Eutheria (; from Greek , 'good, right' and , 'beast'; ) is the

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

consisting of all therian mammals that are more closely related to

placentals Placental mammals (infraclass Placentalia ) are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupialia. Placentalia contains the vast majority of extant mammals, which are partly distinguishe ...

than to marsupials. Eutherians are distinguished from noneutherians by various

phenotypic trait A phenotypic trait, simply trait, or character state is a distinct variant of a phenotypic characteristic of an organism; it may be either inherited or determined environmentally, but typically occurs as a combination of the two.Lawrence, Eleano ...

s of the feet, ankles, jaws and teeth. All extant eutherians lack

epipubic bone Epipubic bones are a pair of bones projecting forward from the pelvic bones of modern marsupials, monotremes and fossil mammals like multituberculates, and even basal eutherians (the ancestors of placental mammals, who lack them). They first ...

s, which are present in all other living mammals (marsupials and

monotreme Monotremes () are prototherian mammals of the order Monotremata. They are one of the three groups of living mammals, along with placentals (Eutheria), and marsupials (Metatheria). Monotremes are typified by structural differences in their brain ...

s). This allows for expansion of the abdomen during pregnancy. The oldest-known eutherian species is '' Juramaia sinensis'', dated at from the early

Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...

( Oxfordian) of China. Eutheria was named in 1872 by

Theodore Gill Theodore Nicholas Gill (March 21, 1837 – September 25, 1914) was an American ichthyologist, mammalogist, malacologist and librarian. Career Born and educated in New York City under private tutors, Gill early showed interest in natural histor ...

; in 1880

Thomas Henry Huxley Thomas Henry Huxley (4 May 1825 – 29 June 1895) was an English biologist and anthropologist specialising in comparative anatomy. He has become known as "Darwin's Bulldog" for his advocacy of Charles Darwin's theory of evolution. The storie ...

defined it to encompass a more broadly defined group than Placentalia.

Characteristics

Cambridge Natural History Mammalia Fig 068

Distinguishing features are: *an enlarged

malleolus A malleolus is the bony prominence on each side of the human ankle. Each leg is supported by two bones, the tibia on the inner side (medial) of the leg and the fibula on the outer side (lateral) of the leg. The medial malleolus is the promine ...

("little hammer") at the bottom of the

tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...

, the larger of the two shin bones *the

joint A joint or articulation (or articular surface) is the connection made between bones, ossicles, or other hard structures in the body which link an animal's skeletal system into a functional whole.Saladin, Ken. Anatomy & Physiology. 7th ed. McGraw ...

between the first

metatarsal bone The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the ...

and the entocuneiform bone (the innermost of the three

cuneiform bones There are three cuneiform ("wedge-shaped") bones in the human foot: * the first or medial cuneiform * the second or intermediate cuneiform, also known as the middle cuneiform * the third or lateral cuneiform They are located between the navicu ...

) in the foot is offset farther back than the joint between the second metatarsal and middle cuneiform bones—in metatherians these joints are level with each other *various features of jaws and teeth

Evolutionary history

Eutheria contains several

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...

as well as larger groups, many with complicated

taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...

histories still not fully understood. Members of the

Adapisoriculidae Adapisoriculidae is an extinct family of non-placental eutherian mammals present during the Paleogene and possibly the Late Cretaceous. They were once thought to be members of the order Erinaceomorpha, closely related to the hedgehog family ( E ...

Cimolesta Cimolesta is an extinct order of non-placental eutherian mammals. Cimolestans had a wide variety of body shapes, dentition and lifestyles, though the majority of them were small to medium-sized general mammals that bore superficial resemblances ...

and

Leptictida Leptictida (''leptos iktis'' "small/slender weasel") is a possibly paraphyletic extinct order of eutherian mammals. Their classification is contentious: according to cladistic studies, they may be (distantly) related to Euarchontoglires (rodents ...

have been previously placed within the out-dated placental group

Insectivora The order Insectivora (from Latin ''insectum'' "insect" and ''vorare'' "to eat") is a now-abandoned biological grouping within the class of mammals. Some species have now been moved out, leaving the remaining ones in the order Eulipotyphla, wi ...

, while Zhelestids have been considered primitive

ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...

s. However, more recent studies have suggested these enigmatic taxa represent

stem group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. ...

eutherians, more basal to Placentalia. The weakly favoured cladogram favours Boreoeuthearia as a basal eutherian clade as sister to the Atlantogenata.

References

{{Taxonbar, from=Q17092469 Mammal taxonomy Oxfordian first appearances Extant Late Jurassic first appearances Taxa described in 1872