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Santanaraptor
''Santanaraptor'' (meaning "Santana Formation thief") is a genus of Tyrannosauroidea, tyrannosauroid Theropoda, theropod dinosaur that lived in South America during the Early Cretaceous (late Aptian-early Albian), about 112 million years ago. Discovery The type species is ''S. placidus'', first described by Alexander Kellner, Kellner in 1999. The species epithet refers to , who founded the . Description The holotype (National Museum of Brazil, MN 4802-V) is a juvenile partial skeleton consisting of three caudal vertebrae with chevrons, ischia, femora, tibia, fibula, pes, and soft tissue. The fossilized tissue includes a thin Epidermis (zoology), epidermis, muscle fibers, and possibly blood vessels. Skin impressions under the left foot are also preserved, showing Scale (anatomy), scales. It was unearthed in 1996 from the Romualdo Formation (Santana Group) in the Ceará State, Northeast Region, Brazil, northeastern Brazil. While primarily known from hindquarter elements, the indiv ...
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Tyrannosauroidea
Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent beginning in the Jurassic Period. By the end of the Cretaceous Period, tyrannosauroids were the dominant large predators in the Northern Hemisphere, culminating in the gigantic ''Tyrannosaurus''. Fossils of tyrannosauroids have been recovered on what are now the continents of North America, Europe and Asia, with fragmentary remains possibly attributable to tyrannosaurs also known from South America and Australia. Tyrannosauroids were bipedal carnivores, as were most theropods, and were characterized by numerous synapomorphy, skeletal features, especially of the skull and pelvis. Early in their existence, tyrannosauroids were small predators with long, three-fingered forelimbs. Late Cretaceous genera became much larger, including some of the ...
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Tyrannosauroidea
Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent beginning in the Jurassic Period. By the end of the Cretaceous Period, tyrannosauroids were the dominant large predators in the Northern Hemisphere, culminating in the gigantic ''Tyrannosaurus''. Fossils of tyrannosauroids have been recovered on what are now the continents of North America, Europe and Asia, with fragmentary remains possibly attributable to tyrannosaurs also known from South America and Australia. Tyrannosauroids were bipedal carnivores, as were most theropods, and were characterized by numerous synapomorphy, skeletal features, especially of the skull and pelvis. Early in their existence, tyrannosauroids were small predators with long, three-fingered forelimbs. Late Cretaceous genera became much larger, including some of the ...
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Santanaraptor Placidus Size Chart
''Santanaraptor'' (meaning "Santana Formation thief") is a genus of tyrannosauroid theropod dinosaur that lived in South America during the Early Cretaceous (late Aptian-early Albian), about 112 million years ago. Discovery The type species is ''S. placidus'', first described by Kellner in 1999. The species epithet refers to , who founded the . Description The holotype ( MN 4802-V) is a juvenile partial skeleton consisting of three caudal vertebrae with chevrons, ischia, femora, tibia, fibula, pes, and soft tissue. The fossilized tissue includes a thin epidermis, muscle fibers, and possibly blood vessels. Skin impressions under the left foot are also preserved, showing scales. It was unearthed in 1996 from the Romualdo Formation (Santana Group) in the Ceará State, northeastern Brazil. While primarily known from hindquarter elements, the individual represented by the fossil may have reached in length and in mass. The fossil consists of bones from the pelvis, hindlimbs, and ta ...
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Romualdo Formation
The Romualdo Formation is a geologic Konservat-Lagerstätte in northeastern Brazil's Araripe Basin where the states of Pernambuco, Piauí and Ceará come together. The geological formation, previously designated as the Romualdo Member of the Santana Formation, named after the village of Santana do Cariri, lies at the base of the Araripe Plateau. It was discovered by Johann Baptist von Spix in 1819. The strata were deposited during the Aptian stage of the Early Cretaceous in a lacustrine rift basin with shallow marine incursions of the proto-Atlantic. At that time, the South Atlantic was opening up in a long narrow shallow sea. The Romualdo Formation earns the designation of Lagerstätte due to an exceedingly well preserved and diverse fossil faunal assemblage. Some 25 species of fossil fishes are often found with stomach contents preserved, enabling paleontologists to study predator–prey relationships in this ecosystem. There are also fine examples of pterosaurs, reptiles a ...
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Alexander Kellner
Alexander Wilhelm Armin Kellner (born September 26, 1961) is a Brazilian geologist and paleontologist who is a leading expert in the field of studying pterosaurs. His research has focused mainly on fossil reptiles from the Cretaceous Period, including extinct dinosaurs and crocodylomorphs. Kellner has over 500 publications to his name, has published more than 160 primary studies and two science books. He has participated in paleontological expeditions to many locations including Brazil, Chile, Iran, the United States, Argentina, China, and Antarctica. His scientific achievements include the description of more than thirty species. For his work he has received several honors and prizes, including the TWAS Prize for Earth Sciences from The World Academy of Sciences and admission to the National Order of Scientific Merit (class Comendador), one Brazil's most prestigious awards. Biography Kellner was born in Vaduz, Liechtenstein, son of a German father and Austrian mother. In hi ...
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Ceará
Ceará (, pronounced locally as or ) is one of the 26 states of Brazil, located in the northeastern part of the country, on the Atlantic coast. It is the eighth-largest Brazilian State by population and the 17th by area. It is also one of the main tourist destinations in Brazil. The state capital is the city of Fortaleza, the country's fourth most populous city. The state has 4.3% of the Brazilian population and produces 2.1% of the Brazilian GDP. Literally, the name ''Ceará'' means "sings the jandaia". According to José de Alencar, one of the most important writers of Brazil and an authority in Tupi Guaraní, ''Ceará'' means turquoise or green waters. The state is best known for its extensive coastline, with of sand. There are also mountains and valleys producing tropical fruits. To the south, on the border of Paraíba, Pernambuco and Piauí, is the National Forest of Araripe. Geography Ceará has an area of . It is bounded on the north by the Atlantic Ocean, ...
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Northeast Region, Brazil
The Northeast Region of Brazil ( pt, Região Nordeste do Brasil; ) is one of the five official and political regions of the country according to the Brazilian Institute of Geography and Statistics. Of Brazil's twenty-six states, it comprises nine: Maranhão, Piauí, Ceará, Rio Grande do Norte, Paraíba, Pernambuco, Alagoas, Sergipe and Bahia, along with the Fernando de Noronha archipelago (formerly a separate territory, now part of Pernambuco). Chiefly known as ''Nordeste'' ("Northeast") in Brazil, this region was the first to be colonized by the Portuguese and other European peoples, playing a crucial role in the country's history. ''Nordestes dialects and rich culture, including its folklore, cuisines, music and literature, became the most easily distinguishable across the country. To this day, ''Nordeste'' is known for its history and culture, as well as for its natural environment and its hot weather. ''Nordeste'' stretches from the Atlantic seaboard in the northeast ...
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Brazil
Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area and the seventh most populous. Its capital is Brasília, and its most populous city is São Paulo. The federation is composed of the union of the 26 States of Brazil, states and the Federal District (Brazil), Federal District. It is the largest country to have Portuguese language, Portuguese as an List of territorial entities where Portuguese is an official language, official language and the only one in the Americas; one of the most Multiculturalism, multicultural and ethnically diverse nations, due to over a century of mass Immigration to Brazil, immigration from around the world; and the most populous Catholic Church by country, Roman Catholic-majority country. Bounded by the Atlantic Ocean on the east, Brazil has a Coastline of Brazi ...
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Dorling Kindersley
Dorling Kindersley Limited (branded as DK) is a British multinational publishing company specialising in illustrated reference books for adults and children in 63 languages. It is part of Penguin Random House, a subsidiary of German media conglomerate Bertelsmann. Established in 1974, DK publishes a range of titles in genres including travel (including DK Eyewitness travel), history, geography, science, space, nature, sports, gardening, cookery and parenting. The worldwide co-CEOs of DK is Paul Kelly and Rebecca Smart. DK has offices in New York, Melbourne, London, Munich, New Delhi, Toronto, Madrid, Beijing, and Jiangmen. DK works with licensing partners such as Disney, LEGO, DC Comics, the Royal Horticultural Society, MasterChef, and the Smithsonian Institution. DK has commissioned Mary Berry, Monty Don, Robert Winston, Huw Richards, and Steve Mould for a range of books. History DK was founded in 1974 by Christopher Dorling and Peter Kindersley in London as a book ...
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Aptian
The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), approximately. The Aptian succeeds the Barremian and precedes the Albian, all part of the Lower/Early Cretaceous. The Aptian partly overlaps the upper part of the Western European Urgonian Stage. The Selli Event, also known as OAE1a, was one of two oceanic anoxic events in the Cretaceous Period, which occurred around 120 Ma and lasted approximately 1 to 1.3 million years. The Aptian extinction was a minor extinction event hypothesized to have occurred around 116 to 117 Ma.Archangelsky, Sergio.The Ticó Flora (Patagonia) and the Aptian Extinction Event" ''Acta Paleobotanica'' 41(2), 2001, pp. 115-22. Stratigraphic definitions The Aptian was named after the small city of Apt in the Provence region of France, which is also known for its cry ...
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Pelvis
The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The pelvic region of the trunk includes the bony pelvis, the pelvic cavity (the space enclosed by the bony pelvis), the pelvic floor, below the pelvic cavity, and the perineum, below the pelvic floor. The pelvic skeleton is formed in the area of the back, by the sacrum and the coccyx and anteriorly and to the left and right sides, by a pair of hip bones. The two hip bones connect the spine with the lower limbs. They are attached to the sacrum posteriorly, connected to each other anteriorly, and joined with the two femurs at the hip joints. The gap enclosed by the bony pelvis, called the pelvic cavity, is the section of the body underneath the abdomen and mainly consists of the reproductive organs (sex organs) and the rectum, while the pelvic f ...
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Scale (anatomy)
In most biological nomenclature, a scale ( grc, λεπίς, lepís; la, squāma) is a small rigid plate that grows out of an animal's skin to provide protection. In lepidopteran (butterfly and moth) species, scales are plates on the surface of the insect wing, and provide coloration. Scales are quite common and have evolved multiple times through convergent evolution, with varying structure and function. Scales are generally classified as part of an organism's integumentary system. There are various types of scales according to shape and to class of animal. Fish scales File:Ganoid scales.png, Ganoid scales on a carboniferous fish ''Amblypterus striatus'' File:Denticules cutanés du requin citron Negaprion brevirostris vus au microscope électronique à balayage.jpg, Placoid scales on a lemon shark (''Negaprion brevirostris'') File:RutilusRutilusScalesLateralLine.JPG, Cycloid scales on a common roach (''Rutilus rutilus'') Fish scales are dermally derived, specifically i ...
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