Chañares Formation
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The Chañares Formation is a
Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by t ...
-age
geologic formation A geological formation, or simply formation, is a body of rock having a consistent set of physical characteristics ( lithology) that distinguishes it from adjacent bodies of rock, and which occupies a particular position in the layers of rock exp ...
of the
Ischigualasto-Villa Unión Basin The Ischigualasto-Villa Unión Basin ( es, Cuenca de Ischigualasto-Villa Unión) is a small sedimentary basin located in the Argentine Northwest, Argentina. It is located in the southwestern part of La Rioja Province and the northeastern part of ...
, located in La Rioja Province,
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. It is characterized by drab-colored fine-grained
volcaniclastic Volcaniclastics are geologic materials composed of broken fragments (clasts) of volcanic rock. These encompass all clastic volcanic materials, regardless of what process fragmented the rock, how it was subsequently transported, what environment it ...
claystone Mudrocks are a class of fine-grained siliciclastic sedimentary rocks. The varying types of mudrocks include siltstone, claystone, mudstone, slate, and shale. Most of the particles of which the stone is composed are less than and are too sm ...
s,
siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility.Blatt ''et al.'' 1980, p ...
s, and
sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates) ...
s which were deposited in a
fluvial In geography and geology, fluvial processes are associated with rivers and streams and the deposits and landforms created by them. When the stream or rivers are associated with glaciers, ice sheets, or ice caps, the term glaciofluvial or fluviog ...
to
lacustrine A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much larger ...
environment Environment most often refers to: __NOTOC__ * Natural environment, all living and non-living things occurring naturally * Biophysical environment, the physical and biological factors along with their chemical interactions that affect an organism or ...
. The formation is most prominently exposed within
Talampaya National Park Talampaya National Park ( es, Parque Nacional Talampaya) is a national park located in the east/centre of La Rioja Province, Argentina. It was designated a provincial reserve in 1975, a national park in 1997, and a UNESCO World Heritage Site in 2 ...
, a UNESCO
World Heritage Site A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for h ...
within La Rioja Province. The Chañares formation is the lowermost stratigraphic unit of the Agua de la Peña Group, overlying the Tarjados Formation of the Paganzo Group, and underlying the Los Rastros Formation. Though previously considered
Ladinian The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic). ...
in age, U-Pb dating has determined that most or all of the Chañares Formation dates to the early Carnian stage of the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
.Kent et al, 2014, p.7959 The Chañares Formation has provided a diverse and well-preserved faunal assemblage which has been studied intensively since the 1960s. The most common reptiles were proterochampsids ('' Chanaresuchus'', '' Tropidosuchus'', and '' Gualosuchus)'', which lived alongside true
archosaur Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian d ...
s such as ''
Lewisuchus ''Lewisuchus'' is a genus of archosaur that lived during the Late Triassic (early Carnian). As a silesaurid dinosauriform, it was a member of the group of reptiles most commonly considered to be the closest relatives of dinosaurs (possibly tr ...
'', ''
Lagerpeton ''Lagerpeton'' is a genus of lagerpetid avemetatarsalian, comprising a single species, ''L. chanarensis''. First described from the Chañares Formation of Argentina by A. S. Romer in 1971, ''Lagerpeton'' anatomy is somewhat incompletely kn ...
'', ''
Marasuchus ''Marasuchus'' (meaning " Mara crocodile") is a genus of basal dinosauriform archosaur which is possibly synonymous with ''Lagosuchus''. Both genera lived during the Late Triassic in what is now La Rioja Province, Argentina. ''Marasuchus'' cont ...
'', ''
Gracilisuchus ''Gracilisuchus'' (meaning "slender crocodile") is an extinct genus of tiny pseudosuchian (a group which includes the ancestors of crocodilians) from the Late Triassic of Argentina. It contains a single species, ''G. stipanicicorum'', which is pl ...
'', and ''
Luperosuchus ''Luperosuchus'' (meaning "vexing" or "difficult crocodile") is an extinct genus of loricatan pseudosuchian reptile (historically known as a "rauisuchian") which contains only a single species, ''Luperosuchus fractus.'' It is known from the Cha ...
''.
Cynodont The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety ...
s were abundant, represented by the medium-sized traversodontid '' Massetognathus,'' as well as smaller carnivores such as ''
Chiniquodon ''Chiniquodon'' is an extinct genus of carnivorous cynodonts, which lived during the Late Triassic (Carnian) in South America (Argentina and Brazil) and Africa (Namibia and Madagascar). ''Chiniquodon'' was closely related to the genus '' Aleodo ...
'' and ''
Probainognathus ''Probainognathus'' meaning “progressive jaw” is an extinct genus of cynodonts that lived around 235 to 221.5 million years ago, during the Late Triassic in what is now South America. ''Probainognathus'' is a member of the family Probainogna ...
''. The largest animal in the ecosystem was the giant
dicynodont Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typicall ...
''
Dinodontosaurus ''Dinodontosaurus'' (meaning "terrible-toothed lizard") is a genus of dicynodont therapsid. It was medium to large dicynodont of the Triassic (with skull up to long) and had a beak corneum. It lived in the Middle Triassic but disappeared in the ...
''. An older faunal assemblage, distinguished by the large
erpetosuchid Erpetosuchidae is an extinct family of pseudosuchian archosaurs. Erpetosuchidae was named by D. M. S. Watson in 1917 to include ''Erpetosuchus''. It includes the type species ''Erpetosuchus granti'' from the Late Triassic of Scotland, ''Erpetosu ...
''
Tarjadia ''Tarjadia'' is an Extinction, extinct genus of Erpetosuchidae, erpetosuchid pseudosuchian, distantly related to modern crocodilians. It is known from a single species, ''T. ruthae'', first described in 1998 from the Middle Triassic Chañares Fo ...
'', has been discovered in the earliest part of the formation. The formation as a whole is considered one of the best sources of Carnian-age tetrapods in South America, along with the slightly younger
Ischigualasto Formation The Ischigualasto Formation is a Late Triassic fossiliferous formation and Lagerstätte in the Ischigualasto-Villa Unión Basin of the southwestern La Rioja Province and northeastern San Juan Province in northwestern Argentina. The formation da ...
which lies above the Los Rastros Formation.


Geology

The Chañares Formation is the lowermost unit of the Agua de la Peña Group, representing the onset of the first syn-rift phase of the
Ischigualasto-Villa Unión Basin The Ischigualasto-Villa Unión Basin ( es, Cuenca de Ischigualasto-Villa Unión) is a small sedimentary basin located in the Argentine Northwest, Argentina. It is located in the southwestern part of La Rioja Province and the northeastern part of ...
. It unconformably overlies red
fluvial In geography and geology, fluvial processes are associated with rivers and streams and the deposits and landforms created by them. When the stream or rivers are associated with glaciers, ice sheets, or ice caps, the term glaciofluvial or fluviog ...
(river) sediments of the Tarjados Formation of the Paganzo Group, and is conformably overlain by greenish lake and delta sediments of the Los Rastros Formation. The Chañares Formation has also had an interesting history with the lower part of the Ischichuca Formation. This sequence of sediments on the western edge of the basin has occasionally been considered to take priority over the Chañares Formation and the lower part of the Los Rastros Formation. However, this is rare due to the historical significance of the Chañares Formation. As a result, some authors restrict the Ischichuca Formation to a few layers of lake and delta sediments between the Chañares and Los Rastros formations, while others reject using it in the first place. The Chañares formation was originally thought to be deposited during the
Ladinian The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic). ...
stage of the
Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and ...
. However, Uranium-Lead radiometric dating by Marsicano ''et al.'' (2016) later found that a large portion of the formation was deposited in the early
Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by t ...
(237–234 Ma), near the start of the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
.Marsicano ''et al.'', 2016 2020 U-Pb dating of the overlying lower Los Rastros Formation yielded an age of 234.47 ± 0.44 Ma, making the vast majority of the Chañares Formation lowermost Carnian.Mancuso ''et al.'', 2020 Nevertheless, the Ladinian-Carnian boundary may still lie within the first few meters of the formation, despite the primary fossiliferous sections being well-supported as early Carnian in age. The formation is primarily
sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates) ...
,
siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility.Blatt ''et al.'' 1980, p ...
, and
claystone Mudrocks are a class of fine-grained siliciclastic sedimentary rocks. The varying types of mudrocks include siltstone, claystone, mudstone, slate, and shale. Most of the particles of which the stone is composed are less than and are too sm ...
, arranged in a specific sequence of
facies In geology, a facies ( , ; same pronunciation and spelling in the plural) is a body of rock with specified characteristics, which can be any observable attribute of rocks (such as their overall appearance, composition, or condition of formatio ...
. A distinct and uneven
unconformity An unconformity is a buried erosional or non-depositional surface separating two rock masses or strata of different ages, indicating that sediment deposition was not continuous. In general, the older layer was exposed to erosion for an interval o ...
separates the base of the Chañares Formation from the underlying Tarjados Formation. In the field, this unconformity can be identified by a high-relief layer of
chert Chert () is a hard, fine-grained sedimentary rock composed of microcrystalline or cryptocrystalline quartz, the mineral form of silicon dioxide (SiO2). Chert is characteristically of biological origin, but may also occur inorganically as a prec ...
at the top of the Tarjados Formation.Rogers ''et al.'', 2001Mancuso ''et al.'', 2014Ezcurra ''et al.'', 2017


Fluvial olive-grey facies

Above this unconformity lies the oldest section of the Chañares Formation, a package of olive-grey fluvial sediments. As one goes up the section, increasingly finer beds of sandstone and siltstone are interlayered with coarser lenses, corresponding to periodic
sheet flood Sheet erosion or sheet wash is the even erosion of substrate along a wide area. It occurs in a wide range of settings such as coastal plains, hillslopes, floodplains, beaches, savanna plains and semi-arid plains. Water moving fairly uniformly with ...
s along
braided river A braided river, or braided channel, consists of a network of river channels separated by small, often temporary, islands called braid bars or, in English usage, ''aits'' or ''eyots''. Braided streams tend to occur in rivers with high sediment l ...
s. Weakly-developed palaeosols can be found within this section, filled with root traces, pebbles and small brown
calcareous Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines. In zoology ''Calcareous'' is used as an adje ...
nodules. Winding systems of burrows have also been found in this section, likely created by small
cynodont The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety ...
s. Though fossils are relatively uncommon in the lower fluvial beds of the Chañares Formation, they are
taxonomically In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given ...
and taphonomically distinct from those of succeeding layers. The most common remains are from large
dicynodonts Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were Herbivore, herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horn ...
(possibly referable to ''
Dinodontosaurus ''Dinodontosaurus'' (meaning "terrible-toothed lizard") is a genus of dicynodont therapsid. It was medium to large dicynodont of the Triassic (with skull up to long) and had a beak corneum. It lived in the Middle Triassic but disappeared in the ...
''), the large
erpetosuchid Erpetosuchidae is an extinct family of pseudosuchian archosaurs. Erpetosuchidae was named by D. M. S. Watson in 1917 to include ''Erpetosuchus''. It includes the type species ''Erpetosuchus granti'' from the Late Triassic of Scotland, ''Erpetosu ...
''
Tarjadia ''Tarjadia'' is an Extinction, extinct genus of Erpetosuchidae, erpetosuchid pseudosuchian, distantly related to modern crocodilians. It is known from a single species, ''T. ruthae'', first described in 1998 from the Middle Triassic Chañares Fo ...
'', and small non- massetognathine cynodonts closely related to ''
Aleodon ''Aleodon'' is an extinct genus of cynodonts that lived from the Middle to the Late Triassic. Relatively few analyses have been conducted to identify the phylogenetic placement of ''Aleodon'', however those that have place ''Aleodon'' as a siste ...
'' and '' Scalenodon''. This ecosystem has been termed the ''Tarjadia'' Assemblage Zone, in order to distinguish it from the slightly younger classic Chañares assemblage. No radiometric dating has been done on this section, but it may contain the Ladinian-Carnian boundary based on dates obtained within the younger facies.


Fossiliferous bluish facies

Above the olive-grey fluvial beds is the most fossiliferous and well-studied portion of the formation. This section is characterized by wide and massive layers of very fine bluish-grey sandstone, siltstone, and claystone. These layers have a high concentration of volcanic ash and debris, ranging from glassy shards at the base to weathered
bentonite Bentonite () is an absorbent swelling clay consisting mostly of montmorillonite (a type of smectite) which can either be Na-montmorillonite or Ca-montmorillonite. Na-montmorillonite has a considerably greater swelling capacity than Ca-mon ...
s at the top. Nearby volcanic eruptions likely impacted the local climate and river systems, shifting the depositional regime from a stable braided fluvial system to one dominated by shallow floodplains and
lahar A lahar (, from jv, ꦮ꧀ꦭꦲꦂ) is a violent type of mudflow or debris flow composed of a slurry of pyroclastic material, rocky debris and water. The material flows down from a volcano, typically along a river valley. Lahars are extreme ...
s. Marsicano ''et al.'' (2016) obtained a CA-TIMS U-Pb age of 236.1 ± 0.6 Ma from a siltstone bed immediately below the first major fossil layer within the bluish facies. Ezcurra ''et al.'' (2017) later studied a slightly older sandstone bed using LA-MC-IMP-MS U-Pb dating. They found a date of 236.2 ± 1.1 Ma from a cluster of the three youngest
zircon Zircon () is a mineral belonging to the group of nesosilicates and is a source of the metal zirconium. Its chemical name is zirconium(IV) silicate, and its corresponding chemical formula is Zr SiO4. An empirical formula showing some of the r ...
s. The most productive and historically relevant fossil beds of the Chañares Formation lie within these
volcaniclastic Volcaniclastics are geologic materials composed of broken fragments (clasts) of volcanic rock. These encompass all clastic volcanic materials, regardless of what process fragmented the rock, how it was subsequently transported, what environment it ...
layers. The layers are replete with massive calcareous
concretion A concretion is a hard, compact mass of matter formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular ...
s, some up to 2 meters in width. They commonly preserve articulated skeletal material, often complete skeletons from multiple taxa in a single concretion. Well-preserved fossils found within concretions typically represent smaller taxa, which were buried rapidly after death. Volcanic catastrophes such as lahars, ash falls, or
pyroclastic flow A pyroclastic flow (also known as a pyroclastic density current or a pyroclastic cloud) is a fast-moving current of hot gas and volcanic matter (collectively known as tephra) that flows along the ground away from a volcano at average speeds of bu ...
s are the preferred cause of these rapid
mass mortality event A mass mortality event (MME) is a incident that kills a vast number of individuals of a single species in a short period of time. The event may put a species at risk of extinction or upset an ecosystem. This is distinct from the mass die-off ass ...
s. More fragmentary fossils are also occasionally found outside of concretions, much like fossils of the underlying ''Tarjadia'' AZ. Fossils found outside concretions are typically from large animals, and were probably buried more slowly (albeit still fairly rapidly) by gradual processes. The classic Chañares fauna, recently codified as the ''Massetognathus''-''Chanaresuchus'' Assemblage Zone, is characteristic of this section of the formation. The largest components of the ecosystem include dicynodonts (namely ''
Dinodontosaurus ''Dinodontosaurus'' (meaning "terrible-toothed lizard") is a genus of dicynodont therapsid. It was medium to large dicynodont of the Triassic (with skull up to long) and had a beak corneum. It lived in the Middle Triassic but disappeared in the ...
'') and indeterminate carnivorous
paracrocodylomorphs Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha ...
, similar to taxa from the underlying ''Tarjadia'' AZ. By far the most common fossils belong to three species of
cynodont The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety ...
s: ''
Massetognathus pascuali ''Massetognathus'' ( ; Greek for "chewing muscle jaw") is an extinct genus of plant-eating traversodontid cynodonts. They lived during the Triassic Period about 235 million years ago, and are known from the Chañares Formation in Argentina and t ...
, Probainognathus jenseni,'' and '' Chiniquodon theotonicus.''These three species comprise nearly 3/4ths of the fossils found in the formation, with almost half of the total fossils in the formation referred to ''Massetognathus'' alone. Reptile fossils are less prevalent but more diverse, with the most common belonging to the proterochampsid
archosauriform Archosauriformes (Greek for 'ruling lizards', and Latin for 'form') is a clade of diapsid reptiles that developed from archosauromorph ancestors some time in the Latest Permian (roughly 252 million years ago). It was defined by Jacques Gauthier ...
'' Chanaresuchus''. Other notable taxa include ''
Lagerpeton ''Lagerpeton'' is a genus of lagerpetid avemetatarsalian, comprising a single species, ''L. chanarensis''. First described from the Chañares Formation of Argentina by A. S. Romer in 1971, ''Lagerpeton'' anatomy is somewhat incompletely kn ...
'', ''
Lagosuchus ''Lagosuchus'' is an extinct genus of avemetatarsalian archosaur from the Late Triassic of Argentina. The type species of ''Lagosuchus'', ''Lagosuchus talampayensis,'' is based on a small partial skeleton recovered from the early Carnian-age Cha ...
'', and ''
Lewisuchus ''Lewisuchus'' is a genus of archosaur that lived during the Late Triassic (early Carnian). As a silesaurid dinosauriform, it was a member of the group of reptiles most commonly considered to be the closest relatives of dinosaurs (possibly tr ...
'', which were among the oldest ornithodirans, elaborating on the ancestry of dinosaurs and pterosaurs. Several communal
latrines A latrine is a toilet or an even simpler facility that is used as a toilet within a sanitation system. For example, it can be a communal trench in the earth in a camp to be used as emergency sanitation, a hole in the ground (pit latrine), or m ...
are known from the bluish facies, preserving dicynodont
coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is de ...
s filled with plant remains.


Upper member

The olive-grey fluvial section and the bluish-grey volcaniclastic section collectively form the lower
member Member may refer to: * Military jury, referred to as "Members" in military jargon * Element (mathematics), an object that belongs to a mathematical set * In object-oriented programming, a member of a class ** Field (computer science), entries in ...
of the Chañares Formation. They are overlain by an upper member, which is practically devoid of body fossils. The upper member is generally similar to the bluish facies in appearance, with wide beds of fine-grained volcaniclastic sediments with a pale grey color. Unlike the bluish facies, concretions and body fossils are absent, having been replaced by '' Taenidium'' worm burrows. These burrows likely indicate that the environment had transitioned into a
lacustrine A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much larger ...
(lake) ecosystem by the time that the upper member was deposited. Some outcrops preserve a few sandstone and conglomerate beds near the top of the upper member. These indicate that the depositional regime continued to shift towards the
deltaic A river delta is a landform shaped like a triangle, created by deposition of sediment that is carried by a river and enters slower-moving or stagnant water. This occurs where a river enters an ocean, sea, estuary, lake, reservoir, or (more rare ...
environment of the overlying Los Rastros Formation. The coarser beds also contain a few rare body fossils from fish and
tetrapod Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct theraps ...
s.Gouiric-cavalli ''et al.'', 2017 A white
tuff Tuff is a type of rock made of volcanic ash ejected from a vent during a volcanic eruption. Following ejection and deposition, the ash is lithified into a solid rock. Rock that contains greater than 75% ash is considered tuff, while rock cont ...
near the base of the upper member has been dated to 233.7 ± 0.4 Ma, while a zircon cluster from a sandy tuff at the top of the formation was dated to 233.6 ± 1.1 Ma.


Paleobiota

Plant remains and palynomorphs preserved in the
dicynodont Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typicall ...
coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is de ...
s were described in
2018 File:2018 Events Collage.png, From top left, clockwise: The 2018 Winter Olympics opening ceremony in PyeongChang, South Korea; Protests erupt following the Assassination of Jamal Khashoggi; March for Our Lives protests take place across the United ...
. Though it is difficult to determine the affinities of the larger plant fragments, the palynomorphs are more conclusive. They belong to a broad range of plants, most abundantly pollen from umkomasiales (a type of
seed fern A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosperm ...
), and in smaller portions from podocarpean and voltzialean
conifers Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extan ...
. Spores from humid-loving groups such as
bryophyte The Bryophyta s.l. are a proposed taxonomic division containing three groups of non-vascular land plants (embryophytes): the liverworts, hornworts and mosses. Bryophyta s.s. consists of the mosses only. They are characteristically limited in ...
s, lycopsids, true
fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
s, and
algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...
were also present but rare. The palynomorph taxa generally resemble those of the ''
Dicroidium ''Dicroidium'' is an extinct genus of fork-leaved seed ferns that were widely distributed over Gondwana during the Triassic (). Their fossils are known from South Africa, the Arabian Peninsula, Australia, New Zealand, South America, Madagascar ...
'' flora which is common in other late Ladinian-early Carnian units. More precisely, the flora is intermediate between the temperate Ipswich flora of far southern
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
, and the hot, subtropical Onslow flora which developed along the southern shore of the
Neotethys The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...
. This transitional character is also observed in the flora of the nearby
Ischigualasto Formation The Ischigualasto Formation is a Late Triassic fossiliferous formation and Lagerstätte in the Ischigualasto-Villa Unión Basin of the southwestern La Rioja Province and northeastern San Juan Province in northwestern Argentina. The formation da ...
, as well as the Flagstone Bench Formation of
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
. Curiously, the Los Rastros Formation, which was deposited between the Chañares and Ischigualasto Formations, preserves a typical Ipswich flora. This likely indicates that all three formations lie at a latitude which allows them to quickly shift between the different floras during small climatic changes.Pérez ''et al.'', 2018 Tetrapod burrows, likely produced by small eucynodonts, have been described from the lower section of the formation.Fiorelli ''et al.'', 2018


Therapsids


Reptiles


Fish


See also

*
Ischigualasto Formation The Ischigualasto Formation is a Late Triassic fossiliferous formation and Lagerstätte in the Ischigualasto-Villa Unión Basin of the southwestern La Rioja Province and northeastern San Juan Province in northwestern Argentina. The formation da ...
* Los Rastros Formation *
Santa Maria Formation The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It is primarily Carnian in age (Late Triassic), and is notable for its fossils of cynodonts, "rauisuchian" pseudosuchians, and early dinosaurs and oth ...
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Omingonde Formation The Omingonde Formation is an Early Triassic, Early to Middle Triassic (Anisian to Ladinian) geologic Formation (geology), formation, part of the Karoo Supergroup, in the western Otjozondjupa Region and northeastern Erongo Region of north-central N ...


References


Bibliography

* * * * * * * * * * * * * * * * * * * * * * * ;The Chañares Triassic reptile fauna * * * * * * * * * * * * * * * * * * * * {{DEFAULTSORT:Chanares Formation Geologic formations of Argentina Triassic Argentina Shale formations Tuff formations Fluvial deposits Lacustrine deposits Fossiliferous stratigraphic units of South America Paleontology in Argentina Geology of La Rioja Province, Argentina