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Parasaurus
''Parasaurus'' (meaning "near lizard") is a genus of pareiasaur known from fossils collected in the Kupferschiefer in Germany (Hesse, Thuringia and Lower Saxony), dating to the Late Permian (Wuchiapingian). The type species, ''Parasaurus geinitzi'', described by Hermann von Meyer in 1857, was the first pareiasaur ever described. The seven known specimens were redescribed in 2008.L. A. Tsuji and J. Müller. (2008). A Re-evaluation of ''Parasaurus geinitzi'', the first named pareiasaur (Amniota, Parareptilia). Canadian Journal of Earth Sciences 45(10):1111-1121 Discovery and naming As early as 1848, pareiasaur fossils have been reported from Germany; some of these fossils may have belonged to ''Parasaurus''. It was not until 1857 when von Meyer described these fossils and created the ''Parasaurus'' genus.Meyer, H.V. (1857). Beiträge zur näheren Kenntnis fossiler Reptilien. ''Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde'', 1857: 103–104. von Meyer c ...
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Kupferschiefer
The Kupferschiefer (German for Copper Shale, also called Copper Slate) or Kupfermergel (Copper Marl), (T1 or Z1) is an extensive and remarkable sedimentary unit in Central Europe. The relatively monotonous succession is typically and maximum thick, but extends over an area of across the Southern Permian Basin. The Kupferschiefer can be found in outcrop or in the subsurface straddling six countries, including parts of the southern North Sea. The lateral equivalent outcropping in England is called Marl Slate. Despite its distinctive nature, the Kupferschiefer is not ranked as a formation but is officially declared a sub-unit of the Werra Formation, the lowest formation of the Zechstein Group, overlying the Rotliegend Group. The unit has been dated to 257.3 ± 1.6 Ma, placing it in the Wuchiapingian stage of the Late Permian. The Kupferschiefer comprises black shales, bituminous marls, mudstones and limestones deposited mostly in an open marine setting, with the borders of its ...
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Pareiasaur
Pareiasaurs (meaning "cheek lizards") are an extinct clade of large, herbivorous parareptiles. Members of the group were armoured with scutes which covered large areas of the body. They first appeared in southern Pangea during the Middle Permian, before becoming globally distributed during the Late Permian. Pareiasaurs were the largest reptiles of the Permian, reaching sizes equivalent to those of contemporary therapsids. Pareiasaurs became extinct at the end of the Permian during the Permian-Triassic extinction event. Description Pareiasaurs ranged in size from long, and may have weighed up to . They were stocky, with short tails, small heads, robust limbs, and broad feet. The cow-sized species '' Bunostegos'', which lived 260 million years ago, is the earliest known example of a tetrapod with a fully erect posture as its legs were positioned directly under its body. Pareiasaurs were protected by bony scutes called osteoderms that were set into the skin. Their heavy skulls wer ...
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Wuchiapingian
In the geologic timescale, the Wuchiapingian or Wujiapingian (from in the Liangshan area of Hanzhong, Shaanxi Province) is an age or stage of the Permian. It is also the lower or earlier of two subdivisions of the Lopingian Epoch or Series. The Wuchiapingian spans the time between and million years ago (Ma). It was preceded by the Capitanian and followed by the Changhsingian. Regional stages with which the Wuchiapingian is coeval or overlaps include the Djulfian or Dzhulfian, Longtanian, Rustlerian, Saladoan, and Castilian. Stratigraphic definitions The Wuchiapingian was first used in 1962, when the Lopingian Series of southwestern China was divided in the Changhsingian and Wuchiapingian Formations. In 1973 the Wuchiapingian was first used as a chronostratigraphic unit (i.e. a stage, as opposed to a formation, which is a lithostratigraphic unit). The base of the Wuchiapingian Stage is defined as the place in the stratigraphic record where the conodont species ''Clarkina po ...
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Velosauria
Velosauria is a group of pareiasaur reptiles that existed in the late Permian period. They ranged in size from the 50-centimeter-long '' Pumiliopareia'' to the 3-meter-long '' Scutosaurus''. Velosaurs were some of the largest reptiles of their time. Description Velosaurs were large reptiles that were characterized by their short tails, small heads, broad feet, and their legs, which were directly in between sprawling and semi-erect. Velosaurs' bodies were covered in osteoderms and smaller scales all over. Their heads came in all sorts of different shapes; from the cheek-frilled '' Scutosaurus'' to the spiky-headed '' Elginia'' to the nose-horned ''Arganaceras ''Arganaceras'' ("Argana horn") is a medium-sized pareiasaur from the Late Permian Ikakern Formation of Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It ove ...''. They were herbivorous, and it is believed that their large bodies hou ...
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Bunostegos
''Bunostegos'' ("knobbly kullroof") is an extinct genus of pareiasaur parareptile from the Late Permian of the Agadez Region in Niger. The type species, ''Bunostegos akokanensis'', was named from the Moradi Formation in 2003. It was a cow-sized animal with a distinctive skull that had large bony knobs, similar in form to those of other pareiasaurs but far larger. The species appears to have lived in a desert in the centre of the supercontinent of Pangaea. Analysis of the limb bones (including the scapulocoracoid, humerus, radius, ulna, pelvis, and femur) was published in 2015, and revealed that ''Bunostegos'' walked upright on four limbs, with the body held above ground. This new information directly suggests that it could be the first tetrapod with a fully erect gait. Description The animal has been described as about the size of a modern cow with a knobbly skull and bony plate armor on its back." Its teeth show it to have been a plant eater. It lived in an isolated desert ...
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Embrithosaurus
''Embrithosaurus'' was a pareiasaur from the Permian of South Africa. Description ''Embrithosaurus'' was in length and in weight. The skull is relatively deep and narrow. The body is lightly armoured with thin, smooth dermal scutes. Species * ''E. schwarzi'' (Watson, 1914). The type species. This is the most advanced species of this genus, as indicated by the teeth, which have nine cusps (in three groups of three). In cladistic analyses it is used as the monotypal species for the genus. * ''E. alexanderi'' (Haughton and Boonstra, 1929). This species was made the type for ''"Dolichopareia"''. As the name indicates, the skull is long and narrow. This would seem to indicate a different lifestyle or diet to other parieasaurs. More recently, it has been used as the monotypal species for the genus ''Nochelesaurus'' (it is not clear what the status of ''Embrithosaurus strubeni'' is, this may be a further transitional species). In cladistic analyses, this species is phylogenetically in ...
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Arganaceras
''Arganaceras'' ("Argana horn") is a medium-sized pareiasaur from the Late Permian Ikakern Formation of Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to .... It was about in length and had a horn-like structure on its snout. References Pareiasaurs Lopingian reptiles of Africa Fossils of Morocco Fossil taxa described in 2005 Prehistoric reptile genera {{permian-reptile-stub ...
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Pareiasuchus Nasicornis
''Pareiasaurus'' is an extinct genus of pareiasauromorph reptile from the Permian period. It was a typical member of its family, the pareiasaurids, which take their name from this genus. Fossils have been found in the Beaufort Group. Description ''Pareiasaurus'' is a large quadruped, about long, with elephantine legs, walking in a typically reptilian posture. The skull is broad and the snout short. Its skull had several spine- and wart-like protrusions. ''Pareiasauruss leaf-shaped teeth, ideal for biting through tough plant fibers, indicate it was a herbivore. Even the palate The palate () is the roof of the mouth in humans and other mammals. It separates the oral cavity from the nasal cavity. A similar structure is found in crocodilians, but in most other tetrapods, the oral and nasal cavities are not truly sepa ... had teeth. Species ''P. nasicornis'' (Haughton and Boonstra, 1929) is from the ''Tropidostoma'' Zone, Karoo basin, South Africa. This early form is one ...
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Pareiasuchus Peringueyi
''Pareiasaurus'' is an extinct genus of pareiasauromorph reptile from the Permian period. It was a typical member of its family, the pareiasaurids, which take their name from this genus. Fossils have been found in the Beaufort Group. Description ''Pareiasaurus'' is a large quadruped, about long, with elephantine legs, walking in a typically reptilian posture. The skull is broad and the snout short. Its skull had several spine- and wart-like protrusions. ''Pareiasauruss leaf-shaped teeth, ideal for biting through tough plant fibers, indicate it was a herbivore. Even the palate The palate () is the roof of the mouth in humans and other mammals. It separates the oral cavity from the nasal cavity. A similar structure is found in crocodilians, but in most other tetrapods, the oral and nasal cavities are not truly sepa ... had teeth. Species ''P. nasicornis'' (Haughton and Boonstra, 1929) is from the ''Tropidostoma'' Zone, Karoo basin, South Africa. This early form is one ...
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Shihtienfenia
''Shihtienfenia'' was a pareiasaurid parareptile from the Late Permian of China. Species Lee (1997) refers to ''S. xuecunensis'' as a metaspecies lacking the autapomorphies of ''Shihtienfenia''. Tsuji & Müller (2009) seem to consider it a valid taxon for cladistic analysis, and like Lee 1997 place the two Chinese species close to '' Pareiasuchus''. ''S. permica'' (Young and Yeh, 1963); The skull of this pareiasaur is unknown. It is known originally from a number of isolated vertebrae, jaws, and limb-bones and an incomplete skeleton, all from the Shiqianfeng locality near Baode, Shanxi Shanxi (; ; formerly romanised as Shansi) is a landlocked province of the People's Republic of China and is part of the North China region. The capital and largest city of the province is Taiyuan, while its next most populated prefecture-lev ..., part of the Sunjiagou Formation. ''Shanshisaurus xuecunensis'' Cheng, 1980 and ''Huanghesaurus liuliensis'' Gao, 1983 are synonyms. ''S. comple ...
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Shansisaurus
''Shihtienfenia'' was a pareiasaurid parareptile from the Late Permian of China. Species Lee (1997) refers to ''S. xuecunensis'' as a metaspecies lacking the autapomorphies of ''Shihtienfenia''. Tsuji & Müller (2009) seem to consider it a valid taxon for cladistic analysis, and like Lee 1997 place the two Chinese species close to '' Pareiasuchus''. ''S. permica'' (Young and Yeh, 1963); The skull of this pareiasaur is unknown. It is known originally from a number of isolated vertebrae, jaws, and limb-bones and an incomplete skeleton, all from the Shiqianfeng locality near Baode, Shanxi Shanxi (; ; formerly romanised as Shansi) is a landlocked province of the People's Republic of China and is part of the North China region. The capital and largest city of the province is Taiyuan, while its next most populated prefecture-lev ..., part of the Sunjiagou Formation. ''Shanshisaurus xuecunensis'' Cheng, 1980 and ''Huanghesaurus liuliensis'' Gao, 1983 are synonyms. ''S. comple ...
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Pumiliopareia
''Pumiliopareia'' is an extinct genus of pareiasauridae, pareiasaurid parareptile from the Permian period of South Africa. It is known from a complete skeleton with osteoderms. Description ''Pumiliopareia'' was about 50 cm in length with a 12 cm skull. It is the smallest known member of the pareiasaurs, measuring only a fifth as long as some of its larger relatives. Like ''Anthodon (reptile), Anthodon'', its body was entirely covered with osteoderms. In analyses that support a pareiasaur origin of turtles, the sister taxon of the testudines. However it specifically shares with turtles a single trait only: Ribs greatly expanded anteroposteriorly (i.e. wide). Classification Originally included under the genus ''Nanoparia'', it was given its own name by Lee 1997 who found it did not form a clade with ''Nanoparia luckhoffi'', the type species of that genus, and preferred to have monophyletic genera. ''Nanoparia'' may still be a paraphyletic genus, which is allowed in Linan ...
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