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Timurlengia
''Timurlengia'' is an extinct genus of tyrannosauroid theropod dinosaur found in Uzbekistan, in the Bissekty Formation in the Kyzylkum Desert, hailing from the Turonian age of the early Late Cretaceous. The type species is ''Timurlengia euotica''. Discovery From 1944 onwards, tyrannosauroid fossil material consisting of single bones has been described from the Bissekty Formation by Soviet or Russian researchers. In 2004 a team discovered a braincase, which would have anchored the dinosaur's neck muscles and protected its brain and ear canals. The braincase was stored in a cardboard box in the Zoological Institute of the Russian Academy of Sciences, until tyrannosaur expert Steve Brusatte identified it as a distinctive new species in 2014. In 2016, Stephen Louis Brusatte, Alexander Averianov, Hans-Dieter Sues, Amy Muir, and Ian B. Butler named and described the type species ''Timurlengia euotica''. The genus is named after Timurleng, founder of the Timurid Empire in Central Asia. ...
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Timurlengia Scale
''Timurlengia'' is an extinct genus of tyrannosauroid theropod dinosaur found in Uzbekistan, in the Bissekty Formation in the Kyzylkum Desert, hailing from the Turonian age of the early Late Cretaceous. The type species is ''Timurlengia euotica''. Discovery From 1944 onwards, tyrannosauroid fossil material consisting of single bones has been described from the Bissekty Formation by Soviet or Russian researchers. In 2004 a team discovered a braincase, which would have anchored the dinosaur's neck muscles and protected its brain and ear canals. The braincase was stored in a cardboard box in the Zoological Institute of the Russian Academy of Sciences, until tyrannosaur expert Steve Brusatte identified it as a distinctive new species in 2014. In 2016, Stephen Louis Brusatte, Alexander Averianov, Hans-Dieter Sues, Amy Muir, and Ian B. Butler named and described the type species ''Timurlengia euotica''. The genus is named after Timurleng, founder of the Timurid Empire in Central Asia. ...
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Tyrannosauroid
Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent beginning in the Jurassic Period. By the end of the Cretaceous Period, tyrannosauroids were the dominant large predators in the Northern Hemisphere, culminating in the gigantic ''Tyrannosaurus''. Fossils of tyrannosauroids have been recovered on what are now the continents of North America, Europe and Asia, with fragmentary remains possibly attributable to tyrannosaurs also known from South America and Australia. Tyrannosauroids were bipedal carnivores, as were most theropods, and were characterized by numerous skeletal features, especially of the skull and pelvis. Early in their existence, tyrannosauroids were small predators with long, three-fingered forelimbs. Late Cretaceous genera became much larger, including some of the largest ...
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Bissekty Formation
The Bissekty Formation (sometimes referred to as Bissekt) is a geologic formation and Lagerstätte which crops out in the Kyzyl Kum desert of Uzbekistan, and dates to the Late Cretaceous Period. Laid down in the mid to late Turonian, it is dated to about 92 to 90 Ma (million years ago). Description The lithology of the sediment largely consists of cross bedded sandstones with interbeds of massive sandstone, well cemented intraformational conglomerate, siltstones and mudstones. Most of the fossils are found as clasts within the conglomerates. Fossil content The Bissekty Formation is characterised by a mix of marine, brackish, freshwater, and terrestrial animal fossils. This stands in contrast the strictly marine fossils found in the underlying Dzheirantui Formation, and indicates that the Bissekty was formed during the regression of a saltwater sea. The coastline expanded inland again in the upper portion of the Bissekty, represented by a proportional increase of fully aquat ...
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Timur
Timur ; chg, ''Aqsaq Temür'', 'Timur the Lame') or as ''Sahib-i-Qiran'' ( 'Lord of the Auspicious Conjunction'), his epithet. ( chg, ''Temür'', 'Iron'; 9 April 133617–19 February 1405), later Timūr Gurkānī ( chg, ''Temür Küregen''), was a Turco-Mongol conqueror who founded the Timurid Empire in and around modern-day Afghanistan, Iran, and Central Asia, becoming the first ruler of the Timurid dynasty. An undefeated commander, he is widely regarded as one of the greatest military leaders and tacticians in history, as well as one of the most brutal. Timur is also considered a great patron of art and architecture as he interacted with intellectuals such as Ibn Khaldun, Hafez, and Hafiz-i Abru and his reign introduced the Timurid Renaissance. Born into the Barlas confederation in Transoxiana (in modern-day Uzbekistan) on 9 April 1336, Timur gained control of the western Chagatai Khanate by 1370. From that base, he led military campaigns across Western, South, and ...
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Hans-Dieter Sues
Hans-Dieter Sues (born January 13, 1956) is a German-born American paleontologist who is Senior Scientist and Curator of Vertebrate Paleontology at the National Museum of Natural History of the Smithsonian Institution in Washington, DC. He received his education at Johannes Gutenberg-Universität Mainz (University of Mainz), University of Alberta, and Harvard University (Ph.D., 1984). Before assuming his present position, Sues worked at the Royal Ontario Museum and the University of Toronto and at the Carnegie Museum of Natural History in Pittsburgh. He is interested in the diversity, paleoecology, and evolutionary history of Paleozoic and Mesozoic tetrapods, especially archosaurian reptiles and cynodont therapsids, and the history of biology and paleontology. Sues has discovered numerous new dinosaurs and other extinct terrestrial vertebrates in Paleozoic and Mesozoic continental strata in North America and Europe. He has authored or co-authored over 150 articles and book chapte ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Maxilla
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxillary bones are fused at the intermaxillary suture, forming the anterior nasal spine. This is similar to the mandible (lower jaw), which is also a fusion of two mandibular bones at the mandibular symphysis. The mandible is the movable part of the jaw. Structure In humans, the maxilla consists of: * The body of the maxilla * Four processes ** the zygomatic process ** the frontal process of maxilla ** the alveolar process ** the palatine process * three surfaces – anterior, posterior, medial * the Infraorbital foramen * the maxillary sinus * the incisive foramen Articulations Each maxilla articulates with nine bones: * two of the cranium: the frontal and ethmoid * seven of the face: the nasal, zygomatic, lacrimal, inferior n ...
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Neural Arch
The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic irregular bone whose complex structure is composed primarily of bone, and secondarily of hyaline cartilage. They show variation in the proportion contributed by these two tissue types; such variations correlate on one hand with the cerebral/caudal rank (i.e., location within the backbone), and on the other with phylogenetic differences among the vertebrate taxa. The basic configuration of a vertebra varies, but the bone is its ''body'', with the central part of the body constituting the ''centrum''. The upper (closer to) and lower (further from), respectively, the cranium and its central nervous system surfaces of the vertebra body support attachment to the intervertebral discs. The posterior part of a vertebra forms a vertebral arch (in ...
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Taxon
In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and given a particular ranking, especially if and when it is accepted or becomes established. It is very common, however, for taxonomists to remain at odds over what belongs to a taxon and the criteria used for inclusion. If a taxon is given a formal scientific name, its use is then governed by one of the nomenclature codes specifying which scientific name is correct for a particular grouping. Initial attempts at classifying and ordering organisms (plants and animals) were set forth in Carl Linnaeus's Linnaean taxonomy, system in ''Systema Naturae'', 10th edition (1758), as well as an unpublished work by Bernard de Jussieu, Bernard and Antoine Laurent de Jussieu. The idea of a unit-based system of biological classification was first mad ...
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Supraoccipital Bone
The occipital bone () is a cranial dermal bone and the main bone of the occiput (back and lower part of the skull). It is trapezoidal in shape and curved on itself like a shallow dish. The occipital bone overlies the occipital lobes of the cerebrum. At the base of skull in the occipital bone, there is a large oval opening called the foramen magnum, which allows the passage of the spinal cord. Like the other cranial bones, it is classed as a flat bone. Due to its many attachments and features, the occipital bone is described in terms of separate parts. From its front to the back is the basilar part, also called the basioccipital, at the sides of the foramen magnum are the lateral parts, also called the exoccipitals, and the back is named as the squamous part. The basilar part is a thick, somewhat quadrilateral piece in front of the foramen magnum and directed towards the pharynx. The squamous part is the curved, expanded plate behind the foramen magnum and is the largest part o ...
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Horse
The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million years from a small multi-toed creature, ''Eohippus'', into the large, single-toed animal of today. Humans began domesticating horses around 4000 BCE, and their domestication is believed to have been widespread by 3000 BCE. Horses in the subspecies ''caballus'' are domesticated, although some domesticated populations live in the wild as feral horses. These feral populations are not true wild horses, as this term is used to describe horses that have never been domesticated. There is an extensive, specialized vocabulary used to describe equine-related concepts, covering everything from anatomy to life stages, size, colors, markings, breeds, locomotion, and behavior. Horses are adapted to run, allowing them to quickly escape predators, and po ...
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Specific Name (zoology)
In zoological nomenclature, the specific name (also specific epithet or species epithet) is the second part (the second name) within the scientific name of a species (a binomen). The first part of the name of a species is the name of the genus or the generic name. The rules and regulations governing the giving of a new species name are explained in the article species description. For example, the scientific name for humans is ''Homo sapiens'', which is the species name, consisting of two names: ''Homo'' is the " generic name" (the name of the genus) and ''sapiens'' is the "specific name". Historically, ''specific name'' referred to the combination of what are now called the generic and specific names. Carl Linnaeus, who formalized binomial nomenclature, made explicit distinctions between specific, generic, and trivial names. The generic name was that of the genus, the first in the binomial, the trivial name was the second name in the binomial, and the specific the proper term for ...
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