Labocania
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''Labocania'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
of uncertain affinities from
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
,
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
, which lived 73 million years ago, in the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
stage Stage or stages may refer to: Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly British theatre newspaper * Sta ...
of the late
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
Period. In the summer of 1970, the
National Geographic Society The National Geographic Society (NGS), headquartered in Washington, D.C., United States, is one of the largest non-profit scientific and educational organizations in the world. Founded in 1888, its interests include geography, archaeology, and ...
and the
Los Angeles County Museum of Natural History The Natural History Museum of Los Angeles County is the largest natural and historical museum in the western United States. Its collections include nearly 35 million specimens and artifacts and cover 4.5 billion years of history. This large coll ...
organized a joint
paleontological Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (geology), epoch (roughly 11,700 years before present). It includes t ...
expedition, led by
geologist A geologist is a scientist who studies the solid, liquid, and gaseous matter that constitutes Earth and other terrestrial planets, as well as the processes that shape them. Geologists usually study geology, earth science, or geophysics, althou ...
William J. Morris, to the Arroyo del Rosario in Baja California. While prospecting, volunteer Harley James Garbani discovered the skeleton of a theropod north of Punta Baja near Cerro Rayado. Garbani excavated the site in 1970 and 1971. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
, ''Labocania anomala'', was described and named by
Ralph Molnar Ralph E. Molnar is a paleontologist who had been Curator of Mammals at the Queensland Museum and more recently associated with the Museum of Northern Arizona. He is also a research associate at the Texas natural Science Centre. He co-authored descr ...
in 1974. The generic name references the
La Bocana Roja Formation La Bocana Roja Formation is a geological formation in Baja California, Mexico whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Di ...
, named after ''la Bocana Roja'', "the red estuary". The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
means "anomalous" in Latin, in reference to the distinctive build. The holotype, LACM 20877, was found in a layer of the La Bocana Roja Formation, dating from the late Campanian, about 73 million years old. It consists of a very fragmentary skeleton with skull elements, including a right quadrate, a left frontal, a piece of the left maxilla, a fragment of the dentarium, a chevron, the upper parts of both ischia, the middle shaft of the right pubis, most of the second right metatarsal, a pedal phalanx and several loose teeth. The elements were not articulated, dispersed over a surface of about two square metres, and strongly weathered. The remains were mixed with the ribs of Hadrosauroidea. Though its exact size is hard to establish, ''Labocania'' was probably a medium-sized carnivore, about long. Gregory S. Paul estimated its length at 7 meters (23 ft), its weight at 1.5 tonnes (1.65 short tons) in 2010. In 2016 Molina-Pérez and Larramendi gave a higher estimation of 8.2 meters (27 ft) and 2.6 tonnes (2.86 short tons). The cranial elements are very robust, and the frontals in particular are strongly thickened. The teeth of the maxilla are gradually recurving and rather flat; those of the premaxilla do not have a D-shaped cross-section. Because ''Labocania'' is based on fragmentary material, its affinities are uncertain. Molnar noted certain similarities between ''Labocania'' and
tyrannosaurids Tyrannosauridae (or tyrannosaurids, meaning "tyrant lizards") is a family (biology), family of coelurosaurian Theropoda, theropod dinosaurs that comprises two subfamilies containing up to thirteen genus, genera, including the eponymous ''Tyrannos ...
, especially in the form of the
ischium The ischium () form ...
which features a low triangular obturator process and a circular lateral scar on the upper end, but he did not assign ''Labocania'' to any family, placing it as
Theropoda Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ca ...
'' incertae sedis''. Molnar especially compared ''Labocania'' with '' Indosaurus'' and " Chilantaisaurus" ''maortuensis'', later made the separate genus '' Shaochilong''. ''Labocania'' was considered as a possible tyrannosauroid in the 2004 review of the group by
Thomas R. Holtz Jr. Thomas Richard Holtz Jr. (born September 13, 1965) is an American vertebrate palaeontologist, author, and principal lecturer at the University of Maryland's Department of Geology. He has published extensively on the phylogeny, morphology, ecomorp ...
, who, however, pointed out that the similarities with the Tyrannosauridae were shared with the Coelurosauria in general—no tyrannosauroid
synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have ...
were present—and that ''Labocania'' also showed some abelisaurid traits such as the thick frontals and a reclining quadrate. On the other hand, the L-shaped chevron and the flattened outer side of the second metatarsal indicated a position in the Tetanurae.


References

{{Taxonbar, from=Q1827222 Prehistoric neotheropods Campanian genus first appearances Campanian genus extinctions Late Cretaceous dinosaurs of North America Cretaceous Mexico Fossils of Mexico Fossil taxa described in 1974 Taxa named by Ralph Molnar