Desert Kingdom

A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About one-third of the land surface of the Earth is arid or semi-arid. This includes much of the polar regions, where little precipitation occurs, and which are sometimes called polar deserts or "cold deserts". Deserts can be classified by the amount of precipitation that falls, by the temperature that prevails, by the causes of desertification or by their geographical location.

Deserts are formed by weathering processes as large variations in temperature between day and night put strains on the rocks, which consequently break in pieces. Although rain seldom occurs in deserts, there are occasional downpours that can result in flash floods. Rain falling on hot rocks can cause them to shatter, and the resulting fragments and rubble strewn over the desert floor are further eroded by the wind. This picks up particles of sand and dust, which can remain airborne for extended periods – sometimes causing the formation of sand storms or dust storms. Wind-blown sand grains striking any solid object in their path can abrade the surface. Rocks are smoothed down, and the wind sorts sand into uniform deposits. The grains end up as level sheets of sand or are piled high in billowing sand dunes. Other deserts are flat, stony plains where all the fine material has been blown away and the surface consists of a mosaic of smooth stones. These areas are known as desert pavements, and little further erosion takes place. Other desert features include rock outcrops, exposed bedrock and clays once deposited by flowing water. Temporary lakes may form and salt pans may be left when waters evaporate. There may be underground sources of water, in the form of springs and seepages from aquifers. Where these are found, oases can occur.

Plants and animals living in the desert need special adaptations to survive in the harsh environment. Plants tend to be tough and wiry with small or no leaves, water-resistant cuticles, and often spines to deter herbivory. Some annual plants germinate, bloom and die in the course of a few weeks after rainfall, while other long-lived plants survive for years and have deep root systems able to tap underground moisture. Animals need to keep cool and find enough food and water to survive. Many are nocturnal, and stay in the shade or underground during the heat of the day. They tend to be efficient at conserving water, extracting most of their needs from their food and concentrating their urine. Some animals remain in a state of dormancy for long periods, ready to become active again during the rare rainfall. They then reproduce rapidly while conditions are favorable before returning to dormancy.

People have struggled to live in deserts and the surrounding semi-arid lands for millennia. Nomads have moved their flocks and herds to wherever grazing is available, and oases have provided opportunities for a more settled way of life. The cultivation of semi-arid regions encourages erosion of soil and is one of the causes of increased desertification. Desert farming is possible with the aid of irrigation, and the Imperial Valley in California provides an example of how previously barren land can be made productive by the import of water from an outside source. Many trade routes have been forged across deserts, especially across the Sahara, and traditionally were used by caravans of camels carrying salt, gold, ivory and other goods. Large numbers of slaves were also taken northwards across the Sahara. Some mineral extraction also takes place in deserts, and the uninterrupted sunlight gives potential for the capture of large quantities of solar energy.

Etymology

English ''desert'' and its Romance cognates (including Italian and Portuguese '' deserto'', French '' désert'' and Spanish '' desierto'') all come from the ecclesiastical Latin '' dēsertum'' (originally "an abandoned place"), a participle of '' dēserere'', "to abandon". The correlation between aridity and sparse population is complex and dynamic, varying by culture, era, and technologies; thus the use of the word ''desert'' can cause confusion. In English before the 20th century, ''desert'' was often used in the sense of "unpopulated area", without specific reference to aridity; but today the word is most often used in its climate-science sense (an area of low precipitation). Phrases such as " desert island" and " Great American Desert", or Shakespeare's "deserts of Bohemia" ('' The Winter's Tale'') in previous centuries did not necessarily imply sand or aridity; their focus was the sparse population.

Major deserts

Deserts take up about one third of Earth's land surface. Bottomlands may be salt-covered flats. Eolian processes are major factors in shaping desert landscapes. Polar deserts (also seen as "cold deserts") have similar features, except the main form of precipitation is snow rather than rain. Antarctica is the world's largest cold desert (composed of about 98% thick continental ice sheet and 2% barren rock). Some of the barren rock is to be found in the so-called Dry Valleys of Antarctica that almost never get snow, which can have ice-encrusted saline lakes that suggest evaporation far greater than the rare snowfall due to the strong katabatic winds that even evaporate ice.

Deserts, both hot and cold, play a part in moderating Earth's temperature. This is because they reflect more of the incoming light and their albedo is higher than that of forests or the sea.

Defining characteristics

A desert is a region of land that is very dry because it receives low amounts of precipitation (usually in the form of rain, but it may be snow, mist or fog), often has little coverage by plants, and in which streams dry up unless they are supplied by water from outside the area. Deserts generally receive less than of precipitation each year. The potential evapotranspiration may be large but (in the absence of available water) the actual evapotranspiration may be close to zero. Semi-deserts are regions which receive between and when clad in grass, these are known as steppes.

Water

One of the driest places on Earth is the Atacama Desert. (Excerpt) It is virtually devoid of life because it is blocked from receiving precipitation by the Andes mountains to the east and the Chilean Coast Range to the west. The cold Humboldt Current and the anticyclone of the Pacific are essential to keep the dry climate of the Atacama. The average precipitation in the Chilean region of Antofagasta is just per year. Some weather stations in the Atacama have never received rain. Evidence suggests that the Atacama may not have had any significant rainfall from 1570 to 1971. It is so arid that mountains that reach as high as are completely free of glaciers and, in the southern part from 25°S to 27°S, may have been glacier-free throughout the Quaternary, though permafrost extends down to an altitude of and is continuous above . Nevertheless, there is some plant life in the Atacama, in the form of specialist plants that obtain moisture from dew and the fogs that blow in from the Pacific.

When rain falls in deserts, as it occasionally does, it is often with great violence. The desert surface is evidence of this with dry stream channels known as arroyos or wadis meandering across its surface. These can experience flash floods, becoming raging torrents with surprising rapidity after a storm that may be many kilometers away. Most deserts are in basins with no drainage to the sea but some are crossed by exotic rivers sourced in mountain ranges or other high rainfall areas beyond their borders. The River Nile, the Colorado River and the Yellow River do this, losing much of their water through evaporation as they pass through the desert and raising groundwater levels nearby. There may also be underground sources of water in deserts in the form of springs, aquifers, underground rivers or lakes. Where these lie close to the surface, wells can be dug and oases may form where plant and animal life can flourish. The Nubian Sandstone Aquifer System under the Sahara Desert is the largest known accumulation of fossil water. The Great Man-Made River is a scheme launched by Libya's Muammar Gadaffi to tap this aquifer and supply water to coastal cities. Kharga Oasis in Egypt is long and is the largest oasis in the Libyan Desert. A lake occupied this depression in ancient times and thick deposits of sandy-clay resulted. Wells are dug to extract water from the porous sandstone that lies underneath. Seepages may occur in the walls of canyons and pools may survive in deep shade near the dried up watercourse below.

Lakes may form in basins where there is sufficient precipitation or meltwater from glaciers above. They are usually shallow and saline, and wind blowing over their surface can cause stress, moving the water over nearby low-lying areas. When the lakes dry up, they leave a crust or hardpan behind. This area of deposited clay, silt or sand is known as a playa. The deserts of North America have more than one hundred playas, many of them relics of Lake Bonneville which covered parts of Utah, Nevada and Idaho during the last ice age when the climate was colder and wetter. These include the Great Salt Lake, Utah Lake, Sevier Lake and many dry lake beds. The smooth flat surfaces of playas have been used for attempted vehicle speed records at Black Rock Desert and Bonneville Speedway and the United States Air Force uses Rogers Dry Lake in the Mojave Desert as runways for aircraft and the Space Shuttle.

Classification

Deserts have been defined and classified in a number of ways, generally combining total precipitation, number of days on which this falls, temperature, and humidity, and sometimes additional factors. For example, Phoenix, Arizona, receives less than of precipitation per year, and is immediately recognized as being located in a desert because of its aridity-adapted plants. The North Slope of Alaska's Brooks Range also receives less than of precipitation per year and is often classified as a cold desert. Other regions of the world have cold deserts, including areas of the Himalayas and other high-altitude areas in other parts of the world. Polar deserts cover much of the ice-free areas of the Arctic and Antarctic. A non-technical definition is that deserts are those parts of Earth's surface that have insufficient vegetation cover to support a human population.

Potential evapotranspiration supplements the measurement of precipitation in providing a scientific measurement-based definition of a desert. The water budget of an area can be calculated using the formula ''P'' − ''PE'' ± ''S'', wherein ''P'' is precipitation, ''PE'' is potential evapotranspiration rates and ''S'' is the amount of surface storage of water. Evapotranspiration is the combination of water loss through atmospheric evaporation and through the life processes of plants. Potential evapotranspiration, then, is the amount of water that ''could'' evaporate in any given region. As an example, Tucson, Arizona receives about of rain per year, however about of water could evaporate over the course of a year. In other words, about eight times more water could evaporate from the region than actually falls as rain. Rates of evapotranspiration in cold regions such as Alaska are much lower because of the lack of heat to aid in the evaporation process.

Deserts are sometimes classified as "hot" or "cold", "semiarid" or "coastal". The characteristics of hot deserts include high temperatures in summer; greater evaporation than precipitation, usually exacerbated by high temperatures, strong winds and lack of cloud cover; considerable variation in the occurrence of precipitation, its intensity and distribution; and low humidity. Winter temperatures vary considerably between different deserts and are often related to the location of the desert on the continental landmass and the latitude. Daily variations in temperature can be as great as or more, with heat loss by radiation at night being increased by the clear skies.

Cold deserts, sometimes known as temperate deserts, occur at higher latitudes than hot deserts, and the aridity is caused by the dryness of the air. Some cold deserts are far from the ocean and others are separated by mountain ranges from the sea, and in both cases, there is insufficient moisture in the air to cause much precipitation. The largest of these deserts are found in Central Asia. Others occur on the eastern side of the Rocky Mountains, the eastern side of the southern Andes and in southern Australia. Polar deserts are a particular class of cold desert. The air is very cold and carries little moisture so little precipitation occurs and what does fall, usually snow, is carried along in the often strong wind and may form blizzards, drifts and dunes similar to those caused by dust and sand in other desert regions. In Antarctica, for example, the annual precipitation is about on the central plateau and some ten times that amount on some major peninsulas.

Based on precipitation alone, hyperarid deserts receive less than of rainfall a year; they have no annual seasonal cycle of precipitation and experience twelve-month periods with no rainfall at all. Arid deserts receive between in a year and semiarid deserts between . However, such factors as the temperature, humidity, rate of evaporation and evapotranspiration, and the moisture storage capacity of the ground have a marked effect on the degree of aridity and the plant and animal life that can be sustained. Rain falling in the cold season may be more effective at promoting plant growth, and defining the boundaries of deserts and the semiarid regions that surround them on the grounds of precipitation alone is problematic.

A semi-arid desert or a steppe is a version of the arid desert with much more rainfall, vegetation and higher humidity. These regions feature a semi-arid climate and are less extreme than regular deserts. Like arid deserts, temperatures can vary greatly in semi deserts. They share some characteristics of a true desert and are usually located at the edge of deserts and continental dry areas. They usually receive precipitation from but this can vary due to evapotranspiration and soil nutrition. Semi deserts can be found in the Tabernas Desert (and some of the Spanish Plateau), The Sahel, The Eurasian Steppe, most of Central Asia, the Western US, most of Northern Mexico, portions of South America (especially in Argentina) and the Australian Outback. They usually feature ''BSh'' (hot steppe) or ''BSk'' (temperate steppe) in the Köppen climate classification.

Coastal deserts are mostly found on the western edges of continental land masses in regions where cold currents approach the land or cold water upwellings rise from the ocean depths. The cool winds crossing this water pick up little moisture and the coastal regions have low temperatures and very low rainfall, the main precipitation being in the form of fog and dew. The range of temperatures on a daily and annual scale is relatively low, being and respectively in the Atacama Desert. Deserts of this type are often long and narrow and bounded to the east by mountain ranges. They occur in Namibia, Chile, southern California and Baja California. Other coastal deserts influenced by cold currents are found in Western Australia, the Arabian Peninsula and Horn of Africa, and the western fringes of the Sahara.

In 1961, Peveril Meigs divided desert regions on Earth into three categories according to the amount of precipitation they received. In this now widely accepted system, extremely arid lands have at least twelve consecutive months without precipitation, arid lands have less than of annual precipitation, and semiarid lands have a mean annual precipitation of between . Both extremely arid and arid lands are considered to be deserts while semiarid lands are generally referred to as steppes when they are grasslands.

Deserts are also classified, according to their geographical location and dominant weather pattern, as trade wind, mid-latitude, rain shadow, coastal, monsoon, or polar deserts. Trade wind deserts occur either side of the horse latitudes at 30° to 35° North and South. These belts are associated with the subtropical anticyclone and the large-scale descent of dry air moving from high-altitudes toward the poles. The Sahara Desert is of this type. Mid-latitude deserts occur between 30° and 50° North and South. They are mostly in areas remote from the sea where most of the moisture has already precipitated from the prevailing winds. They include the Tengger and Sonoran Deserts. Monsoon deserts are similar. They occur in regions where large temperature differences occur between sea and land. Moist warm air rises over the land, deposits its water content and circulates back to sea. Further inland, areas receive very little precipitation. The Thar Desert near the India/Pakistan border is of this type.

In some parts of the world, deserts are created by a rain shadow effect. Orographic lift occurs as air masses rise to pass over high ground. In the process they cool and lose much of their moisture by precipitation on the windward slope of the mountain range. When they descend on the leeward side, they warm and their capacity to hold moisture increases so an area with relatively little precipitation occurs. The Taklamakan Desert is an example, lying in the rain shadow of the Himalayas and receiving less than precipitation annually. Other areas are arid by virtue of being a very long way from the nearest available sources of moisture.

Montane deserts are arid places with a very high altitude; the most prominent example is found north of the Himalayas, in the Kunlun Mountains and the Tibetan Plateau. Many locations within this category have elevations exceeding and the thermal regime can be hemiboreal. These places owe their profound aridity (the average annual precipitation is often less than 40 mm or 1.5 in) to being very far from the nearest available sources of moisture and are often in the lee of mountain ranges. Montane deserts are normally cold, or may be scorchingly hot by day and very cold by night as is true of the northeastern slopes of Mount Kilimanjaro.

Polar deserts such as McMurdo Dry Valleys remain ice-free because of the dry katabatic winds that flow downhill from the surrounding mountains. Former desert areas presently in non-arid environments, such as the Sandhills in Nebraska, are known as paleodeserts. In the Köppen climate classification system, deserts are classed as ''BWh'' (hot desert) or ''BWk'' (temperate desert). In the Thornthwaite climate classification system, deserts would be classified as arid megathermal climates.

Polar desert

Polar deserts are a type of cold desert. While they do not lack water, having a persistent cover of snow and ice, this is merely due to marginal evaporation rates and low precipitation.

Biological desert

The concept of "biological desert" redefines the concept of desert, without the characteristic of aridity, not lacking water, but instead lacking life. Such places can be so-called "ocean deserts", which are mostly at the centers of gyres, but also hypoxic or anoxic waters such as dead zones.

Morphology

Weathering processes

Deserts usually have a large diurnal and seasonal temperature range, with high daytime temperatures falling sharply at night. The diurnal range may be as much as and the rock surface experiences even greater temperature differentials. During the day the sky is usually clear and most of the sun's radiation reaches the ground, but as soon as the sun sets, the desert cools quickly by radiating heat into space. In hot deserts, the temperature during daytime can exceed in summer and plunge below freezing point at night during winter.George, 1978. p. 11

Such large temperature variations have a destructive effect on the exposed rocky surfaces. The repeated fluctuations put a strain on exposed rock and the flanks of mountains crack and shatter. Fragmented strata slide down into the valleys where they continue to break into pieces due to the relentless sun by day and chill by night. Successive strata are exposed to further weathering. The relief of the internal pressure that has built up in rocks that have been underground for aeons can cause them to shatter.George, 1978. p. 21 Exfoliation also occurs when the outer surfaces of rocks split off in flat flakes. This is believed to be caused by the stresses put on the rock by repeated thermal expansions and contractions which induces fracturing parallel to the original surface. Chemical weathering processes probably play a more important role in deserts than was previously thought. The necessary moisture may be present in the form of dew or mist. Ground water may be drawn to the surface by evaporation and the formation of salt crystals may dislodge rock particles as sand or disintegrate rocks by exfoliation. Shallow caves are sometimes formed at the base of cliffs by this means.

As the desert mountains decay, large areas of shattered rock and rubble occur. The process continues and the end products are either dust or sand. Dust is formed from solidified clay or volcanic deposits whereas sand results from the fragmentation of harder granites, limestone and sandstone.George, 1978. p. 22 There is a certain critical size (about 0.5 mm) below which further temperature-induced weathering of rocks does not occur and this provides a minimum size for sand grains.

As the mountains are eroded, more and more sand is created. At high wind speeds, sand grains are picked up off the surface and blown along, a process known as saltation. The whirling airborne grains act as a sand blasting mechanism which grinds away solid objects in its path as the kinetic energy of the wind is transferred to the ground.Pye & Tsoar, 2009. p. 4 The sand eventually ends up deposited in level areas known as sand-fields or sand-seas, or piled up in dunes.Pye & Tsoar, 2009. p. 141

Features

Many people think of deserts as consisting of extensive areas of billowing sand dunes because that is the way they are often depicted on TV and in films, but deserts do not always look like this. Across the world, around 20% of desert is sand, varying from only 2% in North America to 30% in Australia and over 45% in Central Asia. Where sand does occur, it is usually in large quantities in the form of sand sheets or extensive areas of dunes.

A sand sheet is a near-level, firm expanse of partially consolidated particles in a layer that varies from a few centimeters to a few meters thick. The structure of the sheet consists of thin horizontal layers of coarse silt and very fine to medium grain sand, separated by layers of coarse sand and pea-gravel which are a single grain thick. These larger particles anchor the other particles in place and may also be packed together on the surface so as to form a miniature desert pavement. Small ripples form on the sand sheet when the wind exceeds . They form perpendicular to the wind direction and gradually move across the surface as the wind continues to blow. The distance between their crests corresponds to the average length of jumps made by particles during saltation. The ripples are ephemeral and a change in wind direction causes them to reorganise.

Sand dunes are accumulations of windblown sand piled up in mounds or ridges. They form downwind of copious sources of dry, loose sand and occur when topographic and climatic conditions cause airborne particles to settle. As the wind blows, saltation and creep take place on the windward side of the dune and individual grains of sand move uphill. When they reach the crest, they cascade down the far side. The upwind slope typically has a gradient of 10° to 20° while the lee slope is around 32°, the angle at which loose dry sand will slip. As this wind-induced movement of sand grains takes place, the dune moves slowly across the surface of the ground. Dunes are sometimes solitary, but they are more often grouped together in dune fields. When these are extensive, they are known as sand seas or ergs.

The shape of the dune depends on the characteristics of the prevailing wind. Barchan dunes are produced by strong winds blowing across a level surface and are crescent-shaped with the concave side away from the wind. When there are two directions from which winds regularly blow, a series of long, linear dunes known as seif dunes may form. These also occur parallel to a strong wind that blows in one general direction. Transverse dunes run at a right angle to the prevailing wind direction. Star dunes are formed by variable winds, and have several ridges and slip faces radiating from a central point. They tend to grow vertically; they can reach a height of , making them the tallest type of dune. Rounded mounds of sand without a slip face are the rare dome dunes, found on the upwind edges of sand seas.

In deserts where large amounts of limestone mountains surround a closed basin, such as at White Sands National Park in south-central New Mexico, occasional storm runoff transports dissolved limestone and gypsum into a low-lying pan within the basin where the water evaporates, depositing the gypsum and forming crystals known as selenite. The crystals left behind by this process are eroded by the wind and deposited as vast white dune fields that resemble snow-covered landscapes. These types of dune are rare, and only form in closed arid basins that retain the highly soluble gypsum that would otherwise be washed into the sea.

A large part of the surface area of the world's deserts consists of flat, stone-covered plains dominated by wind erosion. In " eolian deflation", the wind continually removes fine-grained material, which becomes wind-blown sand. This exposes coarser-grained material, mainly pebbles with some larger stones or cobbles, leaving a desert pavement, an area of land overlaid by closely packed smooth stones forming a tessellated mosaic. Different theories exist as to how exactly the pavement is formed. It may be that after the sand and dust is blown away by the wind the stones jiggle themselves into place; alternatively, stones previously below ground may in some way work themselves to the surface. Very little further erosion takes place after the formation of a pavement, and the ground becomes stable. Evaporation brings moisture to the surface by capillary action and calcium salts may be precipitated, binding particles together to form a desert conglomerate. In time, bacteria that live on the surface of the stones accumulate a film of minerals and clay particles, forming a shiny brown coating known as desert varnish.

Other non-sandy deserts consist of exposed outcrops of bedrock, dry soils or aridisols, and a variety of landforms affected by flowing water, such as alluvial fans, sinks or playas, temporary or permanent lakes, and oases. A hamada is a type of desert landscape consisting of a high rocky plateau where the sand has been removed by aeolian processes. Other landforms include plains largely covered by gravels and angular boulders, from which the finer particles have been stripped by the wind. These are called "reg" in the western Sahara, "serir" in the eastern Sahara, "gibber plains" in Australia and "saï" in central Asia. The Tassili Plateau in Algeria is an impressive jumble of eroded sandstone outcrops, canyons, blocks, pinnacles, fissures, slabs and ravines. In some places the wind has carved holes or arches, and in others, it has created mushroom-like pillars narrower at the base than the top.George, 1978. pp. 29–30 On the Colorado Plateau, it is water that has been the prevailing eroding force. Here, rivers, such as the Colorado, have cut their way over the millennia through the high desert floor, creating canyons that are over a mile (6,000 feet or 1,800 meters) deep in places, exposing strata that are over two billion years old.

Dust storms and sandstorms

Sand and dust storms are natural events that occur in arid regions where the land is not protected by a covering of vegetation. Dust storms usually start in desert margins rather than the deserts themselves where the finer materials have already been blown away. As a steady wind begins to blow, fine particles lying on the exposed ground begin to vibrate. At greater wind speeds, some particles are lifted into the air stream. When they land, they strike other particles which may be jerked into the air in their turn, starting a chain reaction. Once ejected, these particles move in one of three possible ways, depending on their size, shape and density; suspension, saltation or creep. Suspension is only possible for particles less than in diameter. In a dust storm, these fine particles are lifted up and wafted aloft to heights of up to . They reduce visibility and can remain in the atmosphere for days on end, conveyed by the trade winds for distances of up to . Denser clouds of dust can be formed in stronger winds, moving across the land with a billowing leading edge. The sunlight can be obliterated and it may become as dark as night at ground level.George, 1978. pp. 17–20 In a study of a dust storm in China in 2001, it was estimated that 6.5 million tons of dust were involved, covering an area of . The mean particle size was 1.44 μm. A much smaller scale, short-lived phenomenon can occur in calm conditions when hot air near the ground rises quickly through a small pocket of cooler, low-pressure air above forming a whirling column of particles, a dust devil.

Sandstorms occur with much less frequency than dust storms. They are often preceded by severe dust storms and occur when the wind velocity increases to a point where it can lift heavier particles. These grains of sand, up to about in diameter are jerked into the air but soon fall back to earth, ejecting other particles in the process. Their weight prevents them from being airborne for long and most only travel a distance of a few meters (yards). The sand streams along above the surface of the ground like a fluid, often rising to heights of about . In a really severe steady blow, is about as high as the sand stream can rise as the largest sand grains do not become airborne at all. They are transported by creep, being rolled along the desert floor or performing short jumps.

During a sandstorm, the wind-blown sand particles become electrically charged. Such electric fields, which range in size up to 80 kV/m, can produce sparks and cause interference with telecommunications equipment. They are also unpleasant for humans and can cause headaches and nausea. The electric fields are caused by the collision between airborne particles and by the impacts of saltating sand grains landing on the ground. The mechanism is little understood but the particles usually have a negative charge when their diameter is under 250 μm and a positive one when they are over 500 μm.

Ecology and biogeography

Deserts and semi-deserts are home to ecosystems with low or very low biomass and primary productivity in arid or semi-arid climates. They are mostly found in subtropical high-pressure belts and major continental rain shadows. Primary productivity depends on low densities of small photoautotrophs that sustain a sparse trophic network. Plant growth is limited by rainfall, temperature extremes and desiccating winds. Deserts have strong temporal variability in the availability of resources due to the total amount of annual rainfall and the size of individual rainfall events. Resources are often ephemeral or episodic, and this triggers sporadic animal movements and ‘pulse and reserve’ or ‘boom-bust’ ecosystem dynamics. Erosion and sedimentation are high due to the sparse vegetation cover and the activities of large mammals and people. Plants and animals in deserts are mostly adapted to extreme and prolonged water deficits, but their reproductive phenology often responds to short episodes of surplus. Competitive interactions are weak.

Flora

Plants face severe challenges in arid environments. Problems they need to solve include how to obtain enough water, how to avoid being eaten and how to reproduce. Photosynthesis is the key to plant growth. It can only take place during the day as energy from the sun is required, but during the day, many deserts become very hot. Opening stomata to allow in the carbon dioxide necessary for the process causes evapotranspiration, and conservation of water is a top priority for desert vegetation. Some plants have resolved this problem by adopting crassulacean acid metabolism, allowing them to open their stomata during the night to allow CO₂ to enter, and close them during the day, or by using C4 carbon fixation.

Many desert plants have reduced the size of their leaves or abandoned them altogether. Cacti are desert specialists, and in most species, the leaves have been dispensed with and the chlorophyll displaced into the trunks, the cellular structure of which has been modified to allow them to store water. When rain falls, the water is rapidly absorbed by the shallow roots and retained to allow them to survive until the next downpour, which may be months or years away.George, 1978. pp. 122–123 The giant saguaro cacti of the Sonoran Desert form "forests", providing shade for other plants and nesting places for desert birds. Saguaro grows slowly but may live for up to two hundred years. The surface of the trunk is folded like a concertina, allowing it to expand, and a large specimen can hold eight tons of water after a good downpour.

Cacti are present in both North and South America with a post-Gondwana origin. Other xerophytic plants have developed similar strategies by a process known as convergent evolution. They limit water loss by reducing the size and number of stomata, by having waxy coatings and hairy or tiny leaves. Some are deciduous, shedding their leaves in the driest season, and others curl their leaves up to reduce transpiration. Others store water in succulent leaves or stems or in fleshy tubers. Desert plants maximize water uptake by having shallow roots that spread widely, or by developing long taproots that reach down to deep rock strata for ground water. The saltbush in Australia has succulent leaves and secretes salt crystals, enabling it to live in saline areas. In common with cacti, many have developed spines to ward off browsing animals.

Some desert plants produce seed which lies dormant in the soil until sparked into growth by rainfall. With annuals, such plants grow with great rapidity and may flower and set seed within weeks, aiming to complete their development before the last vestige of water dries up. For perennial plants, reproduction is more likely to be successful if the seed germinates in a shaded position, but not so close to the parent plant as to be in competition with it. Some seed will not germinate until it has been blown about on the desert floor to scarify the seed coat. The seed of the mesquite tree, which grows in deserts in the Americas, is hard and fails to sprout even when planted carefully. When it has passed through the gut of a pronghorn it germinates readily, and the little pile of moist dung provides an excellent start to life well away from the parent tree. The stems and leaves of some plants lower the surface velocity of sand-carrying winds and protect the ground from erosion. Even small fungi and microscopic plant organisms found on the soil surface (so-called ''cryptobiotic soil'') can be a vital link in preventing erosion and providing support for other living organisms. Cold deserts often have high concentrations of salt in the soil. Grasses and low shrubs are the dominant vegetation here and the ground may be covered with lichen

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