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Zuul
''Zuul'' is a genus of herbivorous ankylosaurine dinosaur from the Campanian Judith River Formation of Montana. The type species is ''Zuul crurivastator''. It is known from a complete skull and tail, which represents the first ankylosaurin known from a complete skull and tail club, as well as the most complete ankylosaurid specimen thus far recovered from North America. The specimen also preserved ''in situ'' osteoderms, keratin, and skin remains. Discovery and naming In 2014, Theropoda Expeditions LLC was excavating a possible ''Gorgosaurus'' specimen near Havre, Montana. On 16 May, a skid-steer loader removing a -high overburden unexpectedly hit upon an ankylosaurian tail club. An almost complete ankylosaur skeleton proved to be present. As it had not been eroded on the surface, it was in pristine condition. It was found overturned, with the stomach facing upwards. The skeleton was largely articulated, but the skull and some neck vertebrae were separated from the main tors ...
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Zuul Locality
''Zuul'' is a genus of herbivorous Ankylosaurinae, ankylosaurine dinosaur from the Campanian Judith River Formation of Montana. The type species is ''Zuul crurivastator''. It is known from a complete skull and tail, which represents the first Ankylosaurini, ankylosaurin known from a complete skull and tail club, as well as the most complete Ankylosauridae, ankylosaurid specimen thus far recovered from North America. The specimen also preserved ''in situ'' osteoderms, keratin, and skin remains. Discovery and naming In 2014, Theropoda Expeditions LLC was excavating a possible ''Gorgosaurus'' specimen near Havre, Montana. On 16 May, a skid-steer loader removing a -high overburden unexpectedly hit upon an ankylosaurian tail club. An almost complete ankylosaur skeleton proved to be present. As it had not been eroded on the surface, it was in pristine condition. It was found overturned, with the stomach facing upwards. The skeleton was largely articulated, but the skull and some neck ...
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Zuul
''Zuul'' is a genus of herbivorous ankylosaurine dinosaur from the Campanian Judith River Formation of Montana. The type species is ''Zuul crurivastator''. It is known from a complete skull and tail, which represents the first ankylosaurin known from a complete skull and tail club, as well as the most complete ankylosaurid specimen thus far recovered from North America. The specimen also preserved ''in situ'' osteoderms, keratin, and skin remains. Discovery and naming In 2014, Theropoda Expeditions LLC was excavating a possible ''Gorgosaurus'' specimen near Havre, Montana. On 16 May, a skid-steer loader removing a -high overburden unexpectedly hit upon an ankylosaurian tail club. An almost complete ankylosaur skeleton proved to be present. As it had not been eroded on the surface, it was in pristine condition. It was found overturned, with the stomach facing upwards. The skeleton was largely articulated, but the skull and some neck vertebrae were separated from the main tors ...
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Ghostbusters
''Ghostbusters'' is a 1984 American supernatural comedy film directed and produced by Ivan Reitman, and written by Dan Aykroyd and Harold Ramis. It stars Bill Murray, Aykroyd, and Ramis as Peter Venkman, Ray Stantz, and Egon Spengler, three eccentric parapsychologists who start a ghost-catching business in New York City. It also stars Sigourney Weaver and Rick Moranis, and features Annie Potts, William Atherton, and Ernie Hudson in supporting roles. Based on his own fascination with spirituality, Aykroyd conceived ''Ghostbusters'' as a project starring himself and John Belushi, in which they would venture through time and space battling supernatural threats. Following Belushi's death in 1982, and with Aykroyd's concept deemed financially impractical, Ramis was hired to help rewrite the script to set it in New York City and make it more realistic. It was the first comedy film to employ expensive special effects, and Columbia Pictures, concerned about its relatively hig ...
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Victoria Arbour
Victoria Megan Arbour is a Canadian evolutionary biologist and vertebrate palaeontologist at Royal BC Museum, where she is Curator of Palaeontology. An "expert on the armoured dinosaurs known as ankylosaurs", Arbour analyzes fossils and creates 3-D computer models. She named the possible pterosaur ''Gwawinapterus'' from Hornby Island, and a partial ornithischian dinosaur from Sustut Basin, British Columbia (now named '' Ferrisaurus''), and has participated in the naming of the ankylosaurs '' Zuul'', '' Zaraapelta'', ''Crichtonpelta'', and ''Ziapelta''. Early life and education Born in 1983, Arbour is from Halifax, Nova Scotia. Her mother, a math teacher, and father, a soil scientist, supported her science interests. Arbour completed a B.Sc. Honours Thesis supervised by Milton Graves, ''An ornithischian dinosaur from the Sustut Basin, British Columbia, Canada'', and graduated from Dalhousie University in 2006. She completed her master's thesis, ''Evolution, biomechanics, and f ...
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Judith River Formation
The Judith River Formation is a fossil-bearing geologic formation in Montana, and is part of the Judith River Group. It dates to the Late Cretaceous, between 79 and 75.3 million years ago, corresponding to the "Judithian" land vertebrate age. It was laid down during the same time period as portions of the Two Medicine Formation of MontanaSullivan, R.M. and Lucas, S. G. (2006). "The Kirtlandian land-vertebrate "age"–faunal composition, temporal position and biostratigraphic correlation in the nonmarine Upper Cretaceous of western North America." Pp. 7-29 in Lucas, S. G. and Sullivan, R.M. (eds.), ''Late Cretaceous vertebrates from the Western Interior. New Mexico Museum of Natural History and Science Bulletin 35''. and the Oldman Formation of Alberta. It is an historically important formation, explored by early American paleontologists such as Edward Drinker Cope, who named several dinosaurs from scrappy remains found here on his 1876 expedition (such as '' Monoclonius''). Mod ...
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Ankylosauridae
Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Paleogene extinction event. These animals were mainly herbivorous and were obligate quadrupeds, with leaf-shaped teeth and robust, scute-covered bodies. Ankylosaurids possess a distinctly domed and short snout, wedge-shaped osteoderms on their skull, scutes along their torso, and a tail club. Ankylosauridae is exclusively known from the northern hemisphere, with specimens found in western North America, Europe, and East Asia. The first discoveries within this family were of the genus ''Ankylosaurus'', by Peter Kaiser and Barnum Brown in Montana in 1906. Brown went on to name Ankylosauridae and the subfamily Ankylosaurinae in 1908. Anatomy Ankylosaurids are stout, solidly built, armoured dinosaurs. They possess accessory ossifications on ...
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Royal Ontario Museum
The Royal Ontario Museum (ROM) is a museum of art, world culture and natural history in Toronto, Ontario, Canada. It is one of the largest museums in North America and the largest in Canada. It attracts more than one million visitors every year, making the ROM the most-visited museum in Canada. The museum is north of Queen's Park, in the University of Toronto district, with its main entrance on Bloor Street West. Museum subway station is named after the ROM and, since a 2008 renovation, is decorated to resemble the institution's collection at the platform level. Established on April 16, 1912, and opened on March 19, 1914, the museum has maintained close relations with the University of Toronto throughout its history, often sharing expertise and resources. The museum was under the direct control and management of the University of Toronto until 1968, when it became an independent Crown agency of the Government of Ontario. Today, the museum is Canada's largest field-research ...
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Ankylosaurinae
Ankylosaurinae is a subfamily of ankylosaurid dinosaurs, existing from the Early Cretaceous about 105 million years ago until the end of the Late Cretaceous, about 66 mya. Many genera are included in the clade, such as ''Ankylosaurus'', ''Pinacosaurus'', '' Euoplocephalus'', and ''Saichania''. Features Ankylosaurines are defined as being closer relatives to ''Ankylosaurus'' than to ''Shamosaurus''. Diagnostic features of ankylosaurines include the nuchal shelf that obscures the occiput in dorsal view, and the quadrate condyle which is obscured lightly by the quadratojugal boss. Phylogeny The following cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... is based on the 50% majority rule phylogenetic analysis of Arbour & Currie (2015): References Ankyl ...
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Scolosaurus
''Scolosaurus'' is an extinct genus of ankylosaurid dinosaurs within the subfamily Ankylosaurinae. It is known from the lower levels of the Dinosaur Park Formation and upper levels of the Oldman Formation in the Late Cretaceous (latest middle Campanian stage, about 76.5 Ma ago) of Alberta, Canada. It contains two species, ''S. cutleri'' and ''S. thronus''. The type species, ''S. cutleri'', measured up to in length and in body mass. Discovery ''Scolosaurus'' was named by Franz Nopcsa von Felső-Szilvás in 1928 in paleontology, 1928, based on holotype Natural History Museum, London, NHMUK R.5161, a nearly complete specimen that preserves the entire skeleton except for the distal end of the tail, the right forelimb, the right hindlimb, and the skull. The rare preservation of osteoderms and skin impression are also present. The fossil skeleton was discovered by William Edmund Cutler, an independent fossil collector in 1914 at Quarry 80 of the Deadlodge Canyon locality. It was ...
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Ankylosaurini
Ankylosaurinae is a subfamily of ankylosaurid dinosaurs, existing from the Early Cretaceous about 105 million years ago until the end of the Late Cretaceous, about 66 mya. Many genera are included in the clade, such as ''Ankylosaurus'', ''Pinacosaurus'', ''Euoplocephalus'', and ''Saichania''. Features Ankylosaurines are defined as being closer relatives to ''Ankylosaurus'' than to ''Shamosaurus''. Diagnostic features of ankylosaurines include the nuchal shelf that obscures the occiput in dorsal view, and the quadrate condyle which is obscured lightly by the quadratojugal boss. Phylogeny The following cladogram is based on the 50% majority rule phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... of Arbour & Currie (2015): References Ankylosa ...
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Keratin
Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. Alpha-keratin (α-keratin) is a type of keratin found in vertebrates. It is the key structural material making up scales, hair, nails, feathers, horns, claws, hooves, and the outer layer of skin among vertebrates. Keratin also protects epithelial cells from damage or stress. Keratin is extremely insoluble in water and organic solvents. Keratin monomers assemble into bundles to form intermediate filaments, which are tough and form strong unmineralized epidermal appendages found in reptiles, birds, amphibians, and mammals. Excessive keratinization participate in fortification of certain tissues such as in horns of cattle and rhinos, and armadillos' osteoderm. The only other biological matter known to approximate the toughness of keratinized tissue is chitin. Keratin comes in two types, the primitive, softer forms found in all vertebrates and harder, derived forms found only among saur ...
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Campanian
The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian spans the time from 83.6 (± 0.2) to 72.1 (± 0.2) million years ago. It is preceded by the Santonian and it is followed by the Maastrichtian. The Campanian was an age when a worldwide sea level rise covered many coastal areas. The morphology of some of these areas has been preserved: it is an unconformity beneath a cover of marine sedimentary rocks. Etymology The Campanian was introduced in scientific literature by Henri Coquand in 1857. It is named after the French village of Champagne in the department of Charente-Maritime. The original type locality was a series of outcrop near the village of Aubeterre-sur-Dronne in the same region. Definition The base of the Campanian Stage is defined as a place in the stratigraphic colu ...
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