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Vladlenosaurus
''Vladlenosaurus'' is an extinct genus of capitosaur from Russia. It lived during the late Vetlugian (Early Triassic). Based on the type of deposits it was found in, ''Vladlenosaurus'' probably inhabited lacustrine, or lake, habitats. The type species is ''V. alexeyevi'', named in 2000. Description Unlike the flatter, more rounded snouts of other capitosaurs, ''Vladlenosaurus'' had a wedge-shaped snout. This characteristic is also seen in trematosaurians, although it was independently acquired in both cases as a result of convergent evolution. Classification ''Vladlenosaurus'' is similar in appearance to the benthosuchids, a group of trematosauroid temnospondyls, but is more closely related to mastodonsauroids. Among capitosaurs, it shares many features with the basal form ''Wetlugasaurus'', also from the Early Triassic of Russia. ''Vladlenosaurus alexeyevi'' was even considered to be a species of ''Wetlugasaurus'' in a 2006 study. In a 2011 phylogenetic analysis, the first ...
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Capitosauria
Capitosauria is an extinct group of large temnospondyl amphibians with simplified stereospondyl vertebrae. Mainly living as piscivores in lakes and rivers, the Capitosauria and its sister taxon Trematosauria were the only major labyrinthodonts that existed during the Mesozoic in ecological niches broadly similar to those of modern crocodiles, and some grew to very large sizes. At 6 meters in length, the Mid-Triassic '' Mastodonsaurus giganteus'' is not only thought to have been the largest capitosaur, but possibly also the largest amphibian to have lived. The latest known remains are from the Rhaetian of Germany and are referred to ''Cyclotosaurus''. Capitosauria was first named by Schoch and Milner (2000) and further described by Yates and Warren (2000), who assigned ''Lydekkerina'' and Mastodonsauroidea to it. It was described by Damiani (2001) under the name Mastodonsauroidea. In their phylogenetic analysis of temnospondyls, Ruta ''et al.'' (2007) placed ''Lydekkerina'' and it ...
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Benthosuchus
''Benthosuchus'' (meaning "deep water crocodile") is an extinct genus of temnospondyl amphibian from the Early Triassic of Russia. It was primarily aquatic, living in rivers and lakes. Multiple species are known, with the largest reaching about 2.5 meters in length. Russian paleontologist Ivan Yefremov fremovcalled the genus ''Benthosaurus'' "deep water lizard" (from Ancient Greek βένθος (''benthos'') "depth, deep water") in his original 1929 description, "in view of its clearly indicated adaptation to life in deep water" shown by "the position of the orbits and the flatness of the skull." The type species ''B. sushkini'' honored his late teacher Petr Sushkin. The generic name was preoccupied by ''Benthosaurus'' Goode & Bean, 1886, a fish, and he renamed the genus ''Benthosuchus'' ("deep water crocodile") in 1937.Efremov, I. A. (1929). ''Benthosaurus sushkini'', ein neuer Labyrinthodont der permotriassischen Ablagerungen der Sharschenga Flusses. ''Bulletin of the Acad ...
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Edingerella
''Edingerella'' is an extinct genus of temnospondyl amphibian from the Early Triassic of Madagascar. It is a basal capitosaur closely related to '' Watsonisuchus''. Phylogeny Below is a cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... from Fortuny ''et al.'' (2011): References Triassic temnospondyls of Africa Prehistoric animals of Madagascar Capitosaurs Early Triassic amphibians of Africa Prehistoric amphibian genera {{triassic-animal-stub ...
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Odenwaldia
''Odenwaldia'' is an extinct genus of mastodonsauroid temnospondyl within the family Heylerosauridae. History of study ''Odenwaldia'' is only known from one species, ''O. heidelbergensis'', and was named by Morales & Kamphausen (1984). The holotype, a skull roof and counterpiece cast, were collected from the Middle Bundsandstein (Oberes Konglomerat) near Heidelberg, Germany and were first described by Wilhelm Simon in 1961, who thought that the specimen belonged to the trematosaur ''Trematosaurus''. It was then redescribed by Schoch (2008). Description The holotype is the only uncontroversial specimen of this taxon, although others have been referred to the species. ''Odenwaldia'' is diagnosed by several autapomorphies, including (1) small orbits combined with broad interorbital distance; (2) preorbital region slender, with nasals and lacrimals narrower than frontals; and (3) dermal ornament consists of small, similarly sized polygons, but no elongated ridges. In contrast t ...
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Trematosauroidea
Trematosauroidea are an important group of Triassic temnospondyl amphibians. They flourished briefly during the Early Triassic, occurring worldwide before declining at the start of the Middle Triassic, although the group continued until the Late Triassic. They were medium-sized temnospondyls with wedge-shaped tails, narrow skulls, and, in advanced forms, elongated snouts. The latter feature was probably an adaptation for feeding on fish. The largest and most specialized family, the Trematosauridae, are the only batrachomorphs to have adapted to a marine lifestyle with the exception of the modern crab-eating frog. A temnospondyl ilium was described in 2004 from the Callovian Toutunhe Formation in the Junggar Basin of China. Although the isolated bone was impossible to identify on the species level, it was referred to Trematosauroidea. The presence of this bone in the Toutunhe Formation extends the range of trematosauroids into the Middle Jurassic, making it one of only three gro ...
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Cherninia
''Cherninia'' is an extinct genus of mastodonsaurid temnospondyl. The type species, ''Cherninia denwai'', is known from the Denwa Formation of India. It is based on a massive skull, ISI A 54, which was originally considered a species of ''Parotosuchus'' in 1998 before being given its own genus in 2001. Another species, ''Cherninia megarhina'', is known from the Upper Ntawere Formation of Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most central point. Its neighbours are t .... ''C. megarhina'' is based on another large skull, BP/1/4223, which had also been previously referred to ''Parotosuchus''. Though not as well-preserved as the skull of ''C. denwai'', BP/1/4233 was described earlier in 1974. It was described by Sharon Chernin, a paleontologist at the Bernard Price Institute and the namesake of the genus. ...
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Xenotosuchus
''Xenotosuchus'' is an extinct genus of mastodonsaurid temnospondyl within the family Mastodonsauridae known from the Triassic of South Africa. The genus is based on a skull originally described as '' Parotosuchus'', an animal which it resembled in general build and habit. Description Like many mastodontosaurids, it was a large animal with a large head. Its amphibian life history meant that the distinct shape of the skull roof would change from a generalized tadpole-like skull type through to the distinct adult shape. The head bones are covered in large pits and grooves, indicating extensive dermal armour on the head. Both the upper and lower jaw had tusks, those of the upper jaw being situated on a 2nd row of teeth on the vomer and palatine bone. Contrary to related forms like ''Mastodonsaurus'', the tusks of the lower jaw were of moderate size and did not penetrate the premaxilla. See also * Prehistoric amphibian * List of prehistoric amphibians This list of prehistoric ...
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Watsonisuchus
''Watsonisuchus'' is an extinct genus of temnospondyl amphibian from the Early Triassic of Australia, Madagascar, and South Africa. It was up to 122 cm long and had a robust skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ... of 24 cm in length.http://www.angellis.net/Web/PDfiles/amphs.pdf References * ''Dragons in the Dust: The Paleobiology of the Giant Monitor Lizard Megalania'' by Ralph E. Molnar (Pg. 39) Triassic temnospondyls of Africa Prehistoric amphibians of Australia Capitosaurs Triassic temnospondyls of Australia Early Triassic amphibians of Africa Fossil taxa described in 1966 Prehistoric amphibian genera {{Triassic-animal-stub ...
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Rhinesuchidae
Rhinesuchidae is a family of tetrapods that lived primarily in the Permian period. They belonged to the broad group Temnospondyli, a successful and diverse collection of semiaquatic tetrapods which modern amphibians are probably descended from. Rhinesuchids can be differentiated from other temnospondyls by details of their skulls, most notably the interior structure of their otic notches at the back of the skull. They were among the earliest-diverging members of the Stereospondyli, a subgroup of temnospondyls with flat heads and aquatic habits. Although more advanced stereospondyls evolved to reach worldwide distribution in the Triassic period, rhinesuchids primarily lived in the high-latitude environments of Gondwana (what is now South America and Africa) during the Guadalupian and Lopingian epochs of the Permian. The taxonomy of this family has been convoluted, with more than twenty species having been named in the past; a 2017 review recognized only eight of them (distributed a ...
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Lydekkerina
''Lydekkerina'' is an extinct genus of stereospondyl temnospondyl. It is the type genus of the family Lydekkerinidae. Fossils have been collected from Early Triassic deposits in South Africa and Australia. The type species is '' L. huxleyi'', first described in 1889. While most other stereospondyls were semiaquatic, ''Lydekkerina'' was exclusively terrestrial. Description ''Lydekkerina'' was a relatively small temnospondyl, growing up to around in length. Skulls range in length from in the smallest known individual to up to in larger individuals. The skull is wedge-shaped and has a parabolic outline with convex lateral margins. Shallow pits cover the surface of the skull. Teeth line the palate as well as the jaws, and some skulls even bear large ectopterygoid tusks on the underside of the skull. ''Lydekkerina'' can be distinguished from other lydekkerinids on the basis of several skull characteristics. One such feature is the presence of vomerine shagreen, tiny bumps cove ...
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Stereospondyl
The Stereospondyli are a group of extinct temnospondyl amphibians that existed primarily during the Mesozoic period. They are known from all seven continents and were common components of many Triassic ecosystems, likely filling a similar ecological niche to modern crocodilians prior to the diversification of pseudosuchian archosaurs. Classification and anatomy The group was first defined by Zittel (1888) on the recognition of the distinctive vertebral anatomy of the best known stereospondyls of the time, such as ''Mastodonsaurus'' and ''Metoposaurus''. The term 'stereospondylous' as a descriptor of vertebral anatomy was coined the following year by Fraas, referring to a vertebral position consisting largely or entirely of the intercentrum in addition to the neural arch. While the name 'Stereospondyli' is derived from the stereospondylous vertebral condition, there is a diversity of vertebral morphologies among stereospondyls, including the diplospondylous (' tupilakosaurid') co ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about ance ...
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