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Dictyoptera
Dictyoptera (from Greek δίκτυον ''diktyon'' "net" and πτερόν ''pteron'' "wing") is an insect superorder that includes two extant orders of polyneopterous insects: the order Blattodea (termites and cockroaches together) and the order Mantodea (mantises). While all modern Dictyoptera have short ovipositors, the oldest fossils of Dictyoptera have long ovipositors, much like members of the Orthoptera. Classification and phylogeny The use of the term Dictyoptera has changed over the years, and while largely out of use for much of the last century, it is becoming more widely used. It has usually been considered a superorder, with Isoptera, Blattodea and Mantodea being its three orders. In some classifications, however, Dictyoptera is shifted to order status and in others the order Isoptera has been subsumed under Blattodea while retaining Dictyoptera as a superorder. Regardless, in all classifications the constituent groups are the same, just treated at different rank. T ...
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Umenocoleoidea
Dictyoptera (from Greek δίκτυον ''diktyon'' "net" and πτερόν ''pteron'' "wing") is an insect superorder that includes two extant orders of polyneopterous insects: the order Blattodea (termites and cockroaches together) and the order Mantodea (mantises). While all modern Dictyoptera have short ovipositors, the oldest fossils of Dictyoptera have long ovipositors, much like members of the Orthoptera. Classification and phylogeny The use of the term Dictyoptera has changed over the years, and while largely out of use for much of the last century, it is becoming more widely used. It has usually been considered a superorder, with Isoptera, Blattodea and Mantodea being its three orders. In some classifications, however, Dictyoptera is shifted to order status and in others the order Isoptera has been subsumed under Blattodea while retaining Dictyoptera as a superorder. Regardless, in all classifications the constituent groups are the same, just treated at different rank. T ...
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Cockroach
Cockroaches (or roaches) are a paraphyletic group of insects belonging to Blattodea, containing all members of the group except termites. About 30 cockroach species out of 4,600 are associated with human habitats. Some species are well-known as pests. The cockroaches are an ancient group, with their ancestors, known as " roachoids", originating during the Carboniferous period, some 320 million years ago. Those early ancestors, however, lacked the internal ovipositors of modern roaches. Cockroaches are somewhat generalized insects lacking special adaptations (such as the sucking mouthparts of aphids and other true bugs); they have chewing mouthparts and are probably among the most primitive of living Neopteran insects. They are common and hardy insects capable of tolerating a wide range of climates, from Arctic cold to tropical heat. Tropical cockroaches are often much larger than temperate species. Modern cockroaches are not considered to be a monophyletic group, as it has be ...
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Mantodea
Mantises are an order (Mantodea) of insects that contains over 2,400 species in about 460 genera in 33 families. The largest family is the Mantidae ("mantids"). Mantises are distributed worldwide in temperate and tropical habitats. They have triangular heads with bulging eyes supported on flexible necks. Their elongated bodies may or may not have wings, but all Mantodea have forelegs that are greatly enlarged and adapted for catching and gripping prey; their upright posture, while remaining stationary with forearms folded, has led to the common name praying mantis. The closest relatives of mantises are termites and cockroaches (Blattodea), which are all within the superorder Dictyoptera. Mantises are sometimes confused with stick insects (Phasmatodea), other elongated insects such as grasshoppers (Orthoptera), or other more distantly related insects with raptorial forelegs such as mantisflies (Mantispidae). Mantises are mostly ambush predators, but a few ground-dwelling spe ...
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Termite
Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattodea (along with cockroaches). Termites were once classified in a separate order from cockroaches, but recent phylogenetic studies indicate that they evolved from cockroaches, as they are deeply nested within the group, and the sister group to wood eating cockroaches of the genus ''Cryptocercus''. Previous estimates suggested the divergence took place during the Jurassic or Triassic. More recent estimates suggest that they have an origin during the Late Jurassic, with the first fossil records in the Early Cretaceous. About 3,106 species are currently described, with a few hundred more left to be described. Although these insects are often called "white ants", they are not ants, and are not closely related to ants. Like ants and some bees a ...
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Blattodea
Blattodea is an order of insects that contains cockroaches and termites. Formerly, termites were considered a separate order, Isoptera, but genetic and molecular evidence suggests they evolved from within the cockroach lineage, cladistically making them cockroaches as well. The Blattodea and the mantis (order Mantodea) are now all considered part of the superorder Dictyoptera. Blattodea includes approximately 4,400 species of cockroach in almost 500 genera, and about 3,000 species of termite in around 300 genera. Termites are pale-coloured, soft-bodied eusocial insects that live in colonies, whereas cockroaches are darker-coloured (often brown), sclerotized, segmented insects. Within the colony, termites have a caste system, with a pair of mature reproductives, the king and the queen, and numerous sterile workers and soldiers. Cockroaches are not colonial but do have a tendency to aggregate and may be considered pre-social, as all adults are capable of breeding. Other simil ...
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Cryptocercus
''Cryptocercus'' is a genus of Dictyoptera (cockroaches and allies) and the sole member of its own family Cryptocercidae. Species are known as wood roaches or brown-hooded cockroaches. These roaches are subsocial, their young requiring considerable parental interaction. They also share wood-digesting gut bacteria types with wood-eating termites, and are therefore seen as evidence of a close genetic relationship, that termites are essentially evolved from social cockroaches. ''Cryptocercus'' is especially notable for sharing numerous characteristics with termites, and phylogenetic studies have shown this genus is more closely related to termites than it is to other cockroaches.Djernæs, M., et al. 2012Phylogeny of cockroaches (Insecta, Dictyoptera, Blattodea), with placement of aberrant taxa and exploration of out-group sampling.''Systematic Entomology'' 37(1): 65–83. doi: 10.1111/j.1365-3113.2011.00598.x. Species Found in North America and (especially temperate) Asia, there ar ...
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Tryonicidae
The Tryonicidae are a family of cockroaches. Biodiversity and distribution Two genera containing 17 species are currently confirmed as belonging to this family. Table 1: Number of species of Tryonicidae in each region in which it is present (A=adventive, E=endemic, I=indigenous) Notes * Beccaloni & Eggleton's (2011) figures of '10 genera, 47 species' presumably does not take into account Murienne's (2009) publication (they do not cite it) * According to Murienne (2009: 49), the tribe Methanini certainly belongs to the Blattidae: Polyzosteriinae, as probably does the group of New Caledonian endemic genera '' Angustonicus'', '' Pallidionicus'', '' Pellucidonicus'', '' Punctulonicus'', and '' Rothisilpha'' * A report has been published of an unidentified endemic "tryonicine" from New Zealand, in addition to the adventive ''Tryonicus parvus The Tryonicidae are a family of cockroaches. Biodiversity and distribution Two genera containing 17 species are currently confirmed ...
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Roachoid
"Roachoids", also known as "Roachids" or "Blattoids" are members of the stem group of Dictyoptera (the group containing modern cockroaches, termites and praying mantises). They generally resemble cockroaches, but most members, unlike modern dictyopterans, have generally long external ovipositors, and are thought not to have laid ootheca like modern dictyopterans. Systematic position Cockroaches are popularly thought to be an ancient order of insects, with their origins in the Carboniferous. However, since the middle of the 20th century it has been known that the primitive cockroach insects found fossilized in Palaeozoic strata are the forerunners not only of modern cockroaches and termites but also of mantises.Grimaldi, D (1997): A fossil mantis (Insecta: Mantoidea) in Cretaceous amber of New Jersey, with comments on early history of Dictyoptera. ''American Museum Novitates'' 3204: 1–11 The origin of these groups from a blattopteran stock are now generally thought to be in ...
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Alienopteridae
Alienopteridae is an extinct family of dictyopterans, known from the Mid-Cretaceous to Eocene. They are noted for their unusual combination of features not found in other dictyopterans. Taxonomy It was originally assigned to its own order Alienoptera by Bai et al., 2016. It was reassigned to the dictyopteran superfamily Umenocoleoidea as sister family to the beetle-like Umenocoleidae by Vršanský ''et al.'' (2018), and a more recent analysis similarly places Alienopteridae and Umenocoleidae as sister taxa within Dictyoptera, but placing both lineages outside of Blattodea. A 2021 study revived the order Alienoptera for the clade containing Alienopteridae and Umenocoleidae, with a cladistic analysis placing Alienoptera as the sister clade to Mantodea (praying mantises). Distribution The majority of the alienopterid genera are known from the mid Cretaceous (latest Albian-earliest Cenomanian ~ 100 million years ago) Burmese amber found in Myanmar; though an additional two genera ...
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Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian line ...
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Umenocoleidae
Umenocoleidae is an extinct family of dictyopteran insects known from the Cretaceous. They are considered to be closely related to the Alienopteridae. They were originally considered to be beetles due to their beetle-like morphology, with sclerotised elytra-like forewings. This was probably an adaptation for living under bark and in other tight spaces. Systematics After * †'' Umenocoleus'' Chen and Tan 1973 Dalazi Formation, China, Early Cretaceous (Aptian) Zhonggou Formation, China, Early Cretaceous (Albian) * †'' Ponopterix'' Vršanský and Grimaldi 1999 Crato Formation, Brazil, Early Cretaceous (Aptian) * †'' Blattapterix'' Vršanský 2003 Xiagou Formation, China, Early Cretaceous (Aptian) * †'' Elytropterix'' Vršanský 2003 Dzun-Bain Formation, Mongolia, Aptian * †'' Petropterix'' Vršanský 2003 Dzun-Bain Formation, Mongolia, Zaza Formation, Russia, Kitadani Formation, Japan, Early Cretaceous (Aptian) Zhonggou Formation, China, Early Cretaceous (Albian) * ...
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Blaberidae
Giant cockroaches, or blaberids (family Blaberidae) are the second-largest cockroach family by number of species. Notable species Notable species within this family include: * Cape mountain cockroach – ''Aptera fusca'' * Dwarf cave cockroach – ''Blaberus atropos=Blaberus fusca'' * Death's head cockroach – ''Blaberus craniifer'' * False death's head cockroach – ''Blaberus discoidalis'' * Dubia cockroach – ''Blaptica dubia'' * Green banana cockroach – ''Panchlora nivea'' * Madagascar hissing cockroach – ''Gromphadorhina portentosa'' * Speckled cockroach – ''Nauphoeta cinerea'' * Surinam cockroach – ''Pycnoscelus surinamensis'' * Giant burrowing cockroach – ''Macropanesthia rhinoceros'' Genera *Subfamily Attaphilinae **'' Attaphila'' *Subfamily Blaberinae **'' Achroblatta'' **'' Anchoblatta'' **'' Antioquita'' **'' Archimandrita'' **'' Aspiduchus'' **'' Cacoblatta'' **'' Capucinella'' **'' Cariacasia'' **'' Bionoblatta'' **'' Blaberus'' **'' Blaptic ...
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