Alethinophidia
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:''Common names: advanced snakes.'' The Alethinophidia are an
infraorder Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...
of
snakes Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more joi ...
that includes all snakes other than blind snakes and thread snakes. Snakes have long been grouped into families within Alethinophidia based on their morphology, especially that of their teeth. More modern phylogenetic hypotheses using genetic data support the recognition of 19 extant families (see below), although the taxonomy of alethinophidian snakes has long been debated, and ultimately the decision whether to assign a particular clade to a particular Linnaean rank (such as a
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
,
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
, or
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
) is arbitrary.


Etymology

The infraorder name Alethinophidia derives from the two
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
words (), meaning "truthful, genuine", and (), meaning "snake".


Fossil record

Fossils of alethinophidians were found in
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
(Middle
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
) sites of Wadi Milk Formation in Wadi Abu Hashim,
Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...
. ''
Coniophis ''Coniophis'' is an extinct genus of snakes from the late Cretaceous period. The type species, ''Coniophis precedes'', was about 7 cm long and had snake-like teeth and body form, with a skull and a largely lizard-like bone structure. It pr ...
'' presents the vertebral morphology similar to modern-day
Aniliidae The Aniliidae are a monotypic family created for the monotypic genus ''Anilius'' that contains the single species ''A. scytale''. Common names include American pipe snake and false coral snake. It is found in South America. This snake posses ...
. Two extinct families from the same location, the Anomalophiidae and Russellophiidae, also belong to the Alethinophidia. '' Krebsophis'' is the earliest russellophiid. The family Nigerophiidae includes both aquatic '' Nubianophis'' from Wadi Abu Hashim and '' Nigerophis'' from the
Palaeocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''palai ...
of
Niger ) , official_languages = , languages_type = National languagesEoanilius'' (belongs to Aniliidae) appeared in the
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
. It is also existed in
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
and early
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
.


Systematics

* Superfamily
Amerophidia The Amerophidia, also known as amerophidian snakes, are a superfamily of snakes that contains two families: Aniliidae (containing a single species, ''Anilius scytale The Aniliidae are a monotypic family created for the monotypic genus ''Ani ...
** Family:
Aniliidae The Aniliidae are a monotypic family created for the monotypic genus ''Anilius'' that contains the single species ''A. scytale''. Common names include American pipe snake and false coral snake. It is found in South America. This snake posses ...
Stejneger, 1907—red pipesnake ** Family:
Tropidophiidae The Tropidophiidae, common name dwarf boas or thunder snakes, are a family of nonvenomous snakes found from Mexico and the West Indies south to southeastern Brazil. These are small to medium-sized fossorial snakes, some with beautiful and strik ...
Brongersma, 1951—Caribbean dwarf "boas" or thunder snakes * Superfamily
Booidea The Booidea, also known as booid snakes, are a superfamily of snakes that contains boas (family Boidae) and other closely related boa-like snakes (but not pythons, which are in a separate superfamily called Pythonoidea). As of 2017, Booidea conta ...
** Family:
Boidae The Boidae, commonly known as boas or boids, are a family of nonvenomous snakes primarily found in the Americas, as well as Africa, Europe, Asia, and some Pacific Islands. Boas include some of the world's largest snakes, with the green anaconda ...
Gray Grey (more common in British English) or gray (more common in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning literally that it is "without color", because it can be composed o ...
, 1825—boas (see article for comments on former families or subfamilies Calabariidae/inae, Sanziniidae/inae, Charinidae/inae, Erycidae/inae, Candoiidae/inae) * Superfamily
Pythonoidea The Pythonoidea, also known as pythonoid snakes, are a superfamily of snakes that contains pythons (family Pythonidae) and other closely related python-like snakes (but not boas, which are in a separate superfamily called Booidea). As of 2022, ...
** Family:
Pythonidae The Pythonidae, commonly known as pythons, are a family of nonvenomous snakes found in Africa, Asia, and Australia. Among its members are some of the largest snakes in the world. Ten genera and 42 species are currently recognized. Distributi ...
Fitzinger, 1826—pythons ** Family:
Loxocemidae :''Common names: Mexican python, Mexican burrowing python, Mexican burrowing snake.'' ''Loxocemus bicolor'',the sole member of the monotypic family Loxocemidae, is a species of python-like snake found in Mexico and Central America. No subspecies ...
Cope The cope (known in Latin as ''pluviale'' 'rain coat' or ''cappa'' 'cape') is a liturgical vestment, more precisely a long mantle or cloak, open in front and fastened at the breast with a band or clasp. It may be of any liturgical colours, litu ...
, 1861—Mexican burrowing pythons ** Family:
Xenopeltidae ''Xenopeltis'', the sunbeam snakes, are the sole genus of the monotypic family Xenopeltidae, the species of which are found in Southeast Asia. Sunbeam snakes are known for their highly iridescent scales. Two species are recognized, each one with ...
Bonaparte, 1845—sunbeam snakes * Superfamily Uropeltoidea ** Family:
Uropeltidae The Uropeltidae, also known Common name, commonly as the shieldtails or the shield-tailed snakes, are a Family (biology), family of primitive, nonvenomous, burrowing snakes native to peninsular India and Sri Lanka. The name is derived from the Gr ...
Müller, 1832—shield-tailed snakes ** Family:
Cylindrophiidae The Cylindrophiidae are a monotypic family of secretive, semifossorial, non-venomous snakes containing the genus ''Cylindrophis'' found in southeastern Asia. These are burrowing snakes and most have a banded pattern on the belly. Currently, thirt ...
Fitzinger, 1843—Asian pipe snakes ** Family: Anomochilidae Cundall, Wallach and Rossman, 1993—dwarf pipe snakes * Family:
Bolyeriidae :''Common names: Mauritius snakes, Round Island boas, splitjaw snakes.'' The Bolyeriidae are a family of snakes native to Mauritius and a few islands around it, especially Round Island. They also used to be found on the island of Mauritius, but ...
Hoffstetter, 1946—Splitjaw snakes * Family:
Xenophidiidae ''Xenophidion'' is a genus of snakes first described in 1995, and the only genus of the monotypic family Xenophidiidae.. Commonly referred to as spinejaw snakes, this genus is found in Borneo and peninsular Malaysia. Morphology and classificatio ...
Wallach & Günther, 1998—Spine-jawed snakes * Family:
Acrochordidae The Acrochordidae, commonly known as wart snakes, Java wart snakes, file snakes, elephant trunk snakes, or dogface snakes are a monogeneric family created for the genus ''Acrochordus''. This is a group of basal aquatic snakes found in Australia a ...
Bonaparte, 1831—wart or file snakes * Family:
Xenodermidae Xenodermidae is a family of snakes native to East Asia, South Asia, and Southeast Asia. All species in the family Xenodermidae are small or moderately sized snakes, never more than but typically less than in total length (including tail). They ...
Oppel, 1811—odd-scaled snakes * Family:
Pareidae Pareidae is a small family of snakes found largely in southeast Asia, with an isolated subfamily endemic to southwestern India. It encompasses 42 species in four genera divided into two subfamilies: Pareinae and Xylophiinae. Both families are tho ...
Oppel, 1811—snail-eating snakes * Family:
Viperidae The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs tha ...
Oppel, 1811—vipers (including pit vipers) ** Subfamily: Azemiopinae Liem, Marx and Rabb, 1971—Fea's viper ** Subfamily:
Crotalinae The Crotalinae, commonly known as pit vipers,Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . crotaline snakes (from grc, κρόταλον ''krotalon'' castanet), or pit adders, are a subfa ...
Oppel, 1811—pitvipers (including rattlesnakes) ** Subfamily:
Viperinae The Viperinae, or viperines, are a subfamily of vipers endemic to Europe, Asia and Africa. They are distinguished by their lack of the heat-sensing pit organs that characterize their sister group, the subfamily Crotalinae. Currently, 13 gen ...
Oppel, 1811—true vipers * Family:
Homalopsidae The Homalopsidae are a family of snakes which contains about 28 genus, genera and more than 50 species. They are commonly known as Indo-Australian water snakes, mudsnakes, or bockadams. They are also known as ''ular air'' (lit. "water snake") in ...
Günther, 1864—Asian mudsnakes * Superfamily:
Elapoidea The Elapoidea are a Superfamily (biology), superfamily of snakes in the clade Colubroides, traditionally comprising the families Lamprophiidae and Elapidae. Advanced genomic sequence studies, however, have found lamprophiids to be paraphyletic in ...
F. Boie, 1827 (merged with Colubroidea by the
Reptile Database The Reptile Database is a scientific database that collects taxonomic information on all living reptile species (i.e. no fossil species such as dinosaurs). The database focuses on species (as opposed to higher ranks such as families) and has entrie ...
) ** Family: Cyclocoridae Weinell &
Brown Brown is a color. It can be considered a composite color, but it is mainly a darker shade of orange. In the CMYK color model used in printing or painting, brown is usually made by combining the colors orange and black. In the RGB color model used ...
, 2017—Philippine snakes ** Family:
Elapidae Elapidae (, commonly known as elapids ; grc, ἔλλοψ ''éllops'' "sea-fish") is a family of snakes characterized by their permanently erect fangs at the front of the mouth. Most elapids are venomous, with the exception of the genus Emydoceph ...
F. Boie, 1827—Cobras, coral snakes, mambas, taipans,
sea snakes Sea snakes, or coral reef snakes, are elapid snakes that inhabit marine environments for most or all of their lives. They belong to two subfamilies, Hydrophiinae and Laticaudinae. Hydrophiinae also includes Australasian terrestrial snakes, wher ...
, and others ** Family: Pseudaspididae
Cope The cope (known in Latin as ''pluviale'' 'rain coat' or ''cappa'' 'cape') is a liturgical vestment, more precisely a long mantle or cloak, open in front and fastened at the breast with a band or clasp. It may be of any liturgical colours, litu ...
, 1893—mole snake, western keeled snake, and mock vipers ** Family: Prosymnidae Gray, 1849—shovel-snouted snakes ** Family:
Psammophiidae Psammophiidae is a family of elapoid snakes. They were formerly placed as a subfamily of the Lamprophiidae, but have been more recently identified as a distinct family. Genera It contains 8 genera: * '' Dipsina'' Jan, 1862 * ''Hemirhagerrhis'' ...
Dowling, 1967—sand snakes and allies ** Family: Atractaspididae Günther, 1858—African burrowing asps, stiletto snakes, harlequin snakes ** Family:
Pseudoxyrhophiidae The Pseudoxyrhophiidae is a family of elapoid snakes, found mostly in Madagascar. They were formerly placed as a subfamily of the Lamprophiidae, but have been more recently identified as a distinct family. It contains about 22 genera in two s ...
Dowling, 1975—Malagasy hognose snakes, brook snakes, and allies ** Family:
Lamprophiidae The Lamprophiidae are a family of snakes found throughout much of Africa, including the Seychelles. There are 89 species as of July 2022. Biology Lamprophiids are a very diverse group of snakes. Many are terrestrial but some are fossorial (e.g. ...
Fitzinger, 1843—lamprophiids * Superfamily:
Colubroidea Colubroidea is a superfamily of snakes in the clade Colubroides that includes Colubridae, with some studies splitting Colubridae into multiple families that make up Colubroidea. Historically, Colubroidea also included other caenophidian snakes su ...
Oppel, 1811 ** Family:
Colubridae Colubridae (, commonly known as colubrids , from la, coluber, 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest species of the family date back to the Oligocene epoch. Colubrid snakes are found on ever ...
Oppel, 1811—colubrids, typical snakes (subfamilies sometimes considered distinct families) *** Subfamily:
Sibynophiinae Sibynophiinae is a small subfamily of colubroid snakes, sometimes referred to as a family (Sibynophiidae). This group has also been called Scaphiodontophiinae but since the name Sibynophiinae is older, it has priority. They are commonly called h ...
Dunn, 1928—hinged-teeth snakes *** Subfamily:
Natricinae The Natricinae are a subfamily of colubroid snakes, sometimes referred to as a family (Natricidae). The subfamily comprises 37 genera. Members include many very common snake species, such as the European grass snakes, and the North American wa ...
Bonaparte, 1838—keelbacks *** Subfamily:
Pseudoxenodontinae Pseudoxenodontinae is a small subfamily of colubroid snakes, sometimes referred to as a family (Pseudoxenodontidae). They are found in southern and southeastern Asia, from northeast India to southern China (including Taiwan) and south into Indon ...
McDowell, 1987 *** Subfamily:
Dipsadinae Dipsadinae is a large subfamily of colubroid snakes, sometimes referred to as a family (Dipsadidae). They are found in most of the Americas, including the West Indies, and are most diverse in South America. There are more than 700 species. Dips ...
Bonaparte, 1838


See also

*
Scolecophidia The Scolecophidia, commonly known as blind snakes or thread snakes, are an infraorder of snakes. They range in length from . All are fossorial (adapted for burrowing). Five families and 39 genera are recognized. The Scolecophidia infraorder is mos ...
, blind snakes, thread snakes. *
List of snakes List of snakes refers to a variety of different articles and different criteria. these are listed below. Lists General lists *General lists: ** Snake#Taxonomy ** List of reptile genera#Order Squamata ** List of snakes by common name **List of sn ...
, overview of all snake genera.


References


External links


Alethinophidia
a
Palaeos
Accessed 14 August 2007. {{Taxonbar, from=Q940396 Cenomanian first appearances Extant Cretaceous first appearances Taxa named by Franz Nopcsa von Felső-Szilvás