Zhelestid
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Zhelestid
Zhelestidae is a lineage of extinct eutherian mammals. Occurring in the Late Cretaceous from the Turonian to the Maastrichtian, they were an extremely successful group, with representatives present in Europe, Asia, India (and subsequently in Madagascar), Africa and North America, ostensibly rendering them a cosmopolitan clade. They were specialised towards an herbivorous lifestyle and were in fact initially considered stem-ungulates, but the presence of epipubics and "archaic" dental characters render them as non-placental eutherians.

Range

The earliest zhelestid remains occur in the of . By the

UA 8699
UA 8699 (University of Antananarivo specimen 8699)Krause, 2001, p. 497 is a fossil mammalian tooth from the Cretaceous of Madagascar. A broken lower molar about 3.5 mm (0.14 in) long, it is from the Maastrichtian of the Maevarano Formation in northwestern Madagascar. Details of its crown morphology indicate that it is a boreosphenidan, a member of the group that includes living marsupials and placental mammals. David W. Krause, who first described the tooth in 2001, interpreted it as a marsupial on the basis of five shared characters, but in 2003 Averianov and others noted that all those are shared by zhelestid placentals and favored a close relationship between UA 8699 and the Spanish zhelestid '' Lainodon''. Krause used the tooth as evidence that marsupials were present on the southern continents (Gondwana) as early as the late Cretaceous and Averianov and colleagues proposed that the tooth represented another example of faunal exchange between Africa and Europe at ...
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Azilestes
''Azilestes'' ("Mas-d'Azil robber") is a genus of probable Zhelestidae, zhelestid eutherian mammal, a family consisting of small herbivores, that was discovered in the early Maastrichtian Grès de Labarre, Grès de Labarre Formation of France.Gheerbrant E. & Teodori D. 2021. — An enigmatic specialized new eutherian mammal from the Late Cretaceous of Western Europe (Northern Pyrenees), in Folie A., Buffetaut E., Bardet N., Houssaye A., Gheerbrant E. & Laurin M. (eds), Palaeobiology and palaeobiogeography of amphibians and reptiles: An homage to Jean-Claude Rage. Comptes Rendus Palevol 20 (13): 207-223. https://doi.org/10.5852/cr-palevol2021v20a13 It is a monotypic genus, with only type species ''A. ragei'' being known. Only one specimen, the holotype described in 2021, is known. It consists of a partial dentary with teeth. Certain aspects of ''Azilestes'''s dental anatomy are convergent with later groups of herbivorous mammals, including Glires, though phylogenetic analysis stro ...
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Oxlestes
''Oxlestes'' is an extinct mammal from the Late Cretaceous of Asia, more specifically from the Cenomanian of Uzbekistan. A carnivorous species of uncertain affinities, it is notable for its relatively large size, being among the largest of all Mesozoic mammals.L. A. Nessov. 1982. Drevneishie mlekopitaiushchie SSSR ncient mammals of the USSR Palyentologicheskogo Obshchyestva 25:228-243 Due to the limited amount of material, it has been considered a ''nomen dubium''. Description ''Oxlestes'' is currently a monotypic genus, containing a single species, ''O. grandis''. Its specimens were recovered in the Cenomanian-aged Khodzhakul Formation of Uzbekistan. The holotype, CCMGE 6/11758, is composed of an axis vertebra, dentaries, canine teeth and a sagittal crest. The axis is relatively narrow, with a long, pointed anterior process. There are two pairs of distinct foramina anterior and posterior to the dorsal transverse suture; modern placentals only bear one. It is about 1.9 centime ...
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Valentinella
''Valentinella'' is a Late Cretaceous genus of eutherian mammal from France, Europe.Tabuce, R., Vianey-Liaud, M. and Garcia, G., 2004A eutherian mammal in the latest Cretaceous of Vitrolles, southern France ''Acta Palaeontologica Polonica'', 49(3). It was originally known from some damaged lower and upper jaws, and was cautiously referred to the lainodontine zhelestid mammals. This identification was later questioned Gheerbrant, E. and Astibia, H., 2012Addition to the Late Cretaceous Laño mammal faunule (Spain) and to the knowledge of European “Zhelestidae”(Lainodontinae nov.) Bulletin de la Société géologique de France, 183(6), pp.537-546. and even considered a ''nomen dubium'' by some researchers,Archibald, J.D. and Averianov, A., 2012Phylogenetic analysis, taxonomic revision, and dental ontogeny of the Cretaceous Zhelestidae (Mammalia: Eutheria) ''Zoological Journal of the Linnean Society'', 164(2), pp.361-426. and remains uncertain due to the scarcity and fragmentary n ...
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India
India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the south, the Arabian Sea on the southwest, and the Bay of Bengal on the southeast, it shares land borders with Pakistan to the west; China, Nepal, and Bhutan to the north; and Bangladesh and Myanmar to the east. In the Indian Ocean, India is in the vicinity of Sri Lanka and the Maldives; its Andaman and Nicobar Islands share a maritime border with Thailand, Myanmar, and Indonesia. Modern humans arrived on the Indian subcontinent from Africa no later than 55,000 years ago., "Y-Chromosome and Mt-DNA data support the colonization of South Asia by modern humans originating in Africa. ... Coalescence dates for most non-European populations average to between 73–55 ka.", "Modern human beings—''Homo sapiens''—originated in Africa. Then, int ...
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Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area of , about 30% of Earth's total land area and 8.7% of Earth's total surface area. The continent, which has long been home to the majority of the human population, was the site of many of the first civilizations. Its 4.7 billion people constitute roughly 60% of the world's population. In general terms, Asia is bounded on the east by the Pacific Ocean, on the south by the Indian Ocean, and on the north by the Arctic Ocean. The border of Asia with Europe is a historical and cultural construct, as there is no clear physical and geographical separation between them. It is somewhat arbitrary and has moved since its first conception in classical antiquity. The division of Eurasia into two continents reflects East–West cultural, linguistic, ...
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Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia and it is located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. Comprising the westernmost peninsulas of Eurasia, it shares the continental landmass of Afro-Eurasia with both Africa and Asia. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south and Asia to the east. Europe is commonly considered to be Boundaries between the continents of Earth#Asia and Europe, separated from Asia by the drainage divide, watershed of the Ural Mountains, the Ural (river), Ural River, the Caspian Sea, the Greater Caucasus, the Black Sea and the waterways of the Turkish Straits. "Europe" (pp. 68–69); "Asia" (pp. 90–91): "A commonly accepted division between Asia and E ...
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Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the Danian (part of the Paleogene and Paleocene). The Cretaceous–Paleogene extinction event (formerly known as the Cretaceous–Tertiary extinction event) occurred at the end of this age. In this mass extinction, many commonly recognized groups such as non-avian dinosaurs, plesiosaurs and mosasaurs, as well as many other lesser-known groups, died out. The cause of the extinction is most commonly linked to an asteroid about wide colliding with Earth, ending the Cretaceous. Stratigraphic definitions Definition The Maastrichtian was introduced into scientific literature by Belgian geologist André Hubert Dumont in 1849, after studying rock strata of the Chalk Group c ...
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Turonian
The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded by the Cenomanian Stage and underlies the Coniacian Stage. At the beginning of the Turonian an oceanic anoxic event (OAE 2) took place, also referred to as the Cenomanian-Turonian boundary event or the "Bonarelli Event". Stratigraphic definition The Turonian (French: ''Turonien'') was defined by the French paleontologist Alcide d'Orbigny (1802–1857) in 1842. Orbigny named it after the French city of Tours in the region of Touraine (department Indre-et-Loire), which is the original type locality. The base of the Turonian Stage is defined as the place where the ammonite species '' Watinoceras devonense'' first appears in the stratigraphic column. The official reference profile (the GSSP) for the base of the Turonian is located in the Roc ...
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Mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla (hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla ( cetaceans and even-toed ungulates), and the Carnivora (cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, together with Saur ...
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Eutherian
Eutheria (; from Greek , 'good, right' and , 'beast'; ) is the clade consisting of all therian mammals that are more closely related to placentals than to marsupials. Eutherians are distinguished from noneutherians by various phenotypic traits of the feet, ankles, jaws and teeth. All extant eutherians lack epipubic bones, which are present in all other living mammals (marsupials and monotremes). This allows for expansion of the abdomen during pregnancy. The oldest-known eutherian species is '' Juramaia sinensis'', dated at from the early Late Jurassic ( Oxfordian) of China. Eutheria was named in 1872 by Theodore Gill; in 1880 Thomas Henry Huxley defined it to encompass a more broadly defined group than Placentalia. Characteristics Distinguishing features are: *an enlarged malleolus ("little hammer") at the bottom of the tibia, the larger of the two shin bones *the joint between the first metatarsal bone and the entocuneiform bone (the innermost of the three cuneiform ...
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