerves The peripheral nervous system (PNS) is one of two components that make up the nervous system of bilateral animals, with the other part being the central nervous system (CNS). The PNS consists of nerves and ganglia, which lie outside the brain an ...

and muscles beginning at the tip.

Teeth

Elephants usually have 26 teeth: the

incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...

s, known as the

s, 12

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...

premolars, and 12

. Unlike most mammals, which grow baby teeth and then replace them with a single permanent set of adult teeth, elephants are

polyphyodont A polyphyodont is any animal whose teeth are continually replaced. In contrast, diphyodonts are characterized by having only two successive sets of teeth. Polyphyodonts include most toothed fishes, many reptiles such as crocodiles and geckos, ...

s that have cycles of tooth rotation throughout their lives. The chewing teeth are replaced six times in a typical elephant's lifetime. Teeth are not replaced by new ones emerging from the jaws vertically as in most mammals. Instead, new teeth grow in at the back of the mouth and move forward to push out the old ones. The first chewing tooth on each side of the jaw falls out when the elephant is two to three years old. The second set of chewing teeth falls out at four to six years old. The third set falls out at 9–15 years of age and set four lasts until 18–28 years of age. The fifth set of teeth falls out at the early 40s. The sixth (and usually final) set must last the elephant the rest of its life. Elephant teeth have loop-shaped dental ridges, which are thicker and more diamond-shaped in African elephants.Shoshani, pp. 70–71.

Tusks

The tusks of an elephant are modified second incisors in the upper jaw. They replace deciduous milk teeth at 6–12 months of age and grow continuously at about a year. A newly developed tusk has a smooth enamel cap that eventually wears off. The dentine is known as

and its cross-section consists of crisscrossing line patterns, known as "engine turning", which create diamond-shaped areas. As a piece of living tissue, a tusk is relatively soft; it is as hard as the mineral calcite. Much of the tusk can be seen outside; the rest is in a socket in the skull. At least one-third of the tusk contains the

pulp Pulp may refer to: * Pulp (fruit), the inner flesh of fruit Engineering * Dissolving pulp, highly purified cellulose used in fibre and film manufacture * Pulp (paper), the fibrous material used to make paper * Molded pulp, a packaging material ...

and some have nerves stretching to the tip. Thus it would be difficult to remove it without harming the animal. When removed, ivory begins to dry up and crack if not kept cool and moist. Tusks serve multiple purposes. They are used for digging for water, salt, and roots; debarking or marking trees; and for moving trees and branches when clearing a path. When fighting, they are used to attack and defend, and to protect the trunk. Like humans, who are typically right- or left-handed, elephants are usually right- or left-tusked. The dominant tusk, called the master tusk, is generally more worn down, as it is shorter with a rounder tip. For the African elephants, tusks are present in both males and females, and are around the same length in both sexes, reaching up to , but those of males tend to be thicker. In earlier times, elephant tusks weighing over 200 pounds (more than 90 kg) were not uncommon, though it is rare today to see any over . In the Asian species, only the males have large tusks. Female Asians have very small tusks, or none at all.Shoshani, pp. 71–74. Tuskless males exist and are particularly common among Sri Lankan elephants. Asian males can have tusks as long as Africans', but they are usually slimmer and lighter; the largest recorded was long and weighed . Hunting for elephant ivory in Africa and Asia has led to

natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the heritable traits characteristic of a population over generations. Cha ...

for shorter tusks and tusklessness.

Skin

An elephant's skin is generally very tough, at thick on the back and parts of the head. The skin around the mouth, anus, and inside of the ear is considerably thinner. Elephants typically have grey skin, but African elephants look brown or reddish after wallowing in coloured mud. Asian elephants have some patches of depigmentation, particularly on the forehead and ears and the areas around them. Calves have brownish or reddish hair, especially on the head and back. As elephants mature, their hair darkens and becomes sparser, but dense concentrations of hair and bristles remain on the end of the tail as well as the chin,

genitals A sex organ (or reproductive organ) is any part of an animal or plant that is involved in sexual reproduction. The reproductive organs together constitute the reproductive system. In animals, the testis in the male, and the ovary in the female, a ...

and the areas around the eyes and ear openings. Normally the skin of an Asian elephant is covered with more hair than its African counterpart.Shoshani, pp. 66–67. Their hair is thought to be for

thermoregulation Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperature ...

, helping them lose heat in their hot environments. An elephant uses mud as a sunscreen, protecting its skin from

ultraviolet Ultraviolet (UV) is a form of electromagnetic radiation with wavelength from 10 nm (with a corresponding frequency around 30 PHz) to 400 nm (750 THz), shorter than that of visible light, but longer than X-rays. UV radiation ...

light. Although tough, an elephant's skin is very sensitive. Without regular

mud bath A mud bath is a bath of mud, commonly found in areas where hot spring water can combine with volcanic ash. Mud baths have a long history that dates back thousands of years. Mud baths are conceived as public bathing spaces created in open areas ...

s to protect it from burning, insect bites and moisture loss, an elephant's skin suffers serious damage. After bathing, the elephant will usually use its trunk to blow dust onto its body and this dries into a protective crust. Elephants have difficulty releasing heat through the skin because of their low

surface-area-to-volume ratio The surface-area-to-volume ratio, also called the surface-to-volume ratio and variously denoted sa/vol or SA:V, is the amount of surface area per unit volume of an object or collection of objects. SA:V is an important concept in science and engin ...

, which is many times smaller than that of a human. They have even been observed lifting up their legs, presumably in an effort to expose their soles to the air.

Legs, locomotion, and posture

To support the animal's weight, an elephant's limbs are positioned more vertically under the body than in most other mammals. The long bones of the limbs have

cancellous bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, and ...

in place of medullary cavities. This strengthens the bones while still allowing

haematopoiesis Haematopoiesis (, from Greek , 'blood' and 'to make'; also hematopoiesis in American English; sometimes also h(a)emopoiesis) is the formation of blood cellular components. All cellular blood components are derived from haematopoietic stem cell ...

.Shoshani, pp. 69–70. Both the front and hind limbs can support an elephant's weight, although 60% is borne by the front. Since the limb bones are placed on top of each other and under the body, an elephant can stand still for long periods of time without using much energy. Elephants are incapable of rotating their front legs, as the

ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...

and

radius In classical geometry, a radius ( : radii) of a circle or sphere is any of the line segments from its center to its perimeter, and in more modern usage, it is also their length. The name comes from the latin ''radius'', meaning ray but also the ...

are fixed in pronation; the "palm" of the manus faces backward. The pronator quadratus and the pronator teres are either reduced or absent. The circular feet of an elephant have soft tissues or "cushion pads" beneath the manus or pes, which distribute the weight of the animal. They appear to have a sesamoid, an extra "toe" similar in placement to a giant panda's extra "thumb", that also helps in weight distribution. As many as five toenails can be found on both the front and hind feet. Elephants can move both forwards and backwards, but cannot

trot The trot is a ten-beat diagonal horse gait where the diagonal pairs of legs move forward at the same time with a moment of suspension between each beat. It has a wide variation in possible speeds, but averages about . A very slow trot is someti ...

jump Jumping is a form of locomotion or movement in which an organism or non-living (e.g., robotic) mechanical system propels itself through the air along a ballistic trajectory. Jump or Jumping also may refer to: Places * Jump, Kentucky or Jump S ...

, or

gallop The canter and gallop are variations on the fastest gait that can be performed by a horse or other equine. The canter is a controlled three-beat gait, while the gallop is a faster, four-beat variation of the same gait. It is a natural gait pos ...

. They use only two gaits when moving on land: the walk and a faster gait similar to running. In walking, the legs act as pendulums, with the hips and shoulders rising and falling while the foot is planted on the ground. With no "aerial phase", the fast gait does not meet all the criteria of running, although the elephant uses its legs much like other running animals, with the hips and shoulders falling and then rising while the feet are on the ground. Fast-moving elephants appear to 'run' with their front legs, but 'walk' with their hind legs and can reach a top speed of . At this speed, most other

quadrupeds Quadrupedalism is a form of locomotion where four limbs are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four limbs is said to be a quadruped (from Latin ''quattuor' ...

are well into a gallop, even accounting for leg length. Spring-like kinetics could explain the difference between the motion of elephants and other animals. During locomotion, the cushion pads expand and contract, and reduce both the pain and noise that would come from a very heavy animal moving. Elephants are capable swimmers. They have been recorded swimming for up to six hours without touching the bottom, and have swum as far as at a stretch and at speeds of up to .

Organs

The brain of an elephant weighs compared to for a human brain. It is the largest of all terrestrial mammals. While the elephant brain is larger overall, it is proportionally smaller than the human brain when compared to their size. At birth, an elephant's brain already weighs 30–40% of its adult weight. The

cerebrum The cerebrum, telencephalon or endbrain is the largest part of the brain containing the cerebral cortex (of the two cerebral hemispheres), as well as several subcortical structures, including the hippocampus, basal ganglia, and olfactory bulb ...

and cerebellum are well developed, and the

temporal lobe The temporal lobe is one of the four major lobes of the cerebral cortex in the brain of mammals. The temporal lobe is located beneath the lateral fissure on both cerebral hemispheres of the mammalian brain. The temporal lobe is involved in pro ...

s are so large that they bulge out laterally. Their temporal lobes are proportionally larger than in other animals, including humans. The throat of an elephant appears to contain a pouch where it can store water for later use. The larynx of the elephant is the largest known among mammals. The

vocal fold The human voice consists of sound made by a human being using the vocal tract, including talking, singing, laughing, crying, screaming, shouting, humming or yelling. The human voice frequency is specifically a part of human sound product ...

s are long and are attached close to the

epiglottis The epiglottis is a leaf-shaped flap in the throat that prevents food and water from entering the trachea and the lungs. It stays open during breathing, allowing air into the larynx. During swallowing, it closes to prevent aspiration of food in ...

base. When comparing an elephant's vocal folds to those of a human, an elephant's are longer, thicker, and have a larger cross-sectional area. In addition, they are tilted at 45 degrees and positioned more anteriorly than a human's vocal folds. The heart of an elephant weighs . It has a double-pointed

apex The apex is the highest point of something. The word may also refer to: Arts and media Fictional entities * Apex (comics), a teenaged super villainess in the Marvel Universe * Ape-X, a super-intelligent ape in the Squadron Supreme universe *Apex, ...

, an unusual trait among mammals. In addition, the ventricles separate near the top of the heart, a trait they share with sirenians. When standing, the elephant's heart beats approximately 30 times per minute. Unlike many other animals, the heart rate speeds up by 8 to 10 beats per minute when the elephant is lying down. The blood vessels in most of the body are wide and thick and can withstand high blood pressures. The lungs are attached to the diaphragm, and breathing relies mainly on the diaphragm rather than the expansion of the ribcage.Shoshani, pp. 78–79. Connective tissue exists in place of the

pleural cavity The pleural cavity, pleural space, or interpleural space is the potential space between the pleurae of the pleural sac that surrounds each lung. A small amount of serous pleural fluid is maintained in the pleural cavity to enable lubrication b ...

. This may allow the animal to deal with the pressure differences when its body is underwater and its trunk is breaking the surface for air, although this explanation has been questioned. Another possible function for this adaptation is that it helps the animal suck up water through the trunk. Elephants inhale mostly through the trunk, although some air goes through the mouth. They have a

hindgut fermentation Hindgut fermentation is a digestive process seen in monogastric herbivores, animals with a simple, single-chambered stomach. Cellulose is digested with the aid of symbiotic bacteria.penis A penis (plural ''penises'' or ''penes'' () is the primary sexual organ that male animals use to inseminate females (or hermaphrodites) during copulation. Such organs occur in many animals, both vertebrate and invertebrate, but males d ...

can reach a length of and a diameter of at the base. It is S-shaped when fully erect and has a Y-shaped

orifice An orifice is any opening, mouth, hole or vent, as in a pipe, a plate, or a body * Body orifice, any opening in the body of a human or animal *Orifice plate, a restriction used to measure flow or to control pressure or flow, sometimes given specia ...

. The female has a well-developed clitoris at up to . The vulva is located between the hind legs instead of near the tail as in most mammals. Determining pregnancy status can be difficult due to the animal's large abdominal cavity. The female's

mammary gland A mammary gland is an exocrine gland in humans and other mammals that produces milk to feed young offspring. Mammals get their name from the Latin word ''mamma'', "breast". The mammary glands are arranged in organs such as the breasts in pri ...

s occupy the space between the front legs, which puts the suckling calf within reach of the female's trunk. Elephants have a unique organ, the

temporal gland Temporins are a family of peptides isolated originally from the skin secretion of the European red frog, ''Rana temporaria''. Peptides belonging to the temporin family have been isolated also from closely related North American frogs, such as ''R ...

, located in both sides of the head. This organ is associated with sexual behaviour, and males secrete a fluid from it when in

. Females have also been observed with secretions from the temporal glands.Sukumar, p. 149.

Body temperature

Elephants are

homeotherms Warm-blooded is an informal term referring to animal species which can maintain a body temperature higher than their environment. In particular, homeothermic species maintain a stable body temperature by regulating metabolic processes. The onl ...

, and maintain their average body temperature at ~ 36 °C, with minimum 35.2 °C during cool season, and maximum 38.0 °C during hot dry season. Sweat glands are absent in the elephant's skin, but water diffuses through the skin, allowing cooling by evaporative loss. Other physiological or behavioral features may assist with thermoregulation such as flapping ears, mud bathing, spraying water on the skin, seeking shade, and adopting different walking patterns. In addition, the interconnected crevices in the elephant's skin is thought to impede dehydration and improve thermal regulation over a long period of time.

Behaviour and life history

Ecology and activities

The African bush elephant can be found in habitats as diverse as dry

hs, deserts,

es, and lake shores, and in elevations from sea level to mountains above the snow line. Forest elephants mainly live in equatorial forests but will enter

gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...

s and

ecotone An ecotone is a transition area between two biological communities, where two communities meet and integrate. It may be narrow or wide, and it may be local (the zone between a field and forest) or regional (the transition between forest and gras ...

s between forests and savannahs.Shoshani, pp. 42–51. Asian elephants prefer areas with a mix of grasses, low woody plants, and trees, primarily inhabiting dry thorn-scrub forests in southern India and Sri Lanka and

evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, Live oak, acacias, magnolia, and banksia in more temperate zo ...

s in Malaya. Elephants are

and will eat leaves, twigs, fruit, bark, grass and roots. They are born with sterile intestines and require bacteria obtained from their mother's feces to digest vegetation. African elephants are mostly

browsers Browse, browser or browsing may refer to: Programs * Web browser, a program used to access the World Wide Web *Code browser, a program for navigating source code * File browser or file manager, a program used to manage files and related objects * ...

while Asian elephants are mainly grazers. They can consume as much as of food and of water in a day. Elephants tend to stay near water sources. Major feeding bouts take place in the morning, afternoon and night. At midday, elephants rest under trees and may doze off while standing. Sleeping occurs at night while the animal is lying down. Elephants average 3–4 hours of sleep per day. Both males and family groups typically move a day, but distances as far as have been recorded in the Etosha region of Namibia. Elephants go on seasonal migrations in search of food, water, minerals, and mates. At

Chobe National Park Chobe National Park is Botswana's first national park, and also the most biologically diverse. Located in the north of the country, it is Botswana's third largest park, after Central Kalahari Game Reserve and Gemsbok National Park, and has one ...

, Botswana, herds travel to visit the river when the local waterholes dry up. Because of their large size, elephants have a huge impact on their environments and are considered keystone species. Their habit of uprooting trees and undergrowth can transform savannah into grasslands; when they dig for water during drought, they create waterholes that can be used by other animals. They can enlarge waterholes when they bathe and wallow in them. At Mount Elgon, elephants excavate caves that are used by

ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, ...

s, hyraxes, bats, birds and insects. Elephants are important seed dispersers; African forest elephants ingest and defecate seeds, with either no effect or a positive effect on germination. The seeds are typically dispersed in large amounts over great distances. In Asian forests, large seeds require giant herbivores like elephants and

rhinoceros A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species ...

for transport and dispersal. This ecological niche cannot be filled by the next largest herbivore, the

tapir Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a pig, with a short, prehensile nose trunk. Tapirs inhabit jungle and forest regions of South and Central America, with one species inh ...

. Because most of the food elephants eat goes undigested, their dung can provide food for other animals, such as

dung beetle Dung beetles are beetles that feed on feces. Some species of dung beetles can bury dung 250 times their own mass in one night. Many dung beetles, known as ''rollers'', roll dung into round balls, which are used as a food source or breeding cha ...

s and monkeys.Shoshani, pp. 226–29. Elephants can have a negative impact on ecosystems. At

Murchison Falls National Park Murchison Falls National Park (MFNP) is a national park in Uganda and managed by the Ugandan Wildlife Authority. It is in north-western Uganda, spreading inland from the shores of Lake Albert, around the Victoria Nile, up to the Karuma Falls. ...

in Uganda, the overabundance of elephants has threatened several species of small birds that depend on woodlands. Their weight can compact the soil, which causes the rain to run off, leading to

erosion Erosion is the action of surface processes (such as water flow or wind) that removes soil, rock, or dissolved material from one location on the Earth's crust, and then transports it to another location where it is deposited. Erosion is dis ...

.Eltringham, pp. 124–27. Forest elephant

Elephants typically coexist peacefully with other herbivores, which will usually stay out of their way. Some aggressive interactions between elephants and rhinoceros have been recorded. At

Aberdare National Park The Aberdare National Park is a protected area in the Aberdare Mountain Range in central Kenya located east of the East African Rift Valley. It covers the higher areas and the Aberdare Salient to the east. Overview The park is located about 10 ...

, Kenya, a rhino attacked an elephant calf and was killed by the other elephants in the group. At Hluhluwe–Umfolozi Game Reserve,

South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...

, introduced young orphan elephants went on a killing spree that claimed the lives of 36 rhinos during the 1990s, but ended with the introduction of older males. The size of adult elephants makes them nearly invulnerable to

predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...

s. Calves may be preyed on by lions, spotted hyenas, and wild dogs in Africa and tigers in Asia. The lions of Savuti, Botswana, have adapted to hunting elephants, mostly calves, juveniles or even sub-adults, during the dry season, and a pride of 30 lions has been normally recorded killing juvenile individuals between the ages of four and eleven years, and a young bull of about 15 years in an exceptional case. There are rare reports of adult Asian elephants falling prey to

tiger The tiger (''Panthera tigris'') is the largest living cat species and a member of the genus ''Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily preys on ...

s. Elephants appear to distinguish between the growls of larger predators like tigers and smaller predators like leopards (which have not been recorded killing calves); they react to leopards less fearfully and more aggressively. Elephants tend to have high numbers of parasites, particularly nematodes, compared to other herbivores. This is due to lower predation pressures that would otherwise kill off many of the individuals with significant parasite loads.

Social organisation

Female elephants spend their entire lives in tight-knit

matrilineal Matrilineality is the tracing of kinship through the female line. It may also correlate with a social system in which each person is identified with their matriline – their mother's lineage – and which can involve the inheritance ...

family groups, some of which are made up of more than ten members, including three mothers and their dependent offspring, and are led by the

which is often the eldest female. She remains leader of the group until death or if she no longer has the energy for the role;Kingdon, p. 53. a study on zoo elephants showed that when the matriarch died, the levels of faecal

corticosterone Corticosterone, also known as 17-deoxycortisol and 11β,21-dihydroxyprogesterone, is a 21-carbon steroid hormone of the corticosteroid type produced in the cortex of the adrenal glands. It is of minor importance in humans, except in the very rar ...

('stress hormone') dramatically increased in the surviving elephants. When her tenure is over, the matriarch's eldest daughter takes her place; this occurs even if her sister is present. One study found that younger matriarchs are more likely than older ones to under-react to severe danger. Family groups may split after becoming too large for the available resources. The social circle of the female elephant does not necessarily end with the small family unit. In the case of elephants in

Amboseli National Park Amboseli National Park, formerly Maasai Amboseli Game Reserve, is a national park in Kajiado South Constituency in Kajiado County, Kenya. The park is in size at the core of an ecosystem that spreads across the Kenya-Tanzania border. The local ...

, Kenya, a female's life involves interaction with other families, clans, and subpopulations. Families may associate and bond with each other, forming what are known as bond groups which typically made of two family groups. During the dry season, elephant families may cluster together and form another level of social organisation known as the clan. Groups within these clans do not form strong bonds, but they defend their dry-season ranges against other clans. There are typically nine groups in a clan. The Amboseli elephant population is further divided into the "central" and "peripheral" subpopulations.Sukumar, pp. 175–79. Some elephant populations in India and Sri Lanka have similar basic social organisations. There appear to be cohesive family units and loose aggregations. They have been observed to have "nursing units" and "juvenile-care units". In southern India, elephant populations may contain family groups, bond groups and possibly clans. Family groups tend to be small, consisting of one or two adult females and their offspring. A group containing more than two adult females plus offspring is known as a "joint family". Malay elephant populations have even smaller family units and do not have any social organisation higher than a family or bond group. Groups of African forest elephants typically consist of one adult female with one to three offspring. These groups appear to interact with each other, especially at forest clearings. Asian Elephant at Corbett National Park 15

Asian Elephant at Corbett National Park 15

The social life of the adult male is very different. As he matures, a male spends more time at the edge of his group and associates with outside males or even other families. At Amboseli, young males spend over 80% of their time away from their families when they are 14–15. When males permanently leave, they either live alone or with other males. The former is typical of bulls in dense forests. Asian males are usually solitary, but occasionally form groups of two or more individuals; the largest consisted of seven bulls. Larger bull groups consisting of over 10 members occur only among African bush elephants, the largest of which numbered up to 144 individuals. Bulls only return to the herd to breed or to socialize, they do not provide prenatal care to their offspring but rather play a fatherly role to younger bulls to show dominance. Male elephants can be quite sociable when not competing for dominance or mates, and will form long-term relationships. A

dominance hierarchy In biology, a dominance hierarchy (formerly and colloquially called a pecking order) is a type of social hierarchy that arises when members of animal social groups interact, creating a ranking system. A dominant higher-ranking individual is so ...

exists among males, whether they range socially or solitarily. Dominance depends on the age, size and sexual condition, and when in groups, males follow the lead of the dominant bull. Young bulls may seek out the company and leadership of older, more experienced males, whose presence appears to control their aggression and prevent them from exhibiting "deviant" behaviour. Adult males and females come together for reproduction. Bulls associate with family groups if an oestrous cow is present.Sukumar, pp. 179–83.

Sexual behaviour

Musth

Adult males enter a state of increased

known as

. In a population in southern India, males first enter musth at the age of 15, but it is not very intense until they are older than 25. At Amboseli, bulls under 24 do not go into musth, while half of those aged 25–35 and all those over 35 do. Young bulls appear to enter musth during the dry season (January–May), while older bulls go through it during the wet season (June–December). The main characteristic of a bull's musth is a fluid secreted from the

that runs down the side of his face. He may urinate with his penis still in his sheath, which causes the urine to spray on his hind legs. Behaviours associated with musth include walking with the head held high and swinging, picking at the ground with the tusks, marking, rumbling and waving only one ear at a time. This can last from a day to four months. Males become extremely aggressive during musth. Size is the determining factor in

agonistic An agonist is a chemical that activates a receptor to produce a biological response. Receptors are cellular proteins whose activation causes the cell to modify what it is currently doing. In contrast, an antagonist blocks the action of the agon ...

encounters when the individuals have the same condition. In contests between musth and non-musth individuals, musth bulls win the majority of the time, even when the non-musth bull is larger. A male may stop showing signs of musth when he encounters a musth male of higher rank. Those of equal rank tend to avoid each other. Agonistic encounters typically consist of threat displays, chases, and minor sparring with the tusks. Serious fights are rare.Sukumar, pp. 100–08.

Mating

Elephants are polygynous breeders, and copulations are most frequent during the peak of the wet season. A cow in oestrus releases chemical signals (

pheromone A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...

s) in her urine and vaginal secretions to signal her readiness to mate. A bull will follow a potential mate and assess her condition with the

flehmen response The flehmen response (; from German ''flehmen'', to bare the upper teeth, and Upper Saxon German ''flemmen'', to look spiteful), also called the flehmen position, flehmen reaction, flehmen grimace, flehming, or flehmening, is a behavior in which ...

, which requires the male to collect a chemical sample with his trunk and bring it to the

vomeronasal organ The vomeronasal organ (VNO), or Jacobson's organ, is the paired auxiliary olfactory (smell) sense organ located in the soft tissue of the nasal septum, in the nasal cavity just above the roof of the mouth (the hard palate) in various tetrapods. T ...

. The oestrous cycle of a cow lasts 14–16 weeks with a 4–6-week follicular phase and an 8- to 10-week

luteal phase The corpus luteum (Latin for "yellow body"; plural corpora lutea) is a temporary endocrine structure in female ovaries involved in the production of relatively high levels of progesterone, and moderate levels of estradiol, and inhibin A. It is th ...

. While most mammals have one surge of

luteinizing hormone Luteinizing hormone (LH, also known as luteinising hormone, lutropin and sometimes lutrophin) is a hormone produced by gonadotropic cells in the anterior pituitary gland. The production of LH is regulated by gonadotropin-releasing hormone (GnRH) ...

during the follicular phase, elephants have two. The first (or anovulatory) surge, could signal to males that the female is in oestrus by changing her scent, but

ovulation Ovulation is the release of eggs from the ovaries. In women, this event occurs when the ovarian follicles rupture and release the secondary oocyte ovarian cells. After ovulation, during the luteal phase, the egg will be available to be fertilize ...

does not occur until the second (or ovulatory) surge. Fertility rates in cows decline around 45–50 years of age. Bulls engage in a behaviour known as mate-guarding, where they follow oestrous females and defend them from other males. Most mate-guarding is done by musth males, and females actively seek to be guarded by them, particularly older ones. Thus these bulls have more reproductive success. Musth appears to signal to females the condition of the male, as weak or injured males do not have normal musths. For young females, the approach of an older bull can be intimidating, so her relatives stay nearby to provide support and reassurance. During copulation, the male lays his trunk over the female's back. The penis is very mobile, being able to move independently of the pelvis. Before mounting, it curves forward and upward. Copulation lasts about 45 seconds and does not involve

pelvic thrust The pelvic thrust is the thrusting motion of the pelvic region, which is used for a variety of activities, such as dance or sexual activity. Sexual activity The pelvic thrust is used during copulation by many species of mammals, including hum ...

ing or ejaculatory pause. Elephant sperm must swim close to to reach the egg. By comparison, human sperm has to swim around only . Homosexual behaviour is frequent in both sexes. As in heterosexual interactions, this involves mounting. Male elephants sometimes stimulate each other by playfighting and "championships" may form between old bulls and younger males. Female same-sex behaviours have been documented only in captivity where they are known to masturbate one another with their trunks.

Birth and development

Gestation in elephants typically lasts around two years with interbirth intervals usually lasting four to five years. Births tend to take place during the wet season. Calves are born tall and weigh around .Moss, pp. 106–13. Typically, only a single young is born, but twins sometimes occur. The relatively long pregnancy is maintained by five corpus luteums (as opposed to one in most mammals) and gives the foetus more time to develop, particularly the brain and trunk. As such, newborn elephants are

precocial In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...

and quickly stand and walk to follow their mother and family herd. A new calf is usually the centre of attention for herd members. Adults and most of the other young will gather around the newborn, touching and caressing it with their trunks. For the first few days, the mother is intolerant of other herd members near her young.

Alloparenting Alloparenting (also referred to as alloparental care) is a term used to classify any form of parental care provided by an individual towards young that aren't its own direct offspring. These are often referred to as "non-descendant" young, even th ...

– where a calf is cared for by someone other than its mother – takes place in some family groups. Allomothers are typically two to twelve years old. For the first few days, the newborn is unsteady on its feet and needs the support of its mother. It relies on touch, smell, and hearing, as its eyesight is poor. It has little precise control over its trunk, which wiggles around and may cause it to trip. By its second week of life, the calf can walk more firmly and has more control over its trunk. After its first month, a calf can pick up, hold, and put objects in its mouth, but cannot suck water through the trunk and must drink directly through the mouth. It is still dependent on its mother and keeps close to her. For its first three months, a calf relies entirely on milk from its mother for nutrition, after which it begins to forage for vegetation and can use its trunk to collect water. At the same time, improvements in lip and leg coordination occur. Calves continue to suckle at the same rate as before until their sixth month, after which they become more independent when feeding. By nine months, mouth, trunk and foot coordination is perfected. After a year, a calf's abilities to groom, drink, and feed itself are fully developed. It still needs its mother for nutrition and protection from predators for at least another year. Suckling bouts tend to last 2–4 min/hr for a calf younger than a year and it continues to suckle until it reaches three years of age or older. Suckling after two years may serve to maintain growth rate, body condition and reproductive ability.Sukumar, pp. 126–29. Play behaviour in calves differs between the sexes; females run or chase each other while males play-fight. The former are

sexually mature Sexual maturity is the capability of an organism to reproduce. In humans it might be considered synonymous with adulthood, but here puberty is the name for the process of biological sexual maturation, while adulthood is based on cultural definiti ...

by the age of nine years while the latter become mature around 14–15 years. Adulthood starts at about 18 years of age in both sexes. Elephants have long lifespans, reaching 60–70 years of age. Lin Wang, a captive male Asian elephant, lived for 86 years.

Communication

Touching is an important form of communication among elephants. Individuals greet each other by stroking or wrapping their trunks; the latter also occurs during mild competition. Older elephants use trunk-slaps, kicks, and shoves to discipline younger ones. Individuals of any age and sex will touch each other's mouths, temporal glands, and genitals, particularly during meetings or when excited. This allows individuals to pick up chemical cues. Touching is especially important for mother–calf communication. When moving, elephant mothers will touch their calves with their trunks or feet when side-by-side or with their tails if the calf is behind them. If a calf wants to rest, it will press against its mother's front legs and when it wants to suckle, it will touch her breast or leg. Visual displays mostly occur in agonistic situations. Elephants will try to appear more threatening by raising their heads and spreading their ears. They may add to the display by shaking their heads and snapping their ears, as well as throwing dust and vegetation. They are usually bluffing when performing these actions. Excited elephants may raise their trunks. Submissive ones will lower their heads and trunks, as well as flatten their ears against their necks, while those that accept a challenge will position their ears in a V shape. Elephants produce several vocalisations, usually through the larynx, though some may be modified by the trunk.Payne and Langbauer, pp. 120–21. These include trumpets, roars, barks, snorts, growls and rumbles which may be produced for either short or long range communication. Elephants may produce

infrasonic Infrasound, sometimes referred to as low status sound, describes sound waves with a Audio frequency, frequency below the lower limit of human audibility (generally 20 Hertz, Hz). Hearing becomes gradually less sensitive as frequency decreases, ...

rumbles. For Asian elephants, these calls have a frequency of 14–24 Hz, with sound pressure levels of 85–90 dB and last 10–15 seconds. For African elephants, calls range from 15 to 35 Hz with sound pressure levels as high as 117 dB, allowing communication for many kilometres, with a possible maximum range of around . Elephants are known to communicate with seismics, vibrations produced by impacts on the earth's surface or acoustical waves that travel through it. An individual running or mock charging can create seismic signals that can be heard at travel distances of up to . Seismic waveforms produced from predator alarm calls travel .

Intelligence and cognition

Elephants exhibit mirror self-recognition, an indication of self-awareness and cognition that has also been demonstrated in some

ape Apes (collectively Hominoidea ) are a clade of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and as well as Europe in prehistory), which together with its sister g ...

s and

dolphin A dolphin is an aquatic mammal within the infraorder Cetacea. Dolphin species belong to the families Delphinidae (the oceanic dolphins), Platanistidae (the Indian river dolphins), Iniidae (the New World river dolphins), Pontoporiidae (the ...

s. One study of a captive female Asian elephant suggested the animal was capable of learning and distinguishing between several visual and some acoustic discrimination pairs. This individual was even able to score a high accuracy rating when re-tested with the same visual pairs a year later. Elephants are among the species known to use tools. An Asian elephant has been observed modifying branches and using them as flyswatters. Tool modification by these animals is not as advanced as that of chimpanzees. Elephants are popularly thought of as having an excellent memory. This could have a factual basis; they possibly have

cognitive map A cognitive map is a type of mental representation which serves an individual to acquire, code, store, recall, and decode information about the relative locations and attributes of phenomena in their everyday or metaphorical spatial environment. T ...

s to allow them to remember large-scale spaces over long periods of time. Individuals appear to be able to keep track of the current location of their family members. Scientists debate the extent to which elephants feel

emotion Emotions are mental states brought on by neurophysiology, neurophysiological changes, variously associated with thoughts, feelings, behavioral responses, and a degree of pleasure or suffering, displeasure. There is currently no scientific ...

. They appear to show interest in the bones of their own kind, regardless of whether they are related. As with chimpanzees and dolphins, a dying or dead elephant may elicit attention and aid from others, including those from other groups. This has been interpreted as expressing "concern"; however, others would dispute such an interpretation as being anthropomorphic; the ''Oxford Companion to Animal Behaviour'' (1987) advised that "one is well advised to study the behaviour rather than attempting to get at any underlying emotion".

Conservation

Status

African bush elephants were listed as

Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...

by the

(IUCN) in 2021, and African forest elephants were listed as Critically Endangered in the same year. In 1979, Africa had an estimated minimum population of 1.3 million elephants, with a possible upper limit of 3.0 million. By 1989, the population was estimated to be 609,000; with 277,000 in Central Africa, 110,000 in

eastern Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historica ...

, 204,000 in

southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number o ...

, and 19,000 in

western Africa West Africa or Western Africa is the westernmost region of Africa. The United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Mali, Maurita ...

. About 214,000 elephants were estimated to live in the rainforests, fewer than had previously been thought. From 1977 to 1989, elephant populations declined by 74% in East Africa. After 1987, losses in elephant numbers accelerated, and savannah populations from

Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...

Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constituti ...

experienced a decline of 80%. African forest elephants had a total loss of 43%. Population trends in southern Africa were mixed, with anecdotal reports of losses in

Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most central point. Its neighbours are t ...

Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi ...

and

Angola , national_anthem = " Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...

while populations grew in Botswana and Zimbabwe and were stable in South Africa. Conversely, studies in 2005 and 2007 found populations in eastern and southern Africa were increasing by an average annual rate of 4.0%. The IUCN estimated that total population in Africa is estimated at around to 415,000 individuals for both species combined as of 2016. African elephants receive at least some legal protection in every country where they are found, but 70% of their range exists outside protected areas. Successful conservation efforts in certain areas have led to high population densities. As of 2008, local numbers were controlled by contraception or translocation. Large-scale

culling In biology, culling is the process of segregating organisms from a group according to desired or undesired characteristics. In animal breeding, it is the process of removing or segregating animals from a breeding stock based on a specific tr ...

s ceased in 1988 when Zimbabwe abandoned the practice. In 1989, the African elephant was listed under Appendix I by the

Convention on International Trade in Endangered Species of Wild Fauna and Flora CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of intern ...

(CITES), making trade illegal. Appendix II status (which allows restricted trade) was given to elephants in Botswana, Namibia, and Zimbabwe in 1997 and South Africa in 2000. In some countries,

sport hunting Trophy hunting is a form of hunting for sport in which parts of the hunted wild animals are kept and displayed as trophies. The animal being targeted, known as the "game", is typically a mature male specimen from a popular species of collectable ...

of the animals is legal; Botswana, Cameroon,

Gabon Gabon (; ; snq, Ngabu), officially the Gabonese Republic (french: République gabonaise), is a country on the west coast of Central Africa. Located on the equator, it is bordered by Equatorial Guinea to the northwest, Cameroon to the nort ...

, Mozambique, Namibia, South Africa,

Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...

, Zambia, and Zimbabwe have CITES export quotas for elephant trophies. In June 2016, the

First Lady of Kenya The first lady of Kenya is the title held by the wife of the president of Kenya. The country's present first lady is Rachel Ruto, wife of president William Ruto, who took office on 13 September 2022. First ladies of Kenya Daniel arap Moi a ...

Margaret Kenyatta Margaret Gakuo Kenyatta (born 8 April 1964) is a Kenyan educator who served as First Lady of Kenya from 2013 to 2022. She is the wife of Uhuru Kenyatta, the immediate former Kenyan president. Biography Margaret Wanjiru Gakuo was born on 8 Apri ...

, helped launch the East Africa Grass-Root Elephant Education Campaign Walk, organised by elephant conservationist

Jim Nyamu Jim Justus Nyamu, of Nairobi, Kenya, is an elephant research scientist and activist against poaching and trade in ivory. Nyamu is the executive director at the Elephant Neighbors Center (ENC) and is leader of the movement, Ivory Belongs to Elep ...

. The event was conducted to raise awareness of the value of elephants and rhinos, to help mitigate human-elephant conflicts, and to promote anti-poaching activities.Star website
"First Lady launches campaign to save elephants."
4 June 2016. Accessed 5 June 2016. In 2020, the IUCN listed the Asian elephant as

due to an almost 50% population decline over "the last three generations". Asian elephants once ranged from Syria and

Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to the north, Iran to the east, the Persian Gulf and K ...

(the subspecies ''Elephas maximus asurus''), to China (up to the

Yellow River The Yellow River or Huang He (Chinese: , Mandarin: ''Huáng hé'' ) is the second-longest river in China, after the Yangtze River, and the sixth-longest river system in the world at the estimated length of . Originating in the Bayan Ha ...

)Daniel, p. 174. and Java. It is now extinct in these areas, and the current range of Asian elephants is highly fragmented. The total population of Asian elephants is estimated to be around 40,000–50,000, although this may be a loose estimate. Around 60% of the population is in India. Although Asian elephants are declining in numbers overall, particularly in Southeast Asia, the population in the Western Ghats appears to be increasing.

Threats

The

poaching Poaching has been defined as the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set a ...

of elephants for their ivory, meat and hides has been one of the major threats to their existence. Historically, numerous cultures made ornaments and other works of art from elephant ivory, and its use rivalled that of gold.Martin, pp. 202–07 The ivory trade contributed to the African elephant population decline in the late 20th century. This prompted international bans on ivory imports, starting with the United States in June 1989, and followed by bans in other North American countries, western European countries, and Japan. Around the same time, Kenya destroyed all its ivory stocks. CITES approved an international ban on ivory that went into effect in January 1990. Following the bans, unemployment rose in India and China, where the ivory industry was important economically. By contrast, Japan and Hong Kong, which were also part of the industry, were able to adapt and were not badly affected. Zimbabwe, Botswana, Namibia, Zambia, and

Malawi Malawi (; or aláwi Tumbuka: ''Malaŵi''), officially the Republic of Malawi, is a landlocked country in Southeastern Africa that was formerly known as Nyasaland. It is bordered by Zambia to the west, Tanzania to the north and northeas ...

wanted to continue the ivory trade and were allowed to, since their local elephant populations were healthy, but only if their supplies were from elephants that had been culled or died of natural causes. The ban allowed the elephant to recover in parts of Africa. In January 2012, 650 elephants in Bouba Njida National Park, Cameroon, were killed by Chadian raiders. This has been called "one of the worst concentrated killings" since the ivory ban. Asian elephants are potentially less vulnerable to the ivory trade, as females usually lack tusks. Still, members of the species have been killed for their ivory in some areas, such as

Periyar National Park Periyar National Park and Wildlife Sanctuary (PNP) is a protected area located in the districts of Idukki and Pathanamthitta in Kerala, India. It is notable as an elephant reserve and a tiger reserve. The protected area encompasses , of which ...

in India. China was the biggest market for poached ivory but announced they would phase out the legal domestic manufacture and sale of ivory products in May 2015, and in September 2015, China and the United States said "they would enact a nearly complete ban on the import and export of ivory" due to causes of extinction. Other threats to elephants include

and fragmentation. The Asian elephant lives in areas with some of the highest human populations and may be confined to small islands of forest among human-dominated landscapes. Elephants commonly trample and consume crops, which contributes to conflicts with humans, and both elephants and humans have died by the hundreds as a result. Mitigating these conflicts is important for conservation. One proposed solution is the protection of

wildlife corridor A wildlife corridor, habitat corridor, or green corridor is an area of habitat connecting wildlife populations separated by human activities or structures (such as roads, development, or logging). This allows an exchange of individuals between ...

s which gave the animals greater space and maintain the long term viability of large populations.

Association with humans

Working animal

Elephants have been

s since at least the Indus Valley civilization and continue to be used in modern times. There were 13,000–16,500 working elephants employed in Asia in 2000. These animals are typically captured from the wild when they are 10–20 years old when they can be trained quickly and easily, and will have a longer working life. They were traditionally captured with traps and lassos, but since 1950, tranquillisers have been used. Individuals of the Asian species have been often trained as working animals. Asian elephants perform tasks such as hauling loads into remote areas, moving logs to rivers and roads, transporting tourists around national parks, pulling wagons, and leading religious processions. In northern

Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...

, the animals are used to digest coffee beans for

Black Ivory coffee Black Ivory Coffee is a brand of coffee produced by the Black Ivory Coffee Company Ltd in northern Thailand from Arabica coffee beans consumed by elephants and collected from their waste. The taste of Black Ivory coffee is influenced by elephants ...

. They are valued over mechanised tools because they can work in relatively deep water, require relatively little maintenance, need only vegetation and water as fuel and can be trained to memorise specific tasks. Elephants can be trained to respond to over 30 commands.McNeely, pp. 149–50. Musth bulls can be difficult and dangerous to work with and are chained and semi-starved until the condition passes. In India, many working elephants are alleged to have been subject to abuse. They and other captive elephants are thus protected under The Prevention of Cruelty to Animals Act of 1960. In both Myanmar and Thailand,

deforestation Deforestation or forest clearance is the removal of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. The most concentrated ...

and other economic factors have resulted in sizable populations of unemployed elephants resulting in health problems for the elephants themselves as well as economic and safety problems for the people amongst whom they live. The practice of working elephants has also been attempted in Africa. The taming of African elephants in the

Belgian Congo The Belgian Congo (french: Congo belge, ; nl, Belgisch-Congo) was a Belgian colony in Central Africa from 1908 until independence in 1960. The former colony adopted its present name, the Democratic Republic of the Congo (DRC), in 1964. Colo ...

began by decree of

Leopold II of Belgium * german: link=no, Leopold Ludwig Philipp Maria Viktor , house = Saxe-Coburg and Gotha , father = Leopold I of Belgium , mother = Louise of Orléans , birth_date = , birth_place = Brussels, Belgium , death_date = ...

during the 19th century and continues to the present with the Api Elephant Domestication Centre.

Warfare

Historically, elephants were considered formidable instruments of war. They were equipped with armour to protect their sides, and their tusks were given sharp points of iron or brass if they were large enough. War elephants were trained to grasp an enemy soldier and toss him to the person riding on them or to pin the soldier to the ground and impale him. One of the earliest references to war elephants is in the Indian epic ''

Mahabharata The ''Mahābhārata'' ( ; sa, महाभारतम्, ', ) is one of the two major Sanskrit literature, Sanskrit Indian epic poetry, epics of ancient India in Hinduism, the other being the ''Ramayana, Rāmāyaṇa''. It narrates the s ...

'' (written in the 4th century BC, but said to describe events between the 11th and 8th centuries BC). They were not used as much as horse-drawn chariots by either the

Pandava The Pandavas (Sanskrit: पाण्डव, IAST: Pāṇḍava) refers to the five legendary brothers— Yudhishthira, Bhima, Arjuna, Nakula and Sahadeva—who are the central characters of the Hindu epic ''Mahabharata''. They are acknowledge ...

s or

Kaurava ''Kaurava'' is a Sanskrit term which refers to descendants of Kuru, a legendary king of India who is the ancestor of many of the characters of the epic ''Mahabharata''. Usually, the term is used for the 100 sons of King Dhritarashtra and his wi ...

s. During the

Magadha Kingdom Magadha was a region and one of the sixteen sa, script=Latn, Mahajanapadas, label=none, lit=Great Kingdoms of the Second Urbanization (600–200 BCE) in what is now south Bihar (before expansion) at the eastern Ganges Plain. Magadha was ruled ...

(which began in the 6th century BC), elephants began to achieve greater cultural importance than horses, and later Indian kingdoms used war elephants extensively; 3,000 of them were used in the

Nandas The Nanda dynasty ruled in the northern part of the Indian subcontinent during the fourth century BCE, and possibly during the fifth century BCE. The Nandas overthrew the Shaishunaga dynasty in the Magadha region of eastern India, and expanded ...

(5th and 4th centuries BC) army while 9,000 may have been used in the Mauryan army (between the 4th and 2nd centuries BC). The '' Arthashastra'' (written around 300 BC) advised the Mauryan government to reserve some forests for wild elephants for use in the army, and to execute anyone who killed them. From South Asia, the use of elephants in warfare spread west to

Persia Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...

and east to Southeast Asia. The Persians used them during the Achaemenid Empire (between the 6th and 4th centuries BC) while Southeast Asian states first used war elephants possibly as early as the 5th century BC and continued to the 20th century. In his 326 B.C. Indian campaign,

Alexander the Great Alexander III of Macedon ( grc, Ἀλέξανδρος, Alexandros; 20/21 July 356 BC – 10/11 June 323 BC), commonly known as Alexander the Great, was a king of the ancient Greek kingdom of Macedon. He succeeded his father Philip II to ...

confronted elephants for the first time and suffered heavy casualties. Among the reasons for the refusal of the rank-and-file Macedonian soldiers to continue the Indian conquest were rumors of even larger elephant armies in India. Alexander trained his foot soldiers to injure the animals and cause them to panic during wars with both the Persians and Indians.

Ptolemy Claudius Ptolemy (; grc-gre, Πτολεμαῖος, ; la, Claudius Ptolemaeus; AD) was a mathematician, astronomer, astrologer, geographer, and music theorist, who wrote about a dozen scientific treatises, three of which were of importance ...

, who was one of Alexander's generals, used corps of Asian elephants during his reign as the ruler of Egypt (which began in 323 BC). His son and successor Ptolemy II (who began his rule in 285 BC) obtained his supply of elephants further south in

Nubia Nubia () (Nobiin: Nobīn, ) is a region along the Nile river encompassing the area between the first cataract of the Nile (just south of Aswan in southern Egypt) and the confluence of the Blue and White Niles (in Khartoum in central Sudan), or ...

. From then on, war elephants were employed in the Mediterranean and North Africa throughout the classical period. The Greek king Pyrrhus used elephants in his attempted invasion of Rome in 280 BC. While they frightened the Roman horses, they were not decisive and Pyrrhus ultimately lost the battle. The Carthaginian general Hannibal took elephants across the

Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Swi ...

during his war with the Romans and reached the Po Valley in 217 BC with all of them alive, but they later succumbed to disease.Wylie (2000), pp. 146–48. Overall, elephants owed their initial successes to the element of surprise and to the fear that their great size invoked. With time, strategists devised counter-measures and war elephants turned into an expensive liability and were hardly ever used by Romans and Parthians.

Zoos and circuses

Flickr - …trialsanderrors - Terrific flights over ponderous elephants, poster for Forepaugh ^ Sells Brothers, ca

Elephants were historically kept for display in the menageries of Ancient Egypt, China,

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, and Ancient Rome, Rome. The Romans in particular pitted them against humans and other animals in gladiator events. In the Modern history, modern era, elephants have traditionally been a major part of zoos and

es around the world. In circuses, they are trained to perform tricks. The most famous circus elephant was probably Jumbo (1861 – 15 September 1885), who was a major attraction in the Ringling Bros. and Barnum & Bailey Circus, Barnum & Bailey Circus. These animals do not reproduce well in captivity, due to the difficulty of handling musth bulls and limited understanding of female oestrous cycles. Asian elephants were always more common than their African counterparts in modern zoos and circuses. After CITES listed the Asian elephant under Appendix I in 1975, the number of African elephants in zoos increased in the 1980s, although the import of Asians continued. Subsequently, the US received many of its captive African elephants from Zimbabwe, which had an overabundance of the animals.Tuttle, pp. 184–88. Keeping elephants in zoos has met with some controversy. Proponents of zoos argue that they offer researchers easy access to the animals and provide money and expertise for preserving their natural habitats, as well as safekeeping for the species. Critics claim that the animals in zoos are under physical and mental stress. Elephants have been recorded displaying Stereotypy (non-human), stereotypical behaviours in the form of swaying back and forth, trunk swaying, or route tracing. This has been observed in 54% of individuals in UK zoos. Elephants in European zoos appear to have shorter lifespans than their wild counterparts at only 17 years, although other studies suggest that zoo elephants live as long those in the wild. The use of elephants in circuses has also been controversial; the Humane Society of the United States has accused circuses of mistreating and distressing their animals. In testimony to a US federal court in 2009, Barnum & Bailey Circus CEO Kenneth Jeffrey Feld, Kenneth Feld acknowledged that circus elephants are struck behind their ears, under their chins and on their legs with metal-tipped prods, called Elephant goad, bull hooks or ankus. Feld stated that these practices are necessary to protect circus workers and acknowledged that an elephant trainer was reprimanded for using an electric shock device, known as a hot shot or electric prod, on an elephant. Despite this, he denied that any of these practices harm elephants. Some trainers have tried to train elephants without the use of physical punishment. Ralph Helfer is known to have relied on gentleness and reward when training his animals, including elephants and lions. Ringling Bros. and Barnum and Bailey circus retired its touring elephants in May 2016.

Attacks

Elephants can exhibit bouts of aggressive behaviour and engage in destructive actions against humans. In Africa, groups of adolescent elephants damaged homes in villages after cullings in the 1970s and 1980s. Because of the timing, these attacks have been interpreted as vindictive. In parts of India, male elephants regularly enter villages at night, destroying homes and killing people. Elephants killed around 300 people between 2000 and 2004 in Jharkhand while in Assam, 239 people were reportedly killed between 2001 and 2006. Local people have reported their belief that some elephants were drunk during their attacks, although officials have disputed this explanation. Purportedly drunk elephants attacked an Indian village a second time in December 2002, killing six people, which led to the killing of about 200 elephants by locals.

Cultural depictions

In many cultures, elephants represent strength, power, wisdom, longevity, stamina, leadership, sociability, nurturance and loyalty. Several cultural references emphasise the elephant's size and exotic uniqueness. For instance, a "white elephant" is a byword for something expensive, useless, and bizarre. The expression "elephant in the room" refers to an obvious truth that is ignored or otherwise unaddressed. The story of the blind men and an elephant teaches that reality can be observed from different perspectives. Elephants have been represented in art since Paleolithic times. Africa, in particular, contains many rock paintings and engravings of the animals, especially in the Sahara and southern Africa. In Asia, the animals are depicted as Motif (visual arts), motifs in Hinduism, Hindu and Buddhism, Buddhist shrines and temples. Elephants were often difficult to portray by people with no first-hand experience of them. The Ancient Rome, ancient Romans, who kept the animals in captivity, depicted anatomically accurate elephants on mosaics in Tunisia and Sicily. At the beginning of the Middle Ages, when Europeans had little to no access to the animals, elephants were portrayed more like fantasy creatures. They were often depicted with horse- or bovine-like bodies with trumpet-like trunks and tusks like a boar; some were even given hooves. Elephants were commonly featured in motifs by the stonemasons of the Gothic architecture, Gothic churches. As more elephants began to be sent to European kings as gifts during the 15th century, depictions of them became more accurate, including one made by Leonardo da Vinci. Despite this, some Europeans continued to portray them in a more stylised fashion. Max Ernst's 1921 Surrealism, surrealist painting, ''The Elephant Celebes,'' depicts an elephant as a silo with a trunk-like hose protruding from it. Illustration at p

Elephants have been the subject of religious beliefs. The Mbuti people of central Africa believe that the souls of their dead ancestors resided in elephants.McNeely, pp. 158–60. Similar ideas existed among other African societies, who believed that their chiefs would be reincarnation, reincarnated as elephants. During the 10th century AD, the people of Igbo-Ukwu, near the Niger Delta, buried their leaders with elephant tusks. The animals' religious importance is only Totemism, totemic in Africa but is much more significant in Asia. In Sumatra, elephants have been associated with lightning. Likewise in Hinduism, they are linked with thunderstorms as Airavata, the father of all elephants, represents both lightning and rainbows. One of the most important Hindu deities, the elephant-headed Ganesha, is ranked equal with the supreme gods Shiva, Vishnu, and Brahma. Ganesha is associated with writers and merchants and it is believed that he can give people success as well as grant them their desires. In Buddhism, Gautama Buddha, Buddha is said to have been a White elephant (animal), white elephant reincarnated as a human. In Islamic tradition, the year 570 when Muhammad was born is known as the Year of the Elephant. Elephants were thought to be religious themselves by the Romans, who believed that they worshipped the sun and stars. Elephants are ubiquitous in Western

as emblems of the exotic, especially since – as with the giraffe, hippopotamus and

– there are no similar animals familiar to Western audiences. The use of the elephant as a symbol of the Republican Party (United States), U.S. Republican Party began with an Republican elephant, 1874 cartoon by Thomas Nast. As characters, elephants are most common in children's stories, in which they are generally cast as models of exemplary behaviour. They are typically surrogates for humans with ideal human values. Many stories tell of isolated young elephants returning to a close-knit community, such as "The Elephant's Child" from Rudyard Kipling's ''Just So Stories'', The Walt Disney Company, Disney's ''Dumbo,'' and Kathryn and Byron Jackson's ''The Saggy Baggy Elephant''. Other elephant heroes Anthropomorphism, given human qualities include Jean de Brunhoff's Babar the Elephant, Babar, David McKee's Elmer the Patchwork Elephant, Elmer, and Dr. Seuss's Horton the Elephant, Horton.

References

Bibliography

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External links

International Elephant Foundation
{{Featured article Elephants, Herbivorous mammals Tool-using mammals Articles containing video clips Mammal common names Pliocene first appearances

Etymology

Taxonomy

Evolution and extinct relatives

Anatomy

Size

Bones

Head

Ears

Trunk

Teeth

Tusks

Skin

Legs, locomotion, and posture

Organs

Body temperature

Behaviour and life history

Ecology and activities

Social organisation

Sexual behaviour

Musth

Mating

Birth and development

Communication

Intelligence and cognition

Conservation

Status

Threats

Association with humans

Working animal

Warfare

Zoos and circuses

Attacks

Cultural depictions

See also

References

Bibliography

Further reading

External links