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Pareiasauria
Pareiasaurs (meaning "cheek lizards") are an extinct clade of large, herbivorous parareptiles. Members of the group were armoured with scutes which covered large areas of the body. They first appeared in southern Pangea during the Middle Permian, before becoming globally distributed during the Late Permian. Pareiasaurs were the largest reptiles of the Permian, reaching sizes equivalent to those of contemporary therapsids. Pareiasaurs became extinct at the end of the Permian during the Permian-Triassic extinction event. Description Pareiasaurs ranged in size from long, and may have weighed up to . They were stocky, with short tails, small heads, robust limbs, and broad feet. The cow-sized species '' Bunostegos'', which lived 260 million years ago, is the earliest known example of a tetrapod with a fully erect posture as its legs were positioned directly under its body. Pareiasaurs were protected by bony scutes called osteoderms that were set into the skin. Their heavy skulls wer ...
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Bunostegos
''Bunostegos'' ("knobbly kullroof") is an extinct genus of pareiasaur parareptile from the Late Permian of the Agadez Region in Niger. The type species, ''Bunostegos akokanensis'', was named from the Moradi Formation in 2003. It was a cow-sized animal with a distinctive skull that had large bony knobs, similar in form to those of other pareiasaurs but far larger. The species appears to have lived in a desert in the centre of the supercontinent of Pangaea. Analysis of the limb bones (including the scapulocoracoid, humerus, radius, ulna, pelvis, and femur) was published in 2015, and revealed that ''Bunostegos'' walked upright on four limbs, with the body held above ground. This new information directly suggests that it could be the first tetrapod with a fully erect gait. Description The animal has been described as about the size of a modern cow with a knobbly skull and bony plate armor on its back." Its teeth show it to have been a plant eater. It lived in an isolated desert ...
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Velosauria
Velosauria is a group of pareiasaur reptiles that existed in the late Permian period. They ranged in size from the 50-centimeter-long '' Pumiliopareia'' to the 3-meter-long '' Scutosaurus''. Velosaurs were some of the largest reptiles of their time. Description Velosaurs were large reptiles that were characterized by their short tails, small heads, broad feet, and their legs, which were directly in between sprawling and semi-erect. Velosaurs' bodies were covered in osteoderms and smaller scales all over. Their heads came in all sorts of different shapes; from the cheek-frilled '' Scutosaurus'' to the spiky-headed '' Elginia'' to the nose-horned ''Arganaceras ''Arganaceras'' ("Argana horn") is a medium-sized pareiasaur from the Late Permian Ikakern Formation of Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It ove ...''. They were herbivorous, and it is believed that their large bodies hou ...
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Scutosaurus
''Scutosaurus'' ("shield lizard") is an extinct genus of pareiasaur parareptiles. Its genus name refers to large plates of armor scattered across its body. It was a large anapsid reptile that, unlike most reptiles, held its legs underneath its body to support its great weight. Fossils have been found in the Sokolki Assemblage Zone of the Malokinelskaya Formation in European Russia, close to the Ural Mountains, dating to the late Permian (Lopingian) between 264 and 252 million years ago. Research history The first fossils were uncovered by Russian paleontologist Vladimir Prokhorovich Amalitskii while documenting plant and animal species in the Upper Permian sediments in the Northern Dvina River, Arkhangelsk District, Northern European Russia. Amalitskii had discovered the site in 1899, and he and his wife Anne Amalitskii continued to oversee excavation until 1914, recovering numerous nearly complete and articulated (in their natural position) skeletons belonging to a menage ...
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Parasaurus
''Parasaurus'' (meaning "near lizard") is a genus of pareiasaur known from fossils collected in the Kupferschiefer in Germany (Hesse, Thuringia and Lower Saxony), dating to the Late Permian (Wuchiapingian). The type species, ''Parasaurus geinitzi'', described by Hermann von Meyer in 1857, was the first pareiasaur ever described. The seven known specimens were redescribed in 2008.L. A. Tsuji and J. Müller. (2008). A Re-evaluation of ''Parasaurus geinitzi'', the first named pareiasaur (Amniota, Parareptilia). Canadian Journal of Earth Sciences 45(10):1111-1121 Discovery and naming As early as 1848, pareiasaur fossils have been reported from Germany; some of these fossils may have belonged to ''Parasaurus''. It was not until 1857 when von Meyer described these fossils and created the ''Parasaurus'' genus.Meyer, H.V. (1857). Beiträge zur näheren Kenntnis fossiler Reptilien. ''Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde'', 1857: 103–104. von Meyer c ...
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Elginia
''Elginia'' is an extinct genus of pareiasaurid known from the Late Permian of Scotland and China. It was named for the area around Elgin in Scotland, which has yielded many fossils referred to as the Elgin Reptiles. Discovery The type species of ''Elginia, Elginia mirabilis,'' was first described in 1893 by E.T. Newton, after fellow geologists John Horne and Archibald Geikie informed him of several unusual specimens stored at the Elgin Museum in Scotland. The specimens were collected several years earlier from the coarse sandstones of the nearby Cutties Hillock Quarry. The quarry’s sandstone, the Cutties Hillock Sandstone Formation, is often (but not always) considered equivalent to the otherwise fossil-poor Hopeman Sandstone Formation of broader Scottish geology. Tetrapod remains occupy a narrow section of the sandstone, lying above a pebbly layer. The sandstones of Cutties Hillock were deposited at the very end of the Permian Period, based on a dicynodont fauna simi ...
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Middle Permian
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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Parareptilia
Parareptilia ("at the side of reptiles") is a subclass or clade of basal sauropsids (reptiles), typically considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near the end of the Carboniferous period and achieved their highest diversity during the Permian period. Several ecological innovations were first accomplished by parareptiles among reptiles. These include the first reptiles to return to marine ecosystems (mesosaurs), the first bipedal reptiles ( bolosaurids such as ''Eudibamus''), the first reptiles with advanced hearing systems ( nycteroleterids and others), and the first large herbivorous reptiles (the pareiasaurs). The only parareptiles to survive into the Triassic period were the procolophonoids, a group of small generalists, omnivores, and herbivores. The largest family of procolophonoids, the procolophonids, rediversified in the Triassic, but subsequently declined and became extinct by the ...
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Scute
A scute or scutum (Latin: ''scutum''; plural: ''scuta'' "shield") is a bony external plate or scale overlaid with horn, as on the shell of a turtle, the skin of crocodilians, and the feet of birds. The term is also used to describe the anterior portion of the mesonotum in insects as well as some arachnids (e.g., the family Ixodidae, the scale ticks). Properties Scutes are similar to scales and serve the same function. Unlike the scales of lizards and snakes, which are formed from the epidermis, scutes are formed in the lower vascular layer of the skin and the epidermal element is only the top surface . Forming in the living dermis, the scutes produce a horny outer layer that is superficially similar to that of scales. Scutes will usually not overlap as snake scales (but see the pangolin). The outer keratin layer is shed piecemeal, and not in one continuous layer of skin as seen in snakes or lizards. The dermal base may contain bone and produce dermal armour. Scutes with a bony ...
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Pangaea
Pangaea or Pangea () was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from the earlier continental units of Gondwana, Euramerica and Siberia during the Carboniferous approximately 335 million years ago, and began to break apart about 200 million years ago, at the end of the Triassic and beginning of the Jurassic. In contrast to the present Earth and its distribution of continental mass, Pangaea was centred on the equator and surrounded by the superocean Panthalassa and the Paleo-Tethys Ocean, Paleo-Tethys and subsequent Tethys Ocean, Tethys Oceans. Pangaea is the most recent supercontinent to have existed and the first to be reconstructed by geologists. Origin of the concept The name "Pangaea" is derived from Ancient Greek ''pan'' (, "all, entire, whole") and ''Gaia (mythology), Gaia'' or Gaea (, "Mother goddess, Mother Earth, land"). The concept that the continents once formed a contiguous land mass was hypothesised, with ...
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Therapsid
Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more underneath the body, as opposed to the sprawling posture of many reptiles and salamanders. Therapsids evolved from "pelycosaurs", specifically within the Sphenacodontia, more than 279.5 million years ago. They replaced the "pelycosaurs" as the dominant large land animals in the Middle Permian through to the Early Triassic. In the aftermath of the Permian–Triassic extinction event, therapsids declined in relative importance to the rapidly diversifying reptiles during the Middle Triassic. The therapsids include the cynodonts, the group that gave rise to mammals ( Mammaliaformes) in the Late Triassic, around 225 million years ago. Of the non-mammalian therapsids, only cynodonts survived beyond the end of the Triassic, with the only other ...
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Bradysaurus BW
''Bradysaurus'' was a large, early and common pareiasaur, the fossils of which are known from the ''Tapinocephalus'' Assemblage Zone (Capitanian age) of the South African Karoo. Along with the similarly large dinocephalia, the bradysaurs constituted the herbivorous megafauna of the late Middle Permian Period. In life they were probably slow, clumsy and inoffensive animals, that had evolved a covering of armoured scutes to protect them against their predators, the gorgonopsians. Description ''Bradysaurus'' was in length and half a tonne to a tonne in weight. The skull was large (about 42 to 48 centimeters long), broad and rounded at the front. It was coarsely sculptured and knobby, with the sutures between the bones not clearly visible. The marginal teeth were high-crowned, with only a few cusps, which is a primitive characteristic. The feet were short and broad, the phalangeal count being 2,3,3,3,2 on the fore-foot and 2,3,3,4,3 on the hind. The whole body is protect ...
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