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Cattle (''Bos taurus'') are large, domesticated, cloven-hooved,
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
s. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Adult females are referred to as cows and adult males are referred to as bulls. Cattle are commonly raised as
livestock Livestock are the domesticated animals raised in an agricultural setting to provide labor and produce diversified products for consumption such as meat, eggs, milk, fur, leather, and wool. The term is sometimes used to refer solely to animals ...
for meat (
beef Beef is the culinary name for meat from cattle (''Bos taurus''). In prehistoric times, humankind hunted aurochs and later domesticated them. Since that time, numerous breeds of cattle have been bred specifically for the quality or quantity ...
or
veal Veal is the meat of calves, in contrast to the beef from older cattle. Veal can be produced from a calf of either sex and any breed, however most veal comes from young male calves of dairy breeds which are not used for breeding. Generally, v ...
, see beef cattle), for
milk Milk is a white liquid food produced by the mammary glands of mammals. It is the primary source of nutrition for young mammals (including breastfed human infants) before they are able to digestion, digest solid food. Immune factors and immune ...
(see dairy cattle), and for hides, which are used to make
leather Leather is a strong, flexible and durable material obtained from the tanning, or chemical treatment, of animal skins and hides to prevent decay. The most common leathers come from cattle, sheep, goats, equine animals, buffalo, pigs and hogs, ...
. They are used as
riding animal A working animal is an animal, usually domesticated, that is kept by humans and trained to perform tasks instead of being slaughtered to harvest animal products. Some are used for their physical strength (e.g. oxen and draft horses) or for t ...
s and
draft animal A working animal is an animal, usually domesticated, that is kept by humans and trained to perform tasks instead of being slaughtered to harvest animal products. Some are used for their physical strength (e.g. oxen and draft horses) or for t ...
s ( oxen or bullocks, which pull
cart A cart or dray (Australia and New Zealand) is a vehicle designed for transport, using two wheels and normally pulled by one or a pair of draught animals. A handcart is pulled or pushed by one or more people. It is different from the flatbed tr ...
s,
plow A plough or plow ( US; both ) is a farm tool for loosening or turning the soil before sowing seed or planting. Ploughs were traditionally drawn by oxen and horses, but in modern farms are drawn by tractors. A plough may have a wooden, iron or ...
s and other implements). Another product of cattle is their dung, which can be used to create manure or
fuel A fuel is any material that can be made to react with other substances so that it releases energy as thermal energy or to be used for work. The concept was originally applied solely to those materials capable of releasing chemical energy but ...
. In some regions, such as parts of India, cattle have significant religious significance. Cattle, mostly small breeds such as the Miniature Zebu, are also kept as pets. Different types of cattle are common to different geographic areas. Taurine cattle are found primarily in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
and temperate areas of
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
, the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...
, and
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
. Zebus (also called indicine cattle) are found primarily in
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
and tropical areas of Asia, America, and Australia.
Sanga cattle Sanga cattle is the collective name for indigenous cattle of sub-Saharan Africa. They are sometimes identified as a subspecies with the scientific name ''Bos taurus africanus''. Their history of domestication and their origins in relation to ta ...
are found primarily in
sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
. These types (which are sometimes classified as separate species or subspecies) are further divided into over 1000 recognized breeds. Around 10,500 years ago, taurine cattle were domesticated from as few as 80 wild
aurochs The aurochs (''Bos primigenius'') ( or ) is an extinct cattle species, considered to be the wild ancestor of modern domestic cattle. With a shoulder height of up to in bulls and in cows, it was one of the largest herbivores in the Holocen ...
progenitors in
central Anatolia The Central Anatolia Region ( tr, İç Anadolu Bölgesi) is a geographical region of Turkey. The largest city in the region is Ankara. Other big cities are Konya, Kayseri, Eskişehir, Sivas, and Aksaray. Located in Central Turkey, it is bordered ...
, the
Levant The Levant () is an approximate historical geographical term referring to a large area in the Eastern Mediterranean region of Western Asia. In its narrowest sense, which is in use today in archaeology and other cultural contexts, it is eq ...
and
Western Iran Western Iran consists of Armenian Highlands, Northern Zagros and the rich agricultural area of the Khuzestan Plain in the south. It includes the provinces of Kordestan, Kermanshah, Ilam, Hamadan and Lorestan. Some references also count West ...
. Op. cit. in A separate domestication event occurred in the
Indian subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
, which gave rise to zebu. According to the
Food and Agriculture Organization The Food and Agriculture Organization of the United Nations (FAO)french: link=no, Organisation des Nations unies pour l'alimentation et l'agriculture; it, Organizzazione delle Nazioni Unite per l'Alimentazione e l'Agricoltura is an intern ...
(FAO), there are approximately 1.5 billion cattle in the world as of 2018. Cattle are the main source of
greenhouse gas emissions Greenhouse gas emissions from human activities strengthen the greenhouse effect, contributing to climate change. Most is carbon dioxide from burning fossil fuels: coal, oil, and natural gas. The largest emitters include coal in China and lar ...
from livestock, and are responsible for around 10% of global greenhouse gas emissions. In 2009, cattle became one of the first livestock animals to have a fully mapped
genome In the fields of molecular biology and genetics, a genome is all the genetic information of an organism. It consists of nucleotide sequences of DNA (or RNA in RNA viruses). The nuclear genome includes protein-coding genes and non-coding ge ...
.


Taxonomy

Cattle were originally identified as three separate species: ''Bos taurus'', the European or "taurine" cattle (including similar types from Africa and Asia); ''Bos indicus'', the Indicine or "zebu"; and the extinct ''Bos primigenius'', the
aurochs The aurochs (''Bos primigenius'') ( or ) is an extinct cattle species, considered to be the wild ancestor of modern domestic cattle. With a shoulder height of up to in bulls and in cows, it was one of the largest herbivores in the Holocen ...
. The aurochs is ancestral to both zebu and taurine cattle. They were later reclassified as one species, ''Bos taurus'', with the aurochs, zebu, and taurine cattle as subspecies. However, this taxonomy is contentious and some sources prefer the separate species classification, such as the
American Society of Mammalogists The American Society of Mammalogists (ASM) was founded in 1919. Its primary purpose is to encourage the study of mammals, and professions studying them. There are over 4,500 members of this society, and they are primarily professional scientists ...
' Mammal Diversity Database. Complicating the matter is the ability of cattle to
interbreed In biology, a hybrid is the offspring resulting from combining the qualities of two organisms of different breeds, varieties, species or genera through sexual reproduction. Hybrids are not always intermediates between their parents (such as in ...
with other closely related species. Hybrid individuals and even breeds exist, not only between taurine cattle and zebu (such as the
sanga cattle Sanga cattle is the collective name for indigenous cattle of sub-Saharan Africa. They are sometimes identified as a subspecies with the scientific name ''Bos taurus africanus''. Their history of domestication and their origins in relation to ta ...
(''Bos taurus africanus'' x ''Bos indicus''), but also between one or both of these and some other members of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
'' Bos''
yak The domestic yak (''Bos grunniens''), also known as the Tartary ox, grunting ox or hairy cattle, is a species of long-haired domesticated cattle found throughout the Himalayan region of the Indian subcontinent, the Tibetan Plateau, Kachin Sta ...
s (the
dzo A dzo (also spelled zo, zho and dzho, bo, མཛོ་, mdzo) is a hybrid between the yak and domestic cattle. The word dzo technically refers to a male hybrid, while a female is known as a or . In Mongolian, it is called a (хайнаг). ...
or yattle),
banteng The banteng (''Bos javanicus''; ), also known as tembadau, is a species of cattle found in Southeast Asia. The head-and-body length is between . Wild banteng are typically larger and heavier than their domesticated counterparts, but are otherw ...
, and gaur. Hybrids such as the beefalo breed can even occur between taurine cattle and either species of
bison Bison are large bovines in the genus ''Bison'' (Greek: "wild ox" (bison)) within the tribe Bovini. Two extant and numerous extinct species are recognised. Of the two surviving species, the American bison, ''B. bison'', found only in North Ame ...
, leading some authors to consider them part of the genus ''Bos'', as well. The hybrid origin of some types may not be obvious – for example,
genetic testing Genetic testing, also known as DNA testing, is used to identify changes in DNA sequence or chromosome structure. Genetic testing can also include measuring the results of genetic changes, such as RNA analysis as an output of gene expression, or ...
of the
Dwarf Lulu The Dwarf Lulu is a breed of cattle that is native to the mountains of Central Asia. It has taurine, yak and zebu The zebu (; ''Bos indicus'' or ''Bos taurus indicus''), sometimes known in the plural as indicine cattle or humped cattle, is a s ...
breed, the only taurine-type cattle in Nepal, found them to be a mix of taurine cattle, zebu, and yak. However, cattle cannot be successfully hybridized with more distantly related bovines such as
water buffalo The water buffalo (''Bubalus bubalis''), also called the domestic water buffalo or Asian water buffalo, is a large bovid originating in the Indian subcontinent and Southeast Asia. Today, it is also found in Europe, Australia, North America, So ...
or African buffalo. The aurochs originally ranged throughout Europe, North Africa, and much of Asia. In historical times, its range became restricted to Europe, and the last known individual died in Mazovia, Poland, in about 1627. Breeders have attempted to recreate cattle of similar appearance to aurochs by crossing traditional types of domesticated cattle, creating the
Heck cattle Heck cattle are a hardy breed of domestic cattle. These cattle are the result of an attempt by Heinz and Lutz Heck to breed back the extinct aurochs (''Bos primigenius'') from modern aurochs-derived cattle in the 1920s and 1930s. Controver ...
breed. The only pure African taurine breeds (''Bos taurus africanus'') remaining are the N'Dama, Kuri and some varieties of the West African Shorthorn.


Etymology

''Cattle'' did not originate as the term for bovine animals. It was borrowed from
Anglo-Norman Anglo-Norman may refer to: *Anglo-Normans, the medieval ruling class in England following the Norman conquest of 1066 *Anglo-Norman language **Anglo-Norman literature *Anglo-Norman England, or Norman England, the period in English history from 1066 ...
, itself from medieval Latin 'principal sum of money, capital', itself derived in turn from Latin 'head'. ''Cattle'' originally meant movable
personal property property is property that is movable. In common law systems, personal property may also be called chattels or personalty. In civil law systems, personal property is often called movable property or movables—any property that can be moved fr ...
, especially livestock of any kind, as opposed to
real property In English common law, real property, real estate, immovable property or, solely in the US and Canada, realty, is land which is the property of some person and all structures (also called improvements or fixtures) integrated with or affixe ...
(the land, which also included wild or small free-roaming animals such as chickens—they were sold as part of the land). The word is a variant of ''
chattel Chattel may refer to: * Chattel, an alternative name for tangible personal property * A chattel house, a type of West Indian dwelling * A chattel mortgage, a security interest over tangible personal property * Chattel slavery, the most extreme form ...
'' (a unit of personal property) and closely related to ''
capital Capital may refer to: Common uses * Capital city, a municipality of primary status ** List of national capital cities * Capital letter, an upper-case letter Economics and social sciences * Capital (economics), the durable produced goods used f ...
'' in the economic sense. The term replaced earlier
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlement of Britain, Anglo ...
'cattle, property', which survives today as ''fee'' ( cf. , , ). The word ''cow'' came via
Anglo-Saxon The Anglo-Saxons were a Cultural identity, cultural group who inhabited England in the Early Middle Ages. They traced their origins to settlers who came to Britain from mainland Europe in the 5th century. However, the ethnogenesis of the Anglo- ...
(plural ), from
Common Indo-European Proto-Indo-European (PIE) is the reconstructed common ancestor of the Indo-European language family. Its proposed features have been derived by linguistic reconstruction from documented Indo-European languages. No direct record of Proto-Indo-E ...
(
genitive In grammar, the genitive case (abbreviated ) is the grammatical case that marks a word, usually a noun, as modifying another word, also usually a noun—thus indicating an attributive relationship of one noun to the other noun. A genitive can al ...
) 'a bovine animal', cf. fa, gâv, script=Latn, sa, go-, script=Latn, cy, buwch. The plural became or in Middle English, and an additional plural ending was often added, giving , , but also , and others. This is the origin of the now archaic English plural, ''kine''. The
Scots language Scots ( endonym: ''Scots''; gd, Albais, ) is an Anglic language variety in the West Germanic language family, spoken in Scotland and parts of Ulster in the north of Ireland (where the local dialect is known as Ulster Scots). Most commonly ...
singular is or , and the plural is . In older English sources such as the
King James Version The King James Version (KJV), also the King James Bible (KJB) and the Authorized Version, is an Bible translations into English, English translation of the Christian Bible for the Church of England, which was commissioned in 1604 and publis ...
of the Bible, ''cattle'' refers to livestock, as opposed to ''deer'' which refers to wildlife. ''Wild cattle'' may refer to feral cattle or to undomesticated species of the genus '' Bos''. Today, when used without any other qualifier, the modern meaning of ''cattle'' is usually restricted to domesticated bovines.


Terminology

In general, the same words are used in different parts of the world, but with minor differences in the definitions. The terminology described here contrasts the differences in definition between the United Kingdom and other British-influenced parts of the world such as Canada, Australia, New Zealand, Ireland and the United States. * An "intact" (i.e., not castrated) adult male is called a '' bull''. ** A father bull is called a ''sire'' with reference to his offspring. * An adult female that has had a calf (or two, depending on regional usage) is a ''cow''. ** A mother cow is called a ''dam'' with reference to her offspring. Often, mentions of ''dams'' imply cows kept in the herd for repeated breeding (as opposed to heifers or cows sold off sooner). * A young female before she has had a calf of her own and who is under three years of age is called a ''
heifer Heifer may refer to: * Heifer (cow), a young cow before she has had her first calf * Frank Heifer (1854–1893), American outfielder and first baseman * '' The Heifer'' (''La vaquilla''), 1985 Spanish comedy film * Heifer International Heifer ...
'' ( ).Delbridge, Arthur, The Macquarie Dictionary, 2nd ed., Macquarie Library, North Ryde, 1991 A young female that has had only one calf is occasionally called a ''first-calf heifer''. ''Heiferettes'' are either first-calf heifers or a subset thereof without potential to become lineage dams, depending on whose definition is operative. * Young cattle (regardless of sex) are called '' calves'' until they are
weaned Weaning is the process of gradually introducing an infant human or another mammal to what will be its adult diet while withdrawing the supply of its mother's milk. The process takes place only in mammals, as only mammals produce milk. The infan ...
, then ''weaners'' until they are a year old in some areas; in other areas, particularly with male beef cattle, they may be known as ''feeder calves'' or ''feeders''. After that, they are referred to as ''yearlings'' or ''stirks'' if between one and two years of age. * '' Feeder cattle'' or ''store cattle'' are young cattle soon to be either backgrounded or sent to fattening, most especially those intended to be sold to someone else for finishing. In some regions, a distinction between ''stockers'' and ''feeders'' (by those names) is the distinction of backgrounding versus immediate sale to a finisher. * A castrated male is called a ''steer'' in the United States; older steers are often called ''bullocks'' in other parts of the world, but in North America this term refers to a young bull. Piker bullocks are micky bulls (uncastrated young male bulls) that were caught, castrated and then later lost. In Australia, the term ''Japanese ox'' is used for grain-fed steers in the weight range of 500 to 650 kg that are destined for the Japanese meat trade. In North America, draft cattle under four years old are called working steers. Improper or late castration on a bull results in it becoming a coarse steer known as a ''stag'' in Australia, Canada and New Zealand. In some countries, an incompletely castrated male is known also as a ''
rig Rig may refer to: Objects and structures * Rig (fishing), an arrangement of items used for fishing * Drilling rig, a structure housing equipment used to drill or extract oil from underground * Rig (stage lighting) * rig, a horse-drawn carriage ...
''. * A castrated male (occasionally a female or in some areas a bull) kept for draft or riding purposes is called an '' ox'' (plural ''oxen''); ''ox'' may also be used to refer to some carcass products from any adult cattle, such as ox-hide, ox-blood, oxtail, or ox-liver. * A ''springer'' is a cow or heifer close to calving. * In all cattle species, a female twin of a bull usually becomes an infertile partial
intersex Intersex people are individuals born with any of several sex characteristics including chromosome patterns, gonads, or genitals that, according to the Office of the United Nations High Commissioner for Human Rights, "do not fit typical bina ...
, and is called a '' freemartin''. * A wild, young, unmarked bull is known as a ''micky'' in Australia.Coupe, Sheena (ed.), Frontier Country, Vol. 1, Weldon Russell Publishing, Willoughby, 1989, * An unbranded bovine of either sex is called a ''maverick'' in the US and Canada. * ''Neat'' (horned oxen, from which neatsfoot oil is derived), ''beef'' (young ox) and ''beefing'' (young animal fit for slaughtering) are obsolete terms, although ''poll'', ''pollard'' and '' polled cattle'' are still terms in use for naturally hornless animals, or in some areas also for those that have been disbudded or dehorned. * Cattle raised for human consumption are called '' beef cattle''. Within the American beef cattle industry, the older term beef (plural beeves) is still used to refer to an animal of either sex. Some Australian, Canadian, New Zealand and British people use the term ''beast''. * Cattle bred specifically for milk production are called ''milking'' or '' dairy cattle''; a cow kept to provide milk for one family may be called a ''
house cow A house cow is a cow kept to provide milk for a home kitchen. This differentiates them from dairy cows, which are farmed commercially. They can also provide manure, for use as a garden fertilizer, and their offspring can be a source of meat. H ...
'' or ''milker''. A ''fresh cow'' is a dairy term for a cow or first-calf heifer who has recently given birth, or "freshened." * The adjective applying to cattle in general is usually ''bovine''. The terms ''bull'', ''cow'' and ''calf'' are also used by extension to denote the sex or age of other large animals, including whales,
hippopotamus The hippopotamus ( ; : hippopotamuses or hippopotami; ''Hippopotamus amphibius''), also called the hippo, common hippopotamus, or river hippopotamus, is a large semiaquatic mammal native to sub-Saharan Africa. It is one of only two extan ...
es,
camel A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. C ...
s,
elk The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The common ...
and elephants. * Various other terms for cattle or types thereof are
historical History (derived ) is the systematic study and the documentation of the human activity. The time period of event before the invention of writing systems is considered prehistory. "History" is an umbrella term comprising past events as well ...
; these include ''
nowt "Zero" is the usual name for the number 0 in English. In British English "nought" is also used. In American English "naught" is used occasionally for zero, but (as with British English) "naught" is more often used as an archaic word for nothing. "N ...
'', '' nolt'', ''
mart Mart may refer to: * Mart, or marketplace, a location where people regularly gather for the purchase and sale of provisions, livestock, and other goods * Mart (broadcaster), a local broadcasting station in Amsterdam * Mart (given name) * Mart (Syr ...
'', and others.


Singular terminology issue

"Cattle" can only be used in the
plural The plural (sometimes abbreviated pl., pl, or ), in many languages, is one of the values of the grammatical category of number. The plural of a noun typically denotes a quantity greater than the default quantity represented by that noun. This de ...
and not in the
singular Singular may refer to: * Singular, the grammatical number that denotes a unit quantity, as opposed to the plural and other forms * Singular homology * SINGULAR, an open source Computer Algebra System (CAS) * Singular or sounder, a group of boar, ...
: it is a plurale tantum. Thus one may refer to "three cattle" or "some cattle", but not "one cattle". "One head of cattle" is a valid though periphrastic way to refer to one animal of indeterminate or unknown age and sex; otherwise no universally used single-word singular form of ''cattle'' exists in modern English, other than the sex- and age-specific terms such as ''cow'', ''bull'', ''steer'' and ''heifer''. Historically, "ox" was not a sex-specific term for adult cattle, but generally this is now used only for working cattle, especially adult castrated males. The term is also incorporated into the names of other species, such as the musk ox and "grunting ox" (
yak The domestic yak (''Bos grunniens''), also known as the Tartary ox, grunting ox or hairy cattle, is a species of long-haired domesticated cattle found throughout the Himalayan region of the Indian subcontinent, the Tibetan Plateau, Kachin Sta ...
), and is used in some areas to describe certain cattle products such as ox-hide and oxtail. ''Cow'' is in general use as a singular for the collective ''cattle''. The word ''cow'' is easy to use when a singular is needed and the sex is unknown or irrelevant—when "there is a cow in the road", for example. Further, any herd of fully mature cattle in or near a
pasture Pasture (from the Latin ''pastus'', past participle of ''pascere'', "to feed") is land used for grazing. Pasture lands in the narrow sense are enclosed tracts of farmland, grazed by domesticated livestock, such as horses, cattle, sheep, or swine ...
is statistically likely to consist mostly of cows, so the term is probably accurate even in the restrictive sense. Other than the few bulls needed for breeding, the vast majority of male cattle are castrated as calves and are used as oxen or slaughtered for meat before the age of three years. Thus, in a pastured herd, any calves or herd bulls usually are clearly distinguishable from the cows due to distinctively different sizes and clear anatomical differences. Merriam-Webster and Oxford Living Dictionaries recognize the sex-nonspecific use of ''cow'' as an alternate definition, whereas Collins and the OED do not.
Colloquial Colloquialism (), also called colloquial language, everyday language or general parlance, is the linguistic style used for casual (informal) communication. It is the most common functional style of speech, the idiom normally employed in conver ...
ly, more general non
specific Specific may refer to: * Specificity (disambiguation) * Specific, a cure or therapy for a specific illness Law * Specific deterrence, focussed on an individual * Specific finding, intermediate verdict used by a jury in determining the fina ...
terms may denote cattle when a singular form is needed. ''Head of cattle'' is usually used only after a numeral. Australian, New Zealand and British farmers use the term ''beast'' or ''cattle beast''. ''Bovine'' is also used in Britain. The term ''critter'' is common in the western United States and Canada, particularly when referring to young cattle. In some areas of the American South (particularly the Appalachian region), where both dairy and beef cattle are present, an individual animal was once called a "beef critter", though that term is becoming
archaic Archaic is a period of time preceding a designated classical period, or something from an older period of time that is also not found or used currently: *List of archaeological periods **Archaic Sumerian language, spoken between 31st - 26th cent ...
.


Other terminology

Cattle raised for human consumption are called '' beef cattle''. Within the beef cattle industry in parts of the United States, the term ''beef'' (plural ''beeves'') is still used in its archaic sense to refer to an animal of either sex. Cows of certain breeds that are kept for the milk they give are called '' dairy cows'' or ''milking cows'' (formerly ''milch cows''). Most young male offspring of dairy cows are sold for
veal Veal is the meat of calves, in contrast to the beef from older cattle. Veal can be produced from a calf of either sex and any breed, however most veal comes from young male calves of dairy breeds which are not used for breeding. Generally, v ...
, and may be referred to as veal calves. The term ''dogies'' is used to describe orphaned calves in the context of
ranch A ranch (from es, rancho/Mexican Spanish) is an area of land, including various structures, given primarily to ranching, the practice of raising grazing livestock such as cattle and sheep. It is a subtype of a farm. These terms are most often ...
work in the
American West The Western United States (also called the American West, the Far West, and the West) is the region comprising the westernmost states of the United States. As American settlement in the U.S. expanded westward, the meaning of the term ''the Wes ...
, as in "Keep them dogies moving". In some places, a cow kept to provide milk for one family is called a "house cow". Other obsolete terms for cattle include "neat" (this use survives in " neatsfoot oil", extracted from the feet and legs of cattle), and "beefing" (young animal fit for slaughter). An
onomatopoeic Onomatopoeia is the process of creating a word that phonetically imitates, resembles, or suggests the sound that it describes. Such a word itself is also called an onomatopoeia. Common onomatopoeias include animal noises such as ''oink'', ''m ...
term for one of the most common sounds made by cattle is ''moo'' (also called ''lowing''). There are a number of other sounds made by cattle, including calves ''bawling'', and bulls ''bellowing''. Bawling is most common for cows after weaning of a calf. The bullroarer makes a sound similar to a bull's territorial call.


Characteristics


Anatomy

Cattle are large quadrupedal
ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s with
cloven hooves A cloven hoof, cleft hoof, divided hoof or split hoof is a hoof split into two toes. This is found on members of the mammalian order Even-toed ungulate, Artiodactyla. Examples of mammals that possess this type of hoof are cattle, deer, pigs, ant ...
. Most breeds have
horns Horns or The Horns may refer to: * Plural of Horn (instrument), a group of musical instruments all with a horn-shaped bells * The Horns (Colorado), a summit on Cheyenne Mountain * ''Horns'' (novel), a dark fantasy novel written in 2010 by Joe Hill ...
, which can be as large as the Texas Longhorn or small like a scur. Careful genetic selection has allowed polled (hornless) cattle to become widespread.


Digestive system

Cattle are ruminants, meaning their
digestive system The human digestive system consists of the gastrointestinal tract plus the accessory organs of digestion (the tongue, salivary glands, pancreas, liver, and gallbladder). Digestion involves the breakdown of food into smaller and smaller compone ...
is highly specialized to allow the consumption of difficult to digest plants as food. Cattle have one stomach with four compartments, the
rumen The rumen, also known as a paunch, is the largest stomach compartment in ruminants and the larger part of the reticulorumen, which is the first chamber in the alimentary canal of ruminant animals. The rumen's microbial favoring environment allow ...
, reticulum, omasum, and abomasum, with the rumen being the largest compartment. The reticulum, the smallest compartment, is known as the "honeycomb". The omasum's main function is to absorb water and nutrients from the digestible feed. The omasum is known as the "many plies". The abomasum is like the human stomach; this is why it is known as the "true stomach". Cattle are known for regurgitating and re-chewing their food, known as
cud Cud is a portion of food that returns from a ruminant's stomach to the mouth to be chewed for the second time. More precisely, it is a bolus of semi-degraded food regurgitated from the reticulorumen of a ruminant. Cud is produced during the phy ...
chewing, like most ruminants. While the animal is feeding, the food is swallowed without being chewed and goes into the rumen for storage until the animal can find a quiet place to continue the digestion process. The food is regurgitated, a mouthful at a time, back up to the mouth, where the food, now called the
cud Cud is a portion of food that returns from a ruminant's stomach to the mouth to be chewed for the second time. More precisely, it is a bolus of semi-degraded food regurgitated from the reticulorumen of a ruminant. Cud is produced during the phy ...
, is chewed by the molars, grinding down the coarse vegetation to small particles. The cud is then swallowed again and further digested by specialized
microorganism A microorganism, or microbe,, ''mikros'', "small") and ''organism'' from the el, ὀργανισμός, ''organismós'', "organism"). It is usually written as a single word but is sometimes hyphenated (''micro-organism''), especially in olde ...
s in the rumen. These microbes are primarily responsible for decomposing
cellulose Cellulose is an organic compound with the formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell wall ...
and other
carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may or ma ...
s into
volatile fatty acids Short-chain fatty acids (SCFAs) are fatty acids with fewer than six carbon atoms. Derived from intestinal microbial fermentation of indigestible foods, SCFAs are the main energy source of colonocytes, making them crucial to gastrointestinal health. ...
cattle use as their primary
metabolic Metabolism (, from el, μεταβολή ''metabolē'', "change") is the set of life-sustaining chemical reactions in organisms. The three main functions of metabolism are: the conversion of the energy in food to energy available to run cell ...
fuel. The microbes inside the rumen also synthesize
amino acid Amino acids are organic compounds that contain both amino and carboxylic acid functional groups. Although hundreds of amino acids exist in nature, by far the most important are the alpha-amino acids, which comprise proteins. Only 22 alpha am ...
s from non-protein
nitrogen Nitrogen is the chemical element with the symbol N and atomic number 7. Nitrogen is a nonmetal and the lightest member of group 15 of the periodic table, often called the pnictogens. It is a common element in the universe, estimated at se ...
ous sources, such as
urea Urea, also known as carbamide, is an organic compound with chemical formula . This amide has two amino groups (–) joined by a carbonyl functional group (–C(=O)–). It is thus the simplest amide of carbamic acid. Urea serves an important r ...
and
ammonia Ammonia is an inorganic compound of nitrogen and hydrogen with the formula . A stable binary hydride, and the simplest pnictogen hydride, ammonia is a colourless gas with a distinct pungent smell. Biologically, it is a common nitrogenous was ...
. As these microbes reproduce in the rumen, older generations die and their cells continue on through the digestive tract. These cells are then partially digested in the small intestines, allowing cattle to gain a high-quality protein source. These features allow cattle to thrive on
grasses Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns and ...
and other tough vegetation.


Gestation and size

The
gestation period In mammals, pregnancy is the period of reproduction during which a female carries one or more live offspring from implantation in the uterus through gestation. It begins when a fertilized zygote implants in the female's uterus, and ends once it ...
for a cow is about nine months long. A newborn calf's size can vary among breeds, but a typical calf weighs . Adult size and weight vary significantly among breeds and sex. Steers are generally slaughtered before reaching . Breeding stock may be allowed a longer lifespan, occasionally living as long as 25 years. The oldest recorded cow, Big Bertha, died at the age of 48 in 1993.


Reproduction

On farms it is very common to use artificial insemination (AI), a medically assisted reproduction technique consisting of the artificial deposition of
semen Semen, also known as seminal fluid, is an organic bodily fluid created to contain spermatozoa. It is secreted by the gonads (sexual glands) and other sexual organs of male or hermaphroditic animals and can fertilize the female ovum. Semen i ...
in the female's genital tract. It is used in cases where the
spermatozoa A spermatozoon (; also spelled spermatozoön; ; ) is a motile sperm cell, or moving form of the haploid cell that is the male gamete. A spermatozoon joins an ovum to form a zygote. (A zygote is a single cell, with a complete set of chromosomes, ...
can not reach the
fallopian tubes The fallopian tubes, also known as uterine tubes, oviducts or salpinges (singular salpinx), are paired tubes in the human female that stretch from the uterus to the ovaries. The fallopian tubes are part of the female reproductive system. In ot ...
or by choice of the owner of the animal. It consists of transferring, to the uterine cavity, spermatozoa previously collected and processed, with the selection of morphologically more normal and mobile spermatozoa. A cow's udder contains two pairs of
mammary gland A mammary gland is an exocrine gland in humans and other mammals that produces milk to feed young offspring. Mammals get their name from the Latin word ''mamma'', "breast". The mammary glands are arranged in organs such as the breasts in primat ...
s, (commonly referred to as ''teats'') creating four "quarters". The front ones are referred to as ''fore quarters'' and the rear ones ''rear quarters''. Synchronization of cattle ovulation to benefit
dairy farming Dairy farming is a class of agriculture for long-term production of milk, which is processed (either on the farm or at a dairy plant, either of which may be called a dairy A dairy is a business enterprise established for the harvesting or ...
may be accomplished via
induced ovulation Ovulation induction is the stimulation of ovulation by medication. It is usually used in the sense of stimulation of the development of ovarian follicles Ovulation Induction Retrieved on Mars 7, 2010 to reverse anovulation or oligoovulation. Scop ...
techniques. The
secondary sex ratio The sex ratio (or gender ratio) is usually defined as the ratio of males to females in a population. As explained by Fisher's principle, for evolutionary reasons this is typically about 1:1 in species which reproduce sexually. Many species devia ...
– the ratio of male to female offspring at birth – is approximately 52:48, although it may be influenced by environmental and other factors. Bulls become fertile at about seven months of age. Their fertility is closely related to the size of their
testicle A testicle or testis (plural testes) is the male reproductive gland or gonad in all bilaterians, including humans. It is homologous to the female ovary. The functions of the testes are to produce both sperm and androgens, primarily testostero ...
s, and one simple test of fertility is to measure the circumference of the scrotum: a young bull is likely to be fertile once this reaches ; that of a fully adult bull may be over . A bull has a fibro-elastic penis. Given the small amount of erectile tissue, there is little enlargement after erection. The penis is quite rigid when non-erect, and becomes even more rigid during erection. Protrusion is not affected much by erection, but more by relaxation of the
retractor penis muscle In some animals the male penis possesses a muscle enabling retraction into the prepuce. In particular animals The retractor penis muscle occurs in marsupials and carnivorans, but it is absent in humans. A stag's penis forms an S-shaped curve w ...
and straightening of the
sigmoid flexure The sigmoid colon (or pelvic colon) is the part of the large intestine that is closest to the rectum and anus. It forms a loop that averages about in length. The loop is typically shaped like a Greek letter sigma (ς) or Latin letter S (thus ''s ...
.


Weight

The weight of adult cattle varies, depending on the breed. Smaller kinds, such as Dexter and Jersey adults, range between . Large Continental breeds, such as Charolais, Marchigiana, Belgian Blue and Chianina adults range from . British breeds, such as Hereford, Angus, and Shorthorn, mature at , occasionally higher, particularly with Angus and Hereford. Bulls are larger than cows of the same breed by up to a few hundred kilograms. British Hereford cows weigh ; the bulls weigh . Chianina bulls can weigh up to ; British bulls, such as Angus and Hereford, can weigh as little as and as much as . The world record for the heaviest bull was , a
Chianina The Chianina () is an Italian breed of large white cattle. It was formerly principally a draught breed; it is now raised mainly for beef. It is the largest and one of the oldest cattle breeds in the world. The ''bistecca alla fiorentina'' is ...
named Donetto, when he was exhibited at the
Arezzo Arezzo ( , , ) , also ; ett, 𐌀𐌓𐌉𐌕𐌉𐌌, Aritim. is a city and ''comune'' in Italy and the capital of the province of the same name located in Tuscany. Arezzo is about southeast of Florence at an elevation of above sea level. ...
show in 1955.Friend, John B., Cattle of the World, Blandford Press, Dorset, 1978 The heaviest steer was eight-year-old 'Old Ben', a Shorthorn/
Hereford Hereford () is a cathedral city, civil parish and the county town of Herefordshire, England. It lies on the River Wye, approximately east of the border with Wales, south-west of Worcester and north-west of Gloucester. With a population ...
cross weighing in at in 1910.McWhirter, Norris & Ross, ''Guinness Book of Records'', Redwood Press, Trowbridge, 1968 In the United States, the average weight of beef cattle has steadily increased, especially since the 1970s, requiring the building of new slaughterhouses able to handle larger carcasses. New packing plants in the 1980s stimulated a large increase in cattle weights. Before 1790 beef cattle averaged only net; and thereafter weights climbed steadily.


Cognition

In laboratory studies, young cattle are able to memorize the locations of several food sources and retain this memory for at least 8 hours, although this declined after 12 hours. Fifteen-month-old heifers learn more quickly than adult cows which have had either one or two calvings, but their longer-term memory is less stable. Mature cattle perform well in spatial learning tasks and have a good long-term memory in these tests. Cattle tested in a
radial arm maze The radial arm maze was designed by Olton and Samuelson in 1976 to measure spatial learning and memory in rats. The original apparatus consists of eight equidistantly spaced arms, each about 4 feet long, and all radiating from a small circular cen ...
are able to remember the locations of high-quality food for at least 30 days. Although they initially learn to avoid low-quality food, this memory diminishes over the same duration. Under less artificial testing conditions, young cattle showed they were able to remember the location of feed for at least 48 days. Cattle can make an association between a visual stimulus and food within 1 day—memory of this association can be retained for 1 year, despite a slight decay. Calves are capable of discrimination learning and adult cattle compare favourably with small mammals in their learning ability in the Closed-field Test. They are also able to discriminate between familiar individuals, and among humans. Cattle can tell the difference between familiar and unfamiliar animals of the same species (conspecifics). Studies show they behave less aggressively toward familiar individuals when they are forming a new group. Calves can also discriminate between humans based on previous experience, as shown by approaching those who handled them positively and avoiding those who handled them aversively. Although cattle can discriminate between humans by their faces alone, they also use other cues such as the color of clothes when these are available. In audio play-back studies, calves prefer their own mother's vocalizations compared to the vocalizations of an unfamiliar mother. In laboratory studies using images, cattle can discriminate between images of the heads of cattle and other animal species. They are also able to distinguish between familiar and unfamiliar conspecifics. Furthermore, they are able to
categorize Categorization is the ability and activity of recognizing shared features or similarities between the elements of the experience of the world (such as objects, events, or ideas), organizing and classifying experience by associating them to a ...
images as familiar and unfamiliar individuals. When mixed with other individuals,
cloned Cloning is the process of producing individual organisms with identical or virtually identical DNA, either by natural or artificial means. In nature, some organisms produce clones through asexual reproduction. In the field of biotechnology, c ...
calves from the same donor form subgroups, indicating that kin discrimination occurs and may be a basis of grouping behaviour. It has also been shown using images of cattle that both artificially inseminated and cloned calves have similar cognitive capacities of kin and non-kin discrimination. Cattle can recognize familiar individuals. Visual individual recognition is a more complex mental process than visual discrimination. It requires the recollection of the learned idiosyncratic identity of an individual that has been previously encountered and the formation of a mental representation. By using two-dimensional images of the heads of one cow (face, profiles, views), all the tested heifers showed individual recognition of familiar and unfamiliar individuals from their own breed. Furthermore, almost all the heifers recognized unknown individuals from different breeds, although this was achieved with greater difficulty. Individual recognition was most difficult when the visual features of the breed being tested were quite different from the breed in the image, for example, the breed being tested had no spots whereas the image was of a spotted breed. Cattle use visual/brain lateralisation in their visual scanning of novel and familiar stimuli. Domestic cattle prefer to view novel stimuli with the left eye, i.e. using the right brain hemisphere (similar to horses, Australian magpies, chicks, toads and fish) but use the right eye, i.e. using the left hemisphere, for viewing familiar stimuli.


Temperament and emotions

In cattle, temperament can affect production traits such as carcass and meat quality or milk yield as well as affecting the animal's overall health and reproduction. Cattle temperament is defined as "the consistent behavioral and physiological difference observed between individuals in response to a stressor or environmental challenge and is used to describe the relatively stable difference in the behavioral predisposition of an animal, which can be related to psychobiological mechanisms". Generally, cattle temperament is assumed to be multidimensional. Five underlying categories of temperament traits have been proposed: * shyness–boldness * exploration–avoidance * activity * aggressiveness * sociability In a study on Holstein–Friesian heifers learning to press a panel to open a gate for access to a food reward, the researchers also recorded the heart rate and behavior of the heifers when moving along the race towards the food. When the heifers made clear improvements in learning, they had higher heart rates and tended to move more vigorously along the race. The researchers concluded this was an indication that cattle may react emotionally to their own learning improvement. Negative emotional states are associated with a bias toward negative responses towards ambiguous cues in judgement tasks. After separation from their mothers, Holstein calves showed such a cognitive bias indicative of low mood. A similar study showed that after hot-iron disbudding (
dehorning Dehorning is the process of removing the horns of livestock. Cattle, sheep, and goats are sometimes dehorned for economic and safety reasons. Disbudding is a different process with similar results; it cauterizes and thus destroys horn buds before ...
), calves had a similar negative bias indicating that post-operative pain following this routine procedure results in a negative change in emotional state. In studies of visual discrimination, the position of the ears has been used as an indicator of emotional state. When cattle are stressed other cattle can tell by the chemicals released in their urine. Cattle are very
gregarious Sociality is the degree to which individuals in an animal population tend to associate in social groups (gregariousness) and form cooperative societies. Sociality is a survival response to evolutionary pressures. For example, when a mother wasp ...
and even short-term isolation is considered to cause severe psychological stress. When
Aubrac Aubrac is a small village in the southern Massif Central of France. The name is also applied to the surrounding countryside, which is properly called L'Aubrac in French. The Aubrac region has been a member of the Natura 2000 network since August 200 ...
and Friesian heifers are isolated, they increase their vocalizations and experience increased heart rate and plasma
cortisol Cortisol is a steroid hormone, in the glucocorticoid class of hormones. When used as a medication, it is known as hydrocortisone. It is produced in many animals, mainly by the ''zona fasciculata'' of the adrenal cortex in the adrenal gland ...
concentrations. These physiological changes are greater in Aubracs. When visual contact is re-instated, vocalizations rapidly decline, regardless of the familiarity of the returning cattle, however, heart rate decreases are greater if the returning cattle are familiar to the previously isolated individual. Mirrors have been used to reduce stress in isolated cattle.


Senses

Cattle use all of the five widely recognized sensory modalities. These can assist in some complex behavioural patterns, for example, in grazing behaviour. Cattle eat mixed diets, but when given the opportunity, show a partial preference of approximately 70% clover and 30% grass. This preference has a diurnal pattern, with a stronger preference for clover in the morning, and the proportion of grass increasing towards the evening.


Vision

Vision is the dominant sense in cattle and they obtain almost 50% of their information visually. Cattle are a prey animal and to assist predator detection, their eyes are located on the sides of their head rather than the front. This gives them a wide field of view of 330° but limits binocular vision (and therefore stereopsis) to 30° to 50° compared to 140° in humans. This means they have a blind spot directly behind them. Cattle have good visual acuity, but compared to humans, their visual accommodation is poor. Cattle have two kinds of
color receptors Cone cells, or cones, are photoreceptor cells in the retinas of vertebrate eyes including the human eye. They respond differently to light of different wavelengths, and the combination of their responses is responsible for color vision. Cones f ...
in the
cone cells Cone cells, or cones, are photoreceptor cells in the retinas of vertebrate eyes including the human eye. They respond differently to light of different wavelengths, and the combination of their responses is responsible for color vision. Cones ...
of their
retina The retina (from la, rete "net") is the innermost, light-sensitive layer of tissue of the eye of most vertebrates and some molluscs. The optics of the eye create a focused two-dimensional image of the visual world on the retina, which then ...
s. This means that cattle are dichromatic, as are most other non-primate land mammals. There are two to three rods per cone in the
fovea centralis The fovea centralis is a small, central pit composed of closely packed cones in the eye. It is located in the center of the macula lutea of the retina. The fovea is responsible for sharp central vision (also called foveal vision), which is nec ...
but five to six near the optic papilla. Cattle can distinguish long wavelength colors (yellow, orange and red) much better than the shorter wavelengths (blue, grey and green). Calves are able to discriminate between long (red) and short (blue) or medium (green) wavelengths, but have limited ability to discriminate between the short and medium. They also approach handlers more quickly under red light. Whilst having good color sensitivity, it is not as good as humans or
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus ''Ovis'', in everyday usage it almost always refers to domesticated s ...
. A common misconception about cattle (particularly bulls) is that they are enraged by the color red (something provocative is often said to be "like a red flag to a bull"). This is a myth. In
bullfighting Bullfighting is a physical contest that involves a bullfighter attempting to subdue, immobilize, or kill a bull, usually according to a set of rules, guidelines, or cultural expectations. There are several variations, including some forms wh ...
, it is the movement of the red flag or cape that irritates the bull and incites it to charge.


Taste

Cattle have a well-developed sense of taste and can distinguish the four primary tastes (sweet, salty, bitter and sour). They possess around 20,000 taste buds. The strength of taste perception depends on the individual's current food requirements. They avoid bitter-tasting foods (potentially toxic) and have a marked preference for sweet (high calorific value) and salty foods (
electrolyte An electrolyte is a medium containing ions that is electrically conducting through the movement of those ions, but not conducting electrons. This includes most soluble salts, acids, and bases dissolved in a polar solvent, such as water. Upon dis ...
balance). Their sensitivity to sour-tasting foods helps them to maintain optimal ruminal pH. Plants have low levels of sodium and cattle have developed the capacity of seeking salt by taste and smell. If cattle become depleted of sodium salts, they show increased locomotion directed to searching for these. To assist in their search, the olfactory and gustatory receptors able to detect minute amounts of sodium salts increase their sensitivity as biochemical disruption develops with sodium salt depletion.


Hearing

Cattle hearing ranges from 23 Hz to 35 kHz. Their frequency of best sensitivity is 8 kHz and they have a lowest threshold of −21 db (re 20 μN/m−2), which means their hearing is more acute than horses (lowest threshold of 7 db). Sound localization acuity thresholds are an average of 30°. This means that cattle are less able to localise sounds compared to goats (18°), dogs (8°) and humans (0.8°). Because cattle have a broad foveal fields of view covering almost the entire horizon, they may not need very accurate locus information from their auditory systems to direct their gaze to a sound source. Vocalizations are an important mode of communication amongst cattle and can provide information on the age, sex, dominance status and reproductive status of the caller. Calves can recognize their mothers using vocalizations; vocal behaviour may play a role by indicating estrus and competitive display by bulls.


Olfaction and gustation

Cattle have a range of odiferous glands over their body including interdigital, infraorbital, inguinal and
sebaceous A sebaceous gland is a microscopic exocrine gland in the skin that opens into a hair follicle to secrete an oily or waxy matter, called sebum, which lubricates the hair and skin of mammals. In humans, sebaceous glands occur in the greatest number ...
glands, indicating that olfaction probably plays a large role in their social life. Both the primary olfactory system using the
olfactory bulb The olfactory bulb (Latin: ''bulbus olfactorius'') is a grey matter, neural structure of the vertebrate forebrain involved in olfaction, the sense of odor, smell. It sends olfactory information to be further processed in the amygdala, the orbitof ...
s, and the secondary olfactory system using the
vomeronasal organ The vomeronasal organ (VNO), or Jacobson's organ, is the paired auxiliary olfactory (smell) sense organ located in the soft tissue of the nasal septum, in the nasal cavity just above the roof of the mouth (the hard palate) in various tetrapods. ...
are used. This latter olfactory system is used in the
flehmen The flehmen response (; from German ''flehmen'', to bare the upper teeth, and Upper Saxon German ''flemmen'', to look spiteful), also called the flehmen position, flehmen reaction, flehmen grimace, flehming, or flehmening, is a behavior in which ...
response. There is evidence that when cattle are stressed, this can be recognised by other cattle and this is communicated by alarm substances in the urine. The odour of dog faeces induces behavioural changes prior to cattle feeding, whereas the odours of urine from either stressed or non-stressed conspecifics and blood have no effect. In the laboratory, cattle can be trained to recognise conspecific individuals using olfaction only. In general, cattle use their sense of smell to "expand" on information detected by other sensory modalities. However, in the case of social and reproductive behaviours, olfaction is a key source of information.


Touch

Cattle have tactile sensations detected mainly by
mechanoreceptor A mechanoreceptor, also called mechanoceptor, is a sensory receptor that responds to mechanical pressure or distortion. Mechanoreceptors are innervated by sensory neurons that convert mechanical pressure into electrical signals that, in animals, ...
s,
thermoreceptor A thermoreceptor is a non-specialised sense receptor, or more accurately the receptive portion of a sensory neuron, that codes absolute and relative changes in temperature, primarily within the innocuous range. In the mammalian peripheral nervous s ...
s and nociceptors in the skin and muscles. These are used most frequently when cattle explore their environment.


Magnetoreception

There is conflicting evidence for
magnetoreception Magnetoreception is a sense which allows an organism to detect the Earth's magnetic field. Animals with this sense include some arthropods, molluscs, and vertebrates (fish, amphibians, reptiles, birds, and mammals, though not humans). The se ...
in cattle. One study reported that resting and grazing cattle tend to align their body axes in the geomagnetic north–south direction. In a follow-up study, cattle exposed to various magnetic fields directly beneath or in the vicinity of power lines trending in various magnetic directions exhibited distinct patterns of alignment. However, in 2011, a group of Czech researchers reported their failed attempt to replicate the finding using
Google Earth Google Earth is a computer program that renders a 3D computer graphics, 3D representation of Earth based primarily on satellite imagery. The program maps the Earth by superimposition, superimposing satellite images, aerial photography, and geog ...
images.


Behavior

Under natural conditions, calves stay with their mother until weaning at 8 to 11 months. Heifer and bull calves are equally attached to their mothers in the first few months of life. Cattle are considered to be "hider" type animals, utilizing secluded areas more in the hours before calving and continued to use it more for the hour after calving. Cows that gave birth for the first time show a higher incidence of abnormal maternal behavior. In one study, beef-calves reared on the range were observed to suckle an average of 5.0 times every 24 hours with an average total time of 46 min spent suckling. There was a diurnal rhythm in suckling activity with peaks between 05:00–07:00, 10:00–13:00 and 17:00–21:00.


Reproductive behavior

Semi-wild Highland cattle heifers first give birth at 2 or 3 years of age, and the timing of birth is synchronized with increases in natural food quality. Average calving interval is 391 days, and calving mortality within the first year of life is 5%.


Dominance and leadership

One study showed that over a 4-year period, dominance relationships within a herd of semi-wild highland cattle were very firm. There were few overt aggressive conflicts and the majority of disputes were settled by
agonistic An agonist is a chemical that activates a receptor to produce a biological response. Receptors are cellular proteins whose activation causes the cell to modify what it is currently doing. In contrast, an antagonist blocks the action of the agoni ...
(non-aggressive, competitive) behaviors that involved no physical contact between opponents (e.g. threatening and spontaneous withdrawing). Such agonistic behavior reduces the risk of injury. Dominance status depended on age and sex, with older animals generally being dominant to young ones and males dominant to females. Young bulls gained superior dominance status over adult cows when they reached about 2 years of age. As with many animal dominance hierarchies, dominance-associated aggressiveness does not correlate with rank position, but is closely related to rank distance between individuals. Dominance is maintained in several ways. Cattle often engage in mock fights where they test each other's strength in a non-aggressive way. Licking is primarily performed by subordinates and received by dominant animals. Mounting is a playful behavior shown by calves of both sexes and by bulls and sometimes by cows in estrus, however, this is not a dominance related behavior as has been found in other species. The horns of cattle are "
honest signals Unconscious (or intuitive) communication is the subtle, unintentional, unconscious cues that provide information to another individual. It can be verbal (speech patterns, physical activity while speaking, or the tone of voice of an individual) or ...
" used in mate selection. Furthermore, horned cattle attempt to keep greater distances between themselves and have fewer physical interactions than hornless cattle. This leads to more stable social relationships. In calves, the frequency of agonistic behavior decreases as space allowance increases, but this does not occur for changes in group size. However, in adult cattle, the number of agonistic encounters increases as the group size increases.


Grazing behavior

When grazing, cattle vary several aspects of their bite, i.e. tongue and jaw movements, depending on characteristics of the plant they are eating. Bite area decreases with the density of the plants but increases with their height. Bite area is determined by the sweep of the tongue; in one study observing steers, bite area reached a maximum of approximately . Bite depth increases with the height of the plants. By adjusting their behavior, cattle obtain heavier bites in swards that are tall and sparse compared with short, dense swards of equal mass/area. Cattle adjust other aspects of their grazing behavior in relation to the available food; foraging velocity decreases and intake rate increases in areas of abundant palatable forage. Cattle avoid grazing areas contaminated by the faeces of other cattle more strongly than they avoid areas contaminated by sheep, but they do not avoid pasture contaminated by rabbit faeces.


Genetics

On 24 April 2009, edition of the journal ''Science'', a team of researchers led by the National Institutes of Health and the US Department of Agriculture reported having mapped the
bovine genome The genome of a female Hereford cow was published in 2009. It was sequenced by the Bovine Genome Sequencing and Analysis Consortium, a team of researchers led by the National Institutes of Health and the U.S. Department of Agriculture. It was par ...
. The scientists found cattle have about 22,000 genes, and 80% of their genes are shared with humans, and they share about 1000 genes with dogs and rodents, but are not found in humans. Using this bovine "HapMap", researchers can track the differences between the breeds that affect the quality of meat and milk yields. Behavioral traits of cattle can be as heritable as some production traits, and often, the two can be related. The heritability of fear varies markedly in cattle from low (0.1) to high (0.53); such high variation is also found in pigs and sheep, probably due to differences in the methods used. The heritability of temperament (response to isolation during handling) has been calculated as 0.36 and 0.46 for habituation to handling. Rangeland assessments show that the heritability of aggressiveness in cattle is around 0.36. Quantitative trait loci (QTLs) have been found for a range of production and behavioral characteristics for both dairy and beef cattle.


Domestication and husbandry

Cattle occupy a unique role in
human history Human history, also called world history, is the narrative of humanity's past. It is understood and studied through anthropology, archaeology, genetics, and linguistics. Since the invention of writing, human history has been studied throug ...
, having been domesticated since at least the early
neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several parts ...
age. Archaeozoological and genetic data indicate that cattle were first domesticated from wild
aurochs The aurochs (''Bos primigenius'') ( or ) is an extinct cattle species, considered to be the wild ancestor of modern domestic cattle. With a shoulder height of up to in bulls and in cows, it was one of the largest herbivores in the Holocen ...
(''Bos primigenius'') approximately 10,500 years ago. There were two major areas of domestication: one in the
Near East The ''Near East''; he, המזרח הקרוב; arc, ܕܢܚܐ ܩܪܒ; fa, خاور نزدیک, Xāvar-e nazdik; tr, Yakın Doğu is a geographical term which roughly encompasses a transcontinental region in Western Asia, that was once the hist ...
(specifically
central Anatolia The Central Anatolia Region ( tr, İç Anadolu Bölgesi) is a geographical region of Turkey. The largest city in the region is Ankara. Other big cities are Konya, Kayseri, Eskişehir, Sivas, and Aksaray. Located in Central Turkey, it is bordered ...
, the
Levant The Levant () is an approximate historical geographical term referring to a large area in the Eastern Mediterranean region of Western Asia. In its narrowest sense, which is in use today in archaeology and other cultural contexts, it is eq ...
and
Western Iran Western Iran consists of Armenian Highlands, Northern Zagros and the rich agricultural area of the Khuzestan Plain in the south. It includes the provinces of Kordestan, Kermanshah, Ilam, Hamadan and Lorestan. Some references also count West ...
), giving rise to the taurine line, and a second in the area that is now Pakistan, resulting in the indicine line. Modern mitochondrial DNA variation indicates the taurine line may have arisen from as few as 80
aurochs The aurochs (''Bos primigenius'') ( or ) is an extinct cattle species, considered to be the wild ancestor of modern domestic cattle. With a shoulder height of up to in bulls and in cows, it was one of the largest herbivores in the Holocen ...
tamed in the upper reaches of
Mesopotamia Mesopotamia ''Mesopotamíā''; ar, بِلَاد ٱلرَّافِدَيْن or ; syc, ܐܪܡ ܢܗܪ̈ܝܢ, or , ) is a historical region of Western Asia situated within the Tigris–Euphrates river system, in the northern part of the F ...
near the villages of
Çayönü Tepesi Çayönü Tepesi is a Neolithic settlement in southeastern Turkey which prospered from circa 8,630 to 6,800 BC. It is located forty kilometres north-west of Diyarbakır, at the foot of the Taurus mountains. It lies near the Boğazçay, a tribu ...
in what is now southeastern Turkey and
Dja'de el-Mughara Dja'de el-Mughara is a Pre-Pottery Neolithic settlement in Northern Syria dating back to approximately 9000 BC. The settlement is located about 100 km northeast of the city of Aleppo and was being excavated from 1991 until 2011 when work on the ...
in what is now northern Syria. Although European cattle are largely descended from the taurine lineage, gene flow from African cattle (partially of indicine origin) contributed substantial genomic components to both southern European cattle breeds and their New World descendants. A study on 134 breeds showed that modern taurine cattle originated from Africa, Asia, North and South America, Australia, and Europe. Some researchers have suggested that African taurine cattle are derived from a third independent domestication from North African aurochsen.


Usage as money

As early as 9000 BC both grain and cattle were used as money or as ''barter'' (the '' first grain remains'' found, considered to be evidence of pre-agricultural practice date to 17,000 BC). Some evidence also exists to suggest that other animals, such as camels and goats, may have been used as currency in some parts of the world. One of the advantages of using cattle as currency is that it allows the seller to set a fixed price. It even created the standard pricing. For example, two chickens were traded for one cow as cows were deemed to be more valuable than chickens.


Modern husbandry

Cattle are often raised by allowing herds to graze on the grasses of large tracts of
rangeland Rangelands are grasslands, shrublands, woodlands, wetlands, and deserts that are grazed by domestic livestock or wild animals. Types of rangelands include tallgrass and shortgrass prairies, desert grasslands and shrublands, woodlands, savannas ...
. Raising cattle in this manner allows the use of land that might be unsuitable for growing crops. The most common interactions with cattle involve daily
feeding Eating (also known as consuming) is the ingestion of food, typically to provide a heterotrophic organism with energy and to allow for growth. Animals and other heterotrophs must eat in order to survive — carnivores eat other animals, herbi ...
, cleaning and
milking Milking is the act of removing milk from the mammary glands of cattle, water buffalo, humans, goats, sheep, and, more rarely, camels, horses and donkeys. Milking may be done by hand or by machine, and requires the animal to be currently or recen ...
. Many routine husbandry practices involve
ear tag An ear tag is a plastic or metal object used for identification of domestic livestock and other animals. If the ear tag uses Radio Frequency Identification Device (RFID) technology it is referred to as an electronic ear tag. Electronic ear tags ...
ging,
dehorning Dehorning is the process of removing the horns of livestock. Cattle, sheep, and goats are sometimes dehorned for economic and safety reasons. Disbudding is a different process with similar results; it cauterizes and thus destroys horn buds before ...
, loading, medical operations, artificial insemination, vaccinations and hoof care, as well as training for agricultural shows and preparations. Also, some
cultural differences Cultural diversity is the quality of diverse or different cultures, as opposed to monoculture, the global monoculture, or a homogenization of cultures, akin to cultural evolution. The term "cultural diversity" can also refer to having different cu ...
occur in working with cattle; the cattle husbandry of Fulani men rests on behavioural techniques, whereas in Europe, cattle are controlled primarily by physical means, such as fences. Breeders use cattle husbandry to reduce ''M. bovis'' infection susceptibility by
selective breeding Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant mal ...
and maintaining herd health to avoid concurrent disease. Cattle are farmed for beef, veal, dairy, and leather. They are less commonly used for conservation grazing, or to maintain grassland for wildlife, such as in Epping Forest, England. They are often used in some of the most wild places for livestock. Depending on the breed, cattle can survive on hill grazing, heaths, marshes, moors and semidesert. Modern cattle are more commercial than older breeds and, having become more specialized, are less versatile. For this reason, many smaller farmers still favor old breeds, such as the
Jersey Jersey ( , ; nrf, Jèrri, label=Jèrriais ), officially the Bailiwick of Jersey (french: Bailliage de Jersey, links=no; Jèrriais: ), is an island country and self-governing Crown Dependencies, Crown Dependency near the coast of north-west F ...
dairy breed. In Portugal, Spain,
southern France Southern France, also known as the South of France or colloquially in French language, French as , is a defined geographical area consisting of the regions of France that border the Atlantic Ocean south of the Marais Poitevin,Louis Papy, ''Le midi ...
and some Latin American countries, bulls are used in the activity of
bullfighting Bullfighting is a physical contest that involves a bullfighter attempting to subdue, immobilize, or kill a bull, usually according to a set of rules, guidelines, or cultural expectations. There are several variations, including some forms wh ...
; In many other countries bullfighting is illegal. Other activities such as bull riding are seen as part of a
rodeo Rodeo () is a competitive equestrian sport that arose out of the working practices of cattle herding in Spain and Mexico, expanding throughout the Americas and to other nations. It was originally based on the skills required of the working va ...
, especially in North America.
Bull-leaping Bull-leaping ( grc, ταυροκαθάψια, ) is a term for various types of non-violent bull fighting. Some are based on an ancient ritual from the Minoan civilization involving an acrobat leaping over the back of a charging bull (or cow). ...
, a central ritual in
Bronze Age The Bronze Age is a historic period, lasting approximately from 3300 BC to 1200 BC, characterized by the use of bronze, the presence of writing in some areas, and other early features of urban civilization. The Bronze Age is the second pri ...
Minoan The Minoan civilization was a Bronze Age Aegean civilization on the island of Crete and other Aegean Islands, whose earliest beginnings were from 3500BC, with the complex urban civilization beginning around 2000BC, and then declining from 1450B ...
culture (see Sacred Bull), still exists in southwestern France. In modern times, cattle are also entered into agricultural competitions. These competitions can involve live cattle or cattle carcases in hoof and hook events. In terms of food intake by humans, consumption of cattle is less efficient than of grain or vegetables with regard to
land use Land use involves the management and modification of natural environment or wilderness into built environment such as settlements and semi-natural habitats such as arable fields, pastures, and managed woods. Land use by humans has a long h ...
, and hence cattle grazing consumes more area than such other agricultural production when raised on grains.Edward O. Wilson, ''The Future of Life'', 2003, Vintage Books, 256 pages Nonetheless, cattle and other forms of domesticated animals can sometimes help to use plant resources in areas not easily amenable to other forms of agriculture. Bulls are sometimes used as guard animals. In occasional cases, cattle are kept as pets, and pet cows often have sweet temperaments, enjoying being petted and "kissing" (licking) their owners. But there are costs to keeping them as pets that limit how many people can practically do so; not everyone has space or facilities for a large-animal pet, and some amount of resources are needed to keep one humanely (such as pasture, hay, feed, water, and large-animal veterinary care). In addition, because livestock animals are gregarious, they need at least one companion to avoid being stressed or lonely, so keeping bovine, caprine, or ovine pets requires more than one animal. Most pet cows live on farms that have other livestock anyway, as the
marginal cost In economics, the marginal cost is the change in the total cost that arises when the quantity produced is incremented, the cost of producing additional quantity. In some contexts, it refers to an increment of one unit of output, and in others it r ...
of one or two more animals is then not very large. Farmers have traditionally often been averse to making pets out of livestock, on the principle that each animal must pay its way somehow if the farm is to survive financially, and also because there are sufficient opportunities for moments of petting and animal appreciation among the herd anyway, even when none of them are pets per se.


Sleep

The average sleep time of a domestic cow is about 4 hours a day. Cattle do have a
stay apparatus The stay apparatus is a group of ligaments, tendons and muscles which "lock" major joints in the limbs of the horse. It is best known as the mechanism by which horses can enter a light sleep while still standing up. It does, however, exist in oth ...
, but do not sleep standing up; they lie down to sleep deeply. In spite of the
urban legend An urban legend (sometimes contemporary legend, modern legend, urban myth, or urban tale) is a genre of folklore comprising stories or fallacious claims circulated as true, especially as having happened to a "friend of a friend" or a family m ...
, cows cannot be tipped over by people pushing on them.


Economy

The meat of adult cattle is known as
beef Beef is the culinary name for meat from cattle (''Bos taurus''). In prehistoric times, humankind hunted aurochs and later domesticated them. Since that time, numerous breeds of cattle have been bred specifically for the quality or quantity ...
, and that of calves is
veal Veal is the meat of calves, in contrast to the beef from older cattle. Veal can be produced from a calf of either sex and any breed, however most veal comes from young male calves of dairy breeds which are not used for breeding. Generally, v ...
. Other animal parts are also used as food products, including blood,
liver The liver is a major Organ (anatomy), organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the Protein biosynthesis, synthesis of proteins and biochemicals necessary for ...
,
kidney The kidneys are two reddish-brown bean-shaped organs found in vertebrates. They are located on the left and right in the retroperitoneal space, and in adult humans are about in length. They receive blood from the paired renal arteries; blood ...
,
heart The heart is a muscular organ in most animals. This organ pumps blood through the blood vessels of the circulatory system. The pumped blood carries oxygen and nutrients to the body, while carrying metabolic waste such as carbon dioxide t ...
and oxtail. Cattle also produce
milk Milk is a white liquid food produced by the mammary glands of mammals. It is the primary source of nutrition for young mammals (including breastfed human infants) before they are able to digestion, digest solid food. Immune factors and immune ...
, and dairy cattle are specifically bred to produce the large quantities of milk processed and sold for human consumption. Cattle today are the basis of a multibillion-dollar industry worldwide. The
international trade International trade is the exchange of capital, goods, and services across international borders or territories because there is a need or want of goods or services. (see: World economy) In most countries, such trade represents a significant ...
in beef for 2000 was over $30 billion and represented only 23% of world beef production. Approximately 300 million cattle, including dairy cattle, are slaughtered each year for food. The production of milk, which is also made into cheese, butter,
yogurt Yogurt (; , from tr, yoğurt, also spelled yoghurt, yogourt or yoghourt) is a food produced by bacterial Fermentation (food), fermentation of milk. The bacteria used to make yogurt are known as ''yogurt cultures''. Fermentation of sugars in t ...
, and other dairy products, is comparable in economic size to beef production, and provides an important part of the food supply for many of the world's people. Cattle hides, used for leather to make shoes,
couch A couch, also known as a sofa, settee, or chesterfield, is a cushioned item of furniture for seating multiple people (although it is not uncommon for a single person to use a couch alone). It is commonly found in the form of a bench with uph ...
es and clothing, are another widespread product. Cattle remain broadly used as draft animals in many
developing countries A developing country is a sovereign state with a lesser developed industrial base and a lower Human Development Index (HDI) relative to other countries. However, this definition is not universally agreed upon. There is also no clear agreem ...
, such as India. Cattle are also used in some sporting games, including
rodeo Rodeo () is a competitive equestrian sport that arose out of the working practices of cattle herding in Spain and Mexico, expanding throughout the Americas and to other nations. It was originally based on the skills required of the working va ...
and
bullfighting Bullfighting is a physical contest that involves a bullfighter attempting to subdue, immobilize, or kill a bull, usually according to a set of rules, guidelines, or cultural expectations. There are several variations, including some forms wh ...
.


Cattle meat production

Source: Helgi Library,, Cattle Meat Production , 12 February 2014 World Bank, FAOSTAT About a quarter of the world's meat comes from cattle.


Dairy

Certain breeds of cattle, such as the
Holstein-Friesian Holstein Friesians (often shortened to Holsteins in North America, while the term Friesians is often used in the UK and Ireland) are a breed of dairy cattle that originated in the Dutch provinces of North Holland and Friesland, and Schleswig-Hols ...
, are used to produce
milk Milk is a white liquid food produced by the mammary glands of mammals. It is the primary source of nutrition for young mammals (including breastfed human infants) before they are able to digestion, digest solid food. Immune factors and immune ...
, which can be processed into dairy products such as milk,
cheese Cheese is a dairy product produced in wide ranges of flavors, textures, and forms by coagulation of the milk protein casein. It comprises proteins and fat from milk, usually the milk of cows, buffalo, goats, or sheep. During production, ...
or
yogurt Yogurt (; , from tr, yoğurt, also spelled yoghurt, yogourt or yoghourt) is a food produced by bacterial Fermentation (food), fermentation of milk. The bacteria used to make yogurt are known as ''yogurt cultures''. Fermentation of sugars in t ...
. Dairy cattle are usually kept on specialized dairy farms designed for milk production. Most cows are milked twice per day, with milk processed at a dairy, which may be onsite at the farm or the milk may be shipped to a dairy plant for eventual sale of a dairy product. Lactation is induced in heifers and spayed cows by a combination of physical and psychological stimulation, by drugs, or by a combination of those methods. For mother cows to continue producing milk, they give birth to one calf per year. If the calf is male, it generally is slaughtered at a young age to produce
veal Veal is the meat of calves, in contrast to the beef from older cattle. Veal can be produced from a calf of either sex and any breed, however most veal comes from young male calves of dairy breeds which are not used for breeding. Generally, v ...
. They will continue to produce milk until three weeks before birth. Over the last fifty years, dairy farming has become more intensive to increase the yield of milk produced by each cow. The
Holstein-Friesian Holstein Friesians (often shortened to Holsteins in North America, while the term Friesians is often used in the UK and Ireland) are a breed of dairy cattle that originated in the Dutch provinces of North Holland and Friesland, and Schleswig-Hols ...
is the breed of dairy cow most common in the UK, Europe and the United States. It has been bred selectively to produce the highest yields of milk of any cow. Around 22 litres per day is average in the UK.


Hides

Most cattle are not kept solely for hides, which are usually a by-product of beef production. Hides are most commonly used for leather, which can be made into a variety of products, including shoes. In 2012 India was the world's largest producer of cattle hides.


Feral cattle

Feral cattle are defined as being 'cattle that are not domesticated or cultivated'. Populations of feral cattle are known to come from and exist in: Australia, United States of America, Colombia, Argentina, Spain, France and many islands, including
New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...
,
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
, Galapagos,
Juan Fernández Islands The Juan Fernández Islands ( es, Archipiélago Juan Fernández) are a sparsely inhabited series of islands in the South Pacific Ocean reliant on tourism and fishing. Situated off the coast of Chile, they are composed of three main volcanic i ...
,
Hispaniola Hispaniola (, also ; es, La Española; Latin and french: Hispaniola; ht, Ispayola; tnq, Ayiti or Quisqueya) is an island in the Caribbean that is part of the Greater Antilles. Hispaniola is the most populous island in the West Indies, and th ...
(Dominican Republic and Haiti),
Tristan da Cunha Tristan da Cunha (), colloquially Tristan, is a remote group of volcanic islands in the South Atlantic Ocean. It is the most remote inhabited archipelago in the world, lying approximately from Cape Town in South Africa, from Saint Helena ...
and
Île Amsterdam Île Amsterdam (), also known as Amsterdam Island and New Amsterdam (''Nouvelle-Amsterdam''), is an island of the French Southern and Antarctic Lands in the southern Indian Ocean that together with neighbouring Île Saint-Paul to the south for ...
, two islands of Kuchinoshima and Kazura Island next to Naru Island in Japan.
Chillingham cattle Chillingham cattle, also known as Chillingham wild cattle, are a breed of cattle that live in a large enclosed park at Chillingham Castle, Northumberland, England. In summer 2022 the cattle numbers 138 animals with approximately equal numbers o ...
is sometimes regarded as a feral breed. Aleutian wild cattles can be found on Aleutian Islands. The "Kinmen cattle" which are dominantly found on Kinmen Island, Taiwan is mostly domesticated while smaller portion of the population is believed to live in the wild due to accidental releases. Other notable examples include cattle in the vicinity of Hong Kong (in the Shing Mun Country Park, among Sai Kung District and Lantau Island and on Grass Island, Hong Kong, Grass Island), and semi-feral animals in Yangmingshan, Taiwan.


Environmental impact

Gut flora in cattle include methanogens that methanogenesis, produce methane as a byproduct of enteric fermentation, which cattle belch out. The same volume of atmospheric methane has a 72x higher (over 20 years) global warming potential than Carbon dioxide in Earth's atmosphere, atmospheric carbon dioxide. See Table 8.7. Methane belching from cattle can be reduced with Selective Breeding, genetic selection, immunization against the many methanogens, rumen defaunation (killing the bacteria-killing protozoa), diet modification (e.g. seaweed fortification), decreased antibiotic use, and grazing management, among others. A 2013 report from the
Food and Agriculture Organization The Food and Agriculture Organization of the United Nations (FAO)french: link=no, Organisation des Nations unies pour l'alimentation et l'agriculture; it, Organizzazione delle Nazioni Unite per l'Alimentazione e l'Agricoltura is an intern ...
(FAO) based on 2005 data states that the livestock sector is responsible for 14.5% of greenhouse gas emissions, 65% of which is due to cattle. The IPCC estimates that cattle and other livestock emit about 80 to 93 Megatonnes of methane per year, accounting for an estimated 37% of anthropogenic methane emissions,Steinfeld, H. et al. 2006,
Livestock's Long Shadow: Environmental Issues and Options
Livestock, Environment and Development, FAO.
and additional methane is produced by anaerobic fermentation of manure in manure lagoons and other manure storage structures. Another estimate is 12% of global GHG. While cattle fed forage actually produce more methane than grain-fed cattle, the increase may be offset by the increased carbon recapture of pastures, which recapture three times the CO2 of cropland used for grain. One of the cited changes suggested to reduce greenhouse gas emissions is intensification of the livestock industry, since intensification leads to less land for a given level of production. This assertion is supported by studies of the US beef production system, suggesting practices prevailing in 2007 involved 8.6% less fossil fuel use, 16.3% less greenhouse gas emissions, 12.1% less water use, and 33.0% less land use, per unit mass of beef produced, than those used in 1977. The analysis took into account not only practices in feedlots, but also feed production (with less feed needed in more intensive production systems), forage-based cow-calf operations and back-grounding before cattle enter a feedlot (with more beef produced per head of cattle from those sources, in more intensive systems), and beef from animals derived from the dairy industry. A more controversial suggestion, advocated by George Monbiot in the documentary "Apocalypse Cow", is to stop farming cattle completely, however farmers often have political power so might be able to resist such a big change. Significant numbers of dairy, as well as beef cattle, are confined in concentrated animal feeding operations (CAFOs), defined as "new and existing operations which stable or confine and feed or maintain for a total of 45 days or more in any 12-month period more than the number of animals specified" where "[c]rops, vegetation, forage growth, or post-harvest residues are not sustained in the normal growing season over any portion of the lot or facility." They may be designated as small, medium and large. Such designation of cattle CAFOs is according to cattle type (mature dairy cows, veal calves or other) and cattle numbers, but medium CAFOs are so designated only if they meet certain discharge criteria, and small CAFOs are designated only on a case-by-case basis. A CAFO that discharges pollutants is required to obtain a permit, which requires a plan to manage nutrient runoff, manure, chemicals, contaminants, and other wastewater pursuant to the US Clean Water Act. The regulations involving CAFO permitting have been extensively litigated. Commonly, CAFO wastewater and manure nutrients are applied to land at agronomic rates for use by forages or crops, and it is often assumed that various constituents of wastewater and manure, e.g. organic contaminants and pathogens, will be retained, inactivated or degraded on the land with application at such rates; however, additional evidence is needed to test reliability of such assumptions . Concerns raised by opponents of CAFOs have included risks of contaminated water due to feedlot runoff, soil erosion, human and animal exposure to toxic chemicals, development of antibiotic resistant bacteria and an increase in ''E. coli'' contamination. While research suggests some of these impacts can be mitigated by developing wastewater treatment systems and planting cover crops in larger setback zones, the Union of Concerned Scientists released a report in 2008 concluding that CAFOs are generally unsustainable and Cost externalizing, externalize costs. Another concern is manure, which if not well-managed, can lead to adverse environmental consequences. However, manure also is a valuable source of nutrients and organic matter when used as a fertilizer. Manure was used as a fertilizer on about of US cropland in 2006, with manure from cattle accounting for nearly 70% of manure applications to soybeans and about 80% or more of manure applications to corn, wheat, barley, oats and sorghum. Substitution of manure for synthetic fertilizers in crop production can be environmentally significant, as between 43 and 88 megajoules of fossil fuel energy would be used per kg of nitrogen in manufacture of synthetic nitrogenous fertilizers. Grazing by cattle at low intensities can create a favourable environment for native herbs and forbs by mimicking the native grazers who they displaced; in many world regions, though, cattle are reducing biodiversity due to overgrazing. A survey of refuge managers on 123 National Wildlife Refuges in the US tallied 86 species of wildlife considered positively affected and 82 considered negatively affected by refuge cattle grazing or haying. Proper management of pastures, notably managed intensive rotational grazing and grazing at low intensities can lead to less use of fossil fuel energy, increased recapture of carbon dioxide, fewer ammonia emissions into the atmosphere, reduced soil erosion, better air quality, and less water pollution.


Health

The veterinary discipline dealing with cattle and cattle diseases (bovine veterinary) is called buiatrics. Veterinarians and professionals working on cattle health issues are pooled in the World Association for Buiatrics, founded in 1960. National associations and affiliates also exist. Cattle diseases were in the center of attention in the 1980s and 1990s when the Bovine spongiform encephalopathy (BSE), also known as mad cow disease, was of concern. Cattle might catch and develop various other diseases, like blackleg (disease), blackleg, bluetongue, foot rot too. In most states, as cattle health is not only a veterinarian issue, but also a public health issue, public health and food safety standards and farming regulations directly affect the daily work of farmers who keep cattle. However, said rules change frequently and are often debated. For instance, in the U.K., it was proposed in 2011 that milk from tuberculosis-infected cattle should be allowed to enter the food chain. Internal food safety regulations might affect a country's trade policy as well. For example, the United States has just reviewed its beef import rules according to the "mad cow standards"; while Mexico forbids the entry of cattle who are older than 30 months. Cow urine is commonly used in India for internal medical purposes. It is distilled and then consumed by patients seeking treatment for a wide variety of illnesses. At present, no conclusive medical evidence shows this has any effect. However, an Indian medicine containing cow urine has already obtained U.S. patents. Digital dermatitis is caused by the bacteria from the genus Treponema. It differs from foot rot and can appear under unsanitary conditions such as poor hygiene or inadequate hoof trimming, among other causes. It primarily affects dairy cattle and has been known to lower the quantity of milk produced, however the milk quality remains unaffected. Cattle are also susceptible to ringworm caused by the fungus, ''Trichophyton verrucosum'', a contagious skin disease which may be transferred to humans exposed to infected cows.


Effect of high stocking density

Stocking density refers to the number of animals within a specified area. When stocking density reaches high levels, the behavioural needs of the animals may not be met. This can negatively influence health, welfare and production performance. The effect of overstocking in cows can have a negative effect on Dairy, milk production and reproduction rates which are two very important traits for dairy farmers. Overcrowding of cows in barns has been found to reduced feeding, resting and rumination. Although they consume the same amount of dry matter within the span of a day, they consume the food at a much more rapid rate, and this behaviour in cows can lead to further complications. The feeding behaviour of cows during their post-milking period is very important as it has been proven that the longer animals can eat after milking, the longer they will be standing up and therefore causing less contamination to the teat ends. This is necessary to reduce the risk of mastitis as infection has been shown to increase the chances of embryonic loss. Sufficient rest is important for dairy cows because it is during this period that their resting blood flow increases up to 50%, this is directly proportionate to milk production. Each additional hour of rest can be seen to translate to 2 to 3.5 more pounds of milk per cow daily. Stocking densities of anything over 120% have been shown to decrease the amount of time cows spend lying down. Cortisol is an important stress hormone; its plasma concentrations increase greatly when subjected to high levels of stress.Sjaasted O.V., Howe K., Sand O., (2010) Physiology of Domestic Animals. 3rd edition. Sunderland: Sinaver Association. Inc Increased concentration levels of cortisol have been associated with significant increases in gonadotrophin levels and lowered progestin levels. Reduction of stress is important in the reproductive state of cows as an increase in Gonadotropin, gonadotrophin and lowered progesterone levels may impinge on the Ovulation, ovulatory and Luteal phase, lutenization process and to reduce the chances of successful implantation. A high cortisol level will also stimulate the degradation of fats and proteins which may make it difficult for the animal to sustain its pregnancy if implanted successfully.


Animal welfare concerns

Animal rights activists have criticized the treatment of cattle, claiming that common practices in cattle husbandry, animal slaughter, slaughter and entertainment unnecessarily cause fear, stress, and pain. They advocate for abstaining from the consumption of cattle-related animal products and cattle-based entertainment.


Livestock industry

The following husbandry practices have been criticized by animal welfare and animal rights groups: Livestock branding, branding, castration,
dehorning Dehorning is the process of removing the horns of livestock. Cattle, sheep, and goats are sometimes dehorned for economic and safety reasons. Disbudding is a different process with similar results; it cauterizes and thus destroys horn buds before ...
,
ear tag An ear tag is a plastic or metal object used for identification of domestic livestock and other animals. If the ear tag uses Radio Frequency Identification Device (RFID) technology it is referred to as an electronic ear tag. Electronic ear tags ...
ging, Nose ring (animal), nose ringing, Physical restraint, restraint, Docking (animal), tail docking, the use of
veal Veal is the meat of calves, in contrast to the beef from older cattle. Veal can be produced from a calf of either sex and any breed, however most veal comes from young male calves of dairy breeds which are not used for breeding. Generally, v ...
crates, and cattle prods. There are concerns that the stress and negative health impacts induced by high stocking density such as in concentrated animal feeding operations or feedlots, auctions, and during transport may be detrimental to their welfare, and has also been criticized. The Dairy cattle#Animal welfare, treatment of dairy cows faces additional criticism. To produce milk from dairy cattle, most calves are separated from their mothers soon after birth and fed milk replacement in order to retain the cows' milk for human consumption. Animal welfare advocates are critical of this practice, stating that this breaks the natural bond between the mother and her calf. The Veal#Animal welfare, welfare of veal calves is also a concern. In order to continue lactation, dairy cows are bred every year, usually through artificial insemination. Because of this, some individuals have posited that dairy production is based on the sexual exploitation of cows. Although the natural life expectancy of cattle could be as much as twenty years, after about five years, a cow's milk production has dropped; at which point most dairy cows are sent to slaughter.


Leather

While leather is often a by-product of slaughter, in some countries, such as India and Bangladesh, cows are raised primarily for their leather. These leather industries often make their cows walk long distances across borders to be killed in neighboring provinces and countries where cattle slaughter is legal. Some cows die along the long journey, and sometimes exhausted animals are abused to keep them moving. These practices have faced backlash from various animal rights groups.


Sport

Animal treatment in rodeo is targeted most often at bull riding but also calf roping and steer roping, with the opposition saying that rodeos are unnecessary and cause stress, injury, and death to the animals. In Spain, the Running of the bulls#Opposition, Running of the bulls faces opposition due to the stress and injuries incurred by the bulls during the event. Bullfighting#Animal welfare, Bullfighting is opposed as a blood sport in which bulls are forced to suffer severe stress and death.


Oxen

Oxen (singular ''ox'') are cattle trained as
draft animal A working animal is an animal, usually domesticated, that is kept by humans and trained to perform tasks instead of being slaughtered to harvest animal products. Some are used for their physical strength (e.g. oxen and draft horses) or for t ...
s. Often they are adult, castrated males of larger breeds, although females and bulls are also used in some areas. Usually, an ox is over four years old due to the need for training and to allow it to grow to full size. Oxen are used for plowing, transport, hauling cargo, grain-grinding by trampling or by powering machines, irrigation by powering pumps, and wagon drawing. Oxen were commonly used to skid logs in forests, and sometimes still are, in low-impact, select-cut logging. Oxen are most often used in teams of two, paired, for light work such as
cart A cart or dray (Australia and New Zealand) is a vehicle designed for transport, using two wheels and normally pulled by one or a pair of draught animals. A handcart is pulled or pushed by one or more people. It is different from the flatbed tr ...
ing, with additional pairs added when more power is required, sometimes up to a total of 20 or more. Oxen can be trained to respond to a teamster's signals. These signals are given by verbal commands or by noise (whip cracks). Verbal commands vary according to dialect and local tradition. Oxen can pull harder and longer than horses. Though not as fast as horses, they are less prone to injury because they are more sure-footed. Many oxen are used worldwide, especially in
developing countries A developing country is a sovereign state with a lesser developed industrial base and a lower Human Development Index (HDI) relative to other countries. However, this definition is not universally agreed upon. There is also no clear agreem ...
. About 11.3 million draft oxen are used in sub-Saharan Africa. In India, the number of draft cattle in 1998 was estimated at 65.7 million head. About half the world's crop production is thought to depend on land preparation (such as plowing) made possible by animal traction.


Religion, traditions and folklore


Islamic traditions

The cow is mentioned often in the Quran. The second and longest surah of the Quran is named Al-Baqara ("The Cow"). Out of the 286 verses of the surah, seven mention cows (Al Baqarah 67–73). The name of the surah derives from this passage in which Moses in Islam, Moses orders his people to sacrifice a cow in order to resurrect a man murdered by an unknown person.


Hindu traditions

Veneration of the cow has become a symbol of the identity of Hindus as a community, especially since the end of the 19th century. Slaughter of cows (including oxen, bulls and calves) is forbidden by law in several states of the Indian Union. McDonald's outlets in India do not serve any beef burgers. In Maharaja Ranjit Singh's empire of the early 19th century, the killing of a cow was punishable by death.


Other traditions

* The Evangelist St. Luke is depicted as an ox in Christian art. * In Judaism, as described in Numbers 19:2, the ashes of a sacrificed unblemished red heifer that has never been yoked can be used for ritual purification of people who came into contact with a corpse. * The ox is one of the 12-year cycle of animals which appear in the Chinese zodiac related to the Chinese calendar. See: Ox (Zodiac). * The constellation Taurus (constellation), Taurus represents a bull. * An apocryphal story has it that a cow started the Great Chicago Fire by kicking over a kerosene lamp. Michael Ahern, the reporter who created the cow story, admitted in 1893 that he had fabricated it for more colorful copy. * On 18 February 1930, Elm Farm Ollie became the first cow to fly in an airplane and also the first cow to be milked in an airplane. * The first known law requiring branding in North America was enacted on 5 February 1644, by Connecticut. It said that all cattle and pigs had to have a registered brand or earmark by 1 May 1644. * The is a traditional toy from the Aizu region of Japan that is thought to ward off illness. * The case of ''Sherwood v. Walker''—involving a supposedly barren heifer that was actually pregnant—first enunciated the concept of mutual mistake as a means of destroying the meeting of the minds in contract law. * The Fulani of West Africa are the world's largest nomadic cattle-herders. * The Maasai people, Maasai tribe of East Africa traditionally believe their god Engai entitled them to divine rights to the ownership of all cattle on earth.


In heraldry

Cattle are typically represented in heraldry by the bull. File:Coat of arms of the Azores.svg, Coat of arms of the Azores, Arms of the Azores File:Mecklenburg Arms.svg, Arms of Mecklenburg region, Germany File:Turin coat of arms.svg, Arms of Turin, Italy File:Coat of arms of Kaunas.svg, Arms of Kaunas, Lithuania File:Šiauliai COA great.svg, Arms of Šiauliai, Lithuania File:POL Bielsk Podlaski COA.svg, Arms of Bielsk Podlaski, Poland File:POL COA Ciołek.svg, Ciołek coat of arms, Arms of Ciołek, Poland File:POL Turek COA PioM.svg, Arms of Turek, Poland File:Arms_of_Béarn.svg, Arms of Béarn


Population

The cattle population of Britain rose from 9.8 million in 1878 to 11.7 million in 1908, but beef consumption rose much faster. Britain became the "stud farm of the world" exporting livestock to countries where there were no indigenous cattle. In 1929 80% of the meat trade of the world was products of what were originally English breeds. There were nearly 70 million cattle in the US by the early 1930s. For 2013, the FAO estimated global cattle numbers at 1.47 billion.FAOSTAT. [Agricultural statistics database] Food and Agriculture Organization of the United Nations, Rome. Regionally, the FAO estimate for 2013 includes: Asia 497 million; South America 350 million; Africa 307 million; Europe 122 million; North America 102 million; Central America 47 million; Oceania 40 million; and Caribbean 9 million. As per Foreign Agricultural Service, FAS/United States Department of Agriculture, USDA 2021 data, India had the largest cattle inventory in the world in 2021 followed by Brazil and China
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
's cattle's inventory was reported at 305.5 million head in 2021, accounting for roughly 30% of the world's inventory. India, Brazil and China accounted for roughly 65% of the world's cattle inventory in 2021. It has been estimated that out of all animal species on Earth, ''Bos taurus'' has the largest biomass at roughly 400 million tonnes, followed closely by ''Euphausia superba'' (Antarctic krill) at 379 million tonnes, and ''Homo sapiens'' (humans) at 373 million tonnes.


See also

* 1966 anti-cow slaughter agitation * :Individual cattle * Cattle Health Initiative, British Cattle Health Initiative * Bull-baiting * Bullocky * Bulls and Cows (game) * Cattle age determination * Cowboy * Intensive animal farming * List of cattle breeds * List of domesticated animals


References


Further reading

* Bhattacharya, S. 2003
Cattle ownership makes it a man's world
. ''Newscientist.com''. Retrieved 26 December 2006. * Cattle Today (CT). 2006. Website
Breeds of cattle
''Cattle Today''. Retrieved 26 December 2006 * Clay, J. 2004. ''World Agriculture and the Environment: A Commodity-by-Commodity Guide to Impacts and Practices''. Washington, DC: Island Press. . * Clutton-Brock, J. 1999. ''A Natural History of Domesticated Mammals''. Cambridge: Cambridge University Press. . * Huffman, B. 2006
''The ultimate ungulate page''
''UltimateUngulate.com''. Retrieved 26 December 2006. * Invasive Species Specialist Group (ISSG). 2005
''Bos taurus''
''Global Invasive Species Database''. * Johns, Catherine. 2011 ''Cattle: History, Myth, Art''. London: The British Museum Press. 978-0-7141-5084-0 * Nowak, R.M. and Paradiso, J.L. 1983. ''Walker's Mammals of the World''. Baltimore, MD: The Johns Hopkins University Press. * Oklahoma State University (OSU). 2006
''Breeds of Cattle''
Retrieved 5 January 2007. * Public Broadcasting Service (PBS). 2004

. ''PBS Nature''. Retrieved 5 January 2007. * – A visual textbook containing History/Origin, Phenotype & Statistics of 45 breeds. * Rath, S. 1998. ''The Complete Cow''. Stillwater, MN: Voyageur Press. . * Raudiansky, S. 1992. ''The Covenant of the Wild''. New York: William Morrow and Company, Inc. . * Spectrum Commodities (SC). 2006

''Spectrumcommodities.com''. Retrieved 5 January 2007. * Voelker, W. 1986. ''The Natural History of Living Mammals''. Medford, NJ: Plexus Publishing, Inc. . * Yogananda, P. 1946. ''The Autobiography of a Yogi''. Los Angeles: Self Realization Fellowship. . {{Authority control Cattle, Mammals described in 1758 Herbivorous mammals Articles containing video clips Taxa named by Carl Linnaeus Cosmopolitan mammals