Tyrannosauroidea
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Tyrannosauroidea
Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent beginning in the Jurassic Period. By the end of the Cretaceous Period, tyrannosauroids were the dominant large predators in the Northern Hemisphere, culminating in the gigantic ''Tyrannosaurus''. Fossils of tyrannosauroids have been recovered on what are now the continents of North America, Europe and Asia, with fragmentary remains possibly attributable to tyrannosaurs also known from South America and Australia. Tyrannosauroids were bipedal carnivores, as were most theropods, and were characterized by numerous synapomorphy, skeletal features, especially of the skull and pelvis. Early in their existence, tyrannosauroids were small predators with long, three-fingered forelimbs. Late Cretaceous genera became much larger, including some of the ...
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Santanaraptor
''Santanaraptor'' (meaning "Santana Formation thief") is a genus of Tyrannosauroidea, tyrannosauroid Theropoda, theropod dinosaur that lived in South America during the Early Cretaceous (late Aptian-early Albian), about 112 million years ago. Discovery The type species is ''S. placidus'', first described by Alexander Kellner, Kellner in 1999. The species epithet refers to , who founded the . Description The holotype (National Museum of Brazil, MN 4802-V) is a juvenile partial skeleton consisting of three caudal vertebrae with chevrons, ischia, femora, tibia, fibula, pes, and soft tissue. The fossilized tissue includes a thin Epidermis (zoology), epidermis, muscle fibers, and possibly blood vessels. Skin impressions under the left foot are also preserved, showing Scale (anatomy), scales. It was unearthed in 1996 from the Romualdo Formation (Santana Group) in the Ceará State, Northeast Region, Brazil, northeastern Brazil. While primarily known from hindquarter elements, the indiv ...
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Tyrannosaurus Rex
''Tyrannosaurus'' is a genus of large theropod dinosaur. The species ''Tyrannosaurus rex'' (''rex'' meaning "king" in Latin), often called ''T. rex'' or colloquially ''T-Rex'', is one of the best represented theropods. ''Tyrannosaurus'' lived throughout what is now western North America, on what was then an island continent known as Laramidia. ''Tyrannosaurus'' had a much wider range than other tyrannosaurids. Fossils are found in a variety of rock formations dating to the Maastrichtian age of the Upper Cretaceous period, 68 to 66 million years ago. It was the last known member of the tyrannosaurids and among the last non-avian dinosaurs to exist before the Cretaceous–Paleogene extinction event. Like other tyrannosaurids, ''Tyrannosaurus'' was a bipedal carnivore with a massive skull balanced by a long, heavy tail. Relative to its large and powerful hind limbs, the forelimbs of ''Tyrannosaurus'' were short but unusually powerful for their size, and they had two clawed ...
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Coeluridae
Coeluridae is a historically unnatural group of generally small, carnivorous dinosaurs from the late Jurassic Period. For many years, any small Jurassic or Cretaceous theropod that did not belong to one of the more specialized families recognized at the time was classified with the coelurids, creating a confusing array of 'coelurid' theropods that were not closely related. Although they have been traditionally included in this family, there is no evidence that any of these primitive coelurosaurs form a natural group with ''Coelurus'', the namesake of Coeluridae, to the exclusion of other traditional coelurosaur groups. Classification Before the use of phylogenetic analyses, Coeluridae and Coelurosauria were taxonomic wastebaskets used for small theropods that did not belong to other groups; thus, they accumulated many dubious genera. As late as the 1980s, popular books recognized over a dozen "coelurids", including such disparate forms as the noasaurid ''Laevisuchus'' and the o ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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Labocania
''Labocania'' is a genus of carnivorous theropod dinosaur of uncertain affinities from Baja California, Mexico, which lived 73 million years ago, in the Campanian stage of the late Cretaceous Period. In the summer of 1970, the National Geographic Society and the Los Angeles County Museum of Natural History organized a joint paleontological expedition, led by geologist William J. Morris, to the Arroyo del Rosario in Baja California. While prospecting, volunteer Harley James Garbani discovered the skeleton of a theropod north of Punta Baja near Cerro Rayado. Garbani excavated the site in 1970 and 1971. The type species, ''Labocania anomala'', was described and named by Ralph Molnar Ralph E. Molnar is a paleontologist who had been Curator of Mammals at the Queensland Museum and more recently associated with the Museum of Northern Arizona. He is also a research associate at the Texas natural Science Centre. He co-authored descr ... in 1974. The generic name references the La ...
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Eotyrannus
''Eotyrannus'' (meaning "dawn tyrant") is a genus of tyrannosauroid theropod dinosaur hailing from the Early Cretaceous Wessex Formation beds, included in Wealden Group, located in the southwest coast of the Isle of Wight, United Kingdom. The remains (MIWG1997.550), consisting of assorted skull, axial skeleton and appendicular skeleton elements, from a juvenile or subadult, found in a plant debris clay bed, were described by Hutt ''et al.'' in early 2001. The etymology of the generic name refers to the animal's classification as an early tyrannosaur or "tyrant lizard", while the specific name honors the discoverer of the fossil. Discovery and naming The exact location of the discovery of the holotype specimen has not been revealed due to its importance and the possibility of new material to be collected as the coastline recedes. From what is mentioned in the description, the specimen was found on the southwestern coast of the Isle of Wight, between Atherfield Point and Hanove ...
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Juratyrant
''Juratyrant'' (meaning "Jurassic tyrant") is a tyrannosauroid dinosaur genus from the late Jurassic period (early Tithonian age) of England. The genus contains a single species, ''Juratyrant langhami'', which was once classed as a species of ''Stokesosaurus''. Discovery The species is known from a single specimen consisting of an "associated partial skeleton represented by a complete pelvis" as well as a partially complete leg and neck, back and tail vertebrae. This skeleton was discovered in 1984 in Dorset. The specimen was mentioned in several papers, but was not formally described until 2008. The species was named in honor of commercial fossil collector Peter Langham, who uncovered the specimen. The specimen was discovered in strata of the Kimmeridge Clay dating from the Tithonian, the final stage of the Late Jurassic, and belonging to the ''Pectinatites pectinatus'' ammonite zone, indicating the fossil is between 149.3 and 149 million years old. Description Paul (2010) l ...
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Stokesosaurus
''Stokesosaurus'' (meaning "Stokes' lizard") is a genus of small (around in length), carnivorous early tyrannosauroid theropod dinosaurs from the late Jurassic period of Utah, United States. History From 1960 onwards Utah geologist William Lee Stokes and his assistant James Henry Madsen excavated thousands of disarticulated ''Allosaurus'' bones at the Cleveland-Lloyd Dinosaur Quarry in Emery County, Utah. During the early 1970s, Madsen began to catalogue these finds in detail, discovering that some remains represented species new to science. In 1974 Madsen named and described the type species ''Stokesosaurus clevelandi''. Its generic name honours Stokes. The specific name refers to the town of Cleveland, Utah. The holotype ( UMNH 2938, also known as UMNH VP 7473 and formerly known as UUVP 2938) was uncovered in the Brushy Basin Member of the Morrison Formation dating from the early Tithonian stage, about 150 million years old. It consists of a left ilium or hip bone, bel ...
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Eutyrannosauria
Eutyrannosauria is a clade of tyrannosauroid theropods whose distribution has been found in what is now Asia and North America. The clade consists of an evolutionary grade of tyrannosaurs such as ''Appalachiosaurus'', ''Dryptosaurus'', and ''Bistahieversor'' which led up to the family Tyrannosauridae Tyrannosauridae (or tyrannosaurids, meaning "tyrant lizards") is a family of coelurosaurian theropod dinosaurs that comprises two subfamilies containing up to thirteen genera, including the eponymous ''Tyrannosaurus''. The exact number of genera .... The group was named in 2018 by Delcourt and Grillo in their paper about possible southern hemisphere tyrannosauroids and the phylogeography of tyrannosaurs. Classification Below is a phylogeographic cladogram of Eutyrannosauria after Voris ''et al.'' (2020): References Tyrannosaurs {{Theropod-stub ...
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Megaraptora
Megaraptora is a clade of carnivorous Tetanurae, tetanuran theropod dinosaurs with controversial relations to other theropods. Its Derived (phylogenetics), derived members, the Megaraptoridae are noted for their elongated hand claws and proportionally large arms, which are usually reduced in size in other large theropods. Megaraptorans are incompletely known, and no complete megaraptoran skeleton has been found. However, they still possessed a number of unique features. Their forelimbs were large and strongly built, and the Glossary of dinosaur anatomy#ulna, ulna bone had a unique shape in members of the Family (biology), family Megaraptoridae, a subset of megaraptorans which excludes ''Fukuiraptor'' and ''Phuwiangvenator''. The first two fingers were elongated, with massive curved claws, while the third finger was small. Megaraptoran skull material is very incomplete, but a juvenile ''Megaraptor'' described in 2014 preserved a portion of the snout, which was long and slender. Le ...
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Teihivenator
This list of informally named dinosaurs is a listing of dinosaurs (excluding Aves; birds and their extinct relatives) that have never been given formally published scientific names. This list only includes names that were not properly published ("unavailable names") and have not since been published under a valid name (see list of dinosaur genera for valid names). The following types of names are present on this list: * ''Nomen nudum'', Latin for "naked name": A name that has appeared in print but has not yet been formally published by the standards of the International Commission on Zoological Nomenclature. ''Nomina nuda'' (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. * '' Nomen manuscriptum'', Latin for "manuscript name": A name that appears in manuscript but was not formally published. A ''nomen manuscriptum'' is equivalent to a ''nomen nudum'' for everything except the method of publication, and description. * '' Nomen ex d ...
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Timurlengia
''Timurlengia'' is an extinct genus of tyrannosauroid theropod dinosaur found in Uzbekistan, in the Bissekty Formation in the Kyzylkum Desert, hailing from the Turonian age of the early Late Cretaceous. The type species is ''Timurlengia euotica''. Discovery From 1944 onwards, tyrannosauroid fossil material consisting of single bones has been described from the Bissekty Formation by Soviet or Russian researchers. In 2004 a team discovered a braincase, which would have anchored the dinosaur's neck muscles and protected its brain and ear canals. The braincase was stored in a cardboard box in the Zoological Institute of the Russian Academy of Sciences, until tyrannosaur expert Steve Brusatte identified it as a distinctive new species in 2014. In 2016, Stephen Louis Brusatte, Alexander Averianov, Hans-Dieter Sues, Amy Muir, and Ian B. Butler named and described the type species ''Timurlengia euotica''. The genus is named after Timurleng, founder of the Timurid Empire in Central Asia. ...
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