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Benthosuchus
''Benthosuchus'' (meaning "deep water crocodile") is an extinct genus of temnospondyl amphibian from the Early Triassic of Russia. It was primarily aquatic, living in rivers and lakes. Multiple species are known, with the largest reaching about 2.5 meters in length. Russian paleontologist Ivan Yefremov fremovcalled the genus ''Benthosaurus'' "deep water lizard" (from Ancient Greek βένθος (''benthos'') "depth, deep water") in his original 1929 description, "in view of its clearly indicated adaptation to life in deep water" shown by "the position of the orbits and the flatness of the skull." The type species ''B. sushkini'' honored his late teacher Petr Sushkin. The generic name was preoccupied by ''Benthosaurus'' Goode & Bean, 1886, a fish, and he renamed the genus ''Benthosuchus'' ("deep water crocodile") in 1937.Efremov, I. A. (1929). ''Benthosaurus sushkini'', ein neuer Labyrinthodont der permotriassischen Ablagerungen der Sharschenga Flusses. ''Bulletin of the Acad ...
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Benthosuchus2DB2small
''Benthosuchus'' (meaning "deep water crocodile") is an extinct genus of temnospondyli, temnospondyl amphibian from the Early Triassic of Russia. It was primarily aquatic, living in rivers and lakes. Multiple species are known, with the largest reaching about 2.5 meters in length. Russian paleontologist Ivan Yefremov [Efremov] called the genus ''Benthosaurus'' "deep water lizard" (from Ancient Greek βένθος (''benthos'') "depth, deep water") in his original 1929 description, "in view of its clearly indicated adaptation to life in deep water" shown by "the position of the orbits and the flatness of the skull." The type species ''B. sushkini'' honored his late teacher Petr Sushkin. The generic name was preoccupied by ''Benthosaurus'' Goode & Bean, 1886, a fish, and he renamed the genus ''Benthosuchus'' ("deep water crocodile") in 1937.Efremov, I. A. (1929). ''Benthosaurus sushkini'', ein neuer Labyrinthodont der permotriassischen Ablagerungen der Sharschenga Flusses. ''Bull ...
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Benthosuchus Korobkovi
''Benthosuchus'' (meaning "deep water crocodile") is an extinct genus of temnospondyl amphibian from the Early Triassic of Russia. It was primarily aquatic, living in rivers and lakes. Multiple species are known, with the largest reaching about 2.5 meters in length. Russian paleontologist Ivan Yefremov fremovcalled the genus ''Benthosaurus'' "deep water lizard" (from Ancient Greek βένθος (''benthos'') "depth, deep water") in his original 1929 description, "in view of its clearly indicated adaptation to life in deep water" shown by "the position of the orbits and the flatness of the skull." The type species ''B. sushkini'' honored his late teacher Petr Sushkin. The generic name was preoccupied by ''Benthosaurus'' Goode & Bean, 1886, a fish, and he renamed the genus ''Benthosuchus'' ("deep water crocodile") in 1937.Efremov, I. A. (1929). ''Benthosaurus sushkini'', ein neuer Labyrinthodont der permotriassischen Ablagerungen der Sharschenga Flusses. ''Bulletin of the Academ ...
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Rhinesuchidae
Rhinesuchidae is a family of tetrapods that lived primarily in the Permian period. They belonged to the broad group Temnospondyli, a successful and diverse collection of semiaquatic tetrapods which modern amphibians are probably descended from. Rhinesuchids can be differentiated from other temnospondyls by details of their skulls, most notably the interior structure of their otic notches at the back of the skull. They were among the earliest-diverging members of the Stereospondyli, a subgroup of temnospondyls with flat heads and aquatic habits. Although more advanced stereospondyls evolved to reach worldwide distribution in the Triassic period, rhinesuchids primarily lived in the high-latitude environments of Gondwana (what is now South America and Africa) during the Guadalupian and Lopingian epochs of the Permian. The taxonomy of this family has been convoluted, with more than twenty species having been named in the past; a 2017 review recognized only eight of them (distributed a ...
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Temnospondyli
Temnospondyli (from Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian, and Triassic periods. A few species continued into the Jurassic and Cretaceous periods. Fossils have been found on every continent. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are considered amphibians, many had characteristics, such as scales and armour-like bon ...
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Trematosauroidea
Trematosauroidea are an important group of Triassic temnospondyl amphibians. They flourished briefly during the Early Triassic, occurring worldwide before declining at the start of the Middle Triassic, although the group continued until the Late Triassic. They were medium-sized temnospondyls with wedge-shaped tails, narrow skulls, and, in advanced forms, elongated snouts. The latter feature was probably an adaptation for feeding on fish. The largest and most specialized family, the Trematosauridae, are the only batrachomorphs to have adapted to a marine lifestyle with the exception of the modern crab-eating frog. A temnospondyl ilium was described in 2004 from the Callovian Toutunhe Formation in the Junggar Basin of China. Although the isolated bone was impossible to identify on the species level, it was referred to Trematosauroidea. The presence of this bone in the Toutunhe Formation extends the range of trematosauroids into the Middle Jurassic, making it one of only three gro ...
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Stereospondyli
The Stereospondyli are a group of extinct temnospondyl amphibians that existed primarily during the Mesozoic period. They are known from all seven continents and were common components of many Triassic ecosystems, likely filling a similar ecological niche to modern crocodilians prior to the diversification of pseudosuchian archosaurs. Classification and anatomy The group was first defined by Zittel (1888) on the recognition of the distinctive vertebral anatomy of the best known stereospondyls of the time, such as ''Mastodonsaurus'' and '' Metoposaurus''. The term 'stereospondylous' as a descriptor of vertebral anatomy was coined the following year by Fraas, referring to a vertebral position consisting largely or entirely of the intercentrum in addition to the neural arch. While the name 'Stereospondyli' is derived from the stereospondylous vertebral condition, there is a diversity of vertebral morphologies among stereospondyls, including the diplospondylous (' tupilakosaurid') c ...
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Thoosuchus
''Thoosuchus'' (meaning "active crocodile") is an extinct genus of basal trematosauroid trematosaurian temnospondyl. Fossils have been found from Russia and date back to the Early Triassic. It is the type genus of the family Thoosuchidae, formerly called the subfamily Thoosuchinae and placed within Benthosuchidae. Efremov, I. A. (1940). Preliminary description of the new Permian and Triassic Tetrapoda from U.S.S.R. ''Trudy Paleontologicheskogo Instituta'' 10:1–140. The benthosuchids were originally composed of the majority of basal trematosaurian forms regarded as the ancestors of the trematosaurids. Although the genus was first named in 1940, material from one species, ''E. yakovlevi'', was originally tentatively referred to ''Trematosuchus'' in 1926. For its family, ''Thoosuchus'' was fairly small, reaching a little over 60 cm with a 15 cm skull. ''Thoosuchus'' superficially resembles the more derived trematosaurids, but can be distinguished from them on the ...
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Capitosauria
Capitosauria is an extinct group of large temnospondyl amphibians with simplified stereospondyl vertebrae. Mainly living as piscivores in lakes and rivers, the Capitosauria and its sister taxon Trematosauria were the only major labyrinthodonts that existed during the Mesozoic in ecological niches broadly similar to those of modern crocodiles, and some grew to very large sizes. At 6 meters in length, the Mid-Triassic '' Mastodonsaurus giganteus'' is not only thought to have been the largest capitosaur, but possibly also the largest amphibian to have lived. The latest known remains are from the Rhaetian of Germany and are referred to ''Cyclotosaurus''. Capitosauria was first named by Schoch and Milner (2000) and further described by Yates and Warren (2000), who assigned ''Lydekkerina'' and Mastodonsauroidea to it. It was described by Damiani (2001) under the name Mastodonsauroidea. In their phylogenetic analysis of temnospondyls, Ruta ''et al.'' (2007) placed ''Lydekkerina'' and it ...
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Early Triassic
The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a unit in chronostratigraphy. The Early Triassic is the oldest epoch of the Mesozoic Era. It is preceded by the Lopingian Epoch (late Permian, Paleozoic Era) and followed by the Middle Triassic Epoch. The Early Triassic is divided into the Induan and Olenekian ages. The Induan is subdivided into the Griesbachian and Dienerian subages and the Olenekian is subdivided into the Smithian and Spathian subages. The Lower Triassic series is coeval with the Scythian Stage, which is today not included in the official timescales but can be found in older literature. In Europe, most of the Lower Triassic is composed of Buntsandstein, a lithostratigraphic unit of continental red beds. The Early Triassic and partly also the Middle Triassic span the in ...
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Capitosaurid
Mastodonsauridae is a family of capitosauroid temnospondyls. Fossils belonging to this family have been found in North America, Greenland, Europe, Asia, and Australia. The family Capitosauridae is synonymous with Mastodonsauridae. Description Size Mastodonsaurids were generally large amphibians, with some length estimates ranging from 3 to 6 meters. Some genera, like ''Mastodonsaurus'' and ''Cyclotosaurus'' were specially large, reaching at least 4 meters or more. Other genera like ''Parotosuchus'' and ''Paracyclotosaurus'' only reached 2 meters or more in length, but it's still pretty large compared to modern-day amphibians. Distinguishing features Some mastodonsaurids including ''Parotosuchus'' were covered in a scaly skin, unlike the smooth skin of modern-day amphibians, and probably moved with an eel-like motion in the water. Another peculiar mastodonsaurid is ''Sclerothorax'', with unusual features including its elongated neural spines. The neural spines are tallest at the ...
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Wetlugasaurus
''Wetlugasaurus'' (meaning "Vetluga River lizard") is an extinct genus of Temnospondyli, temnospondyl amphibian from the Early Triassic (Olenekian) Charkabozh Formation, Charkabozh, Kzylsaiskaya Formation, Kzylsaiskaya, Petropavlovka Formation, Petropavlovka, Kamennyi Yar Formation, Kamennyi Yar and Vetluga Series Formations of northern Russia and Greenland. It had a long skull, and reached a total length of . References Further reading

* ''The Age of Dinosaurs in Russia and Mongolia'' by Michael J. Benton, Mikhail A. Shishkin, David M. Unwin, and Evgenii N. Kurochkin. p. 35-59. * ''Bibliography Of Fossil Vertebrates 1934-1938'' by C. I. Camp Capitosaurs Prehistoric amphibian genera Triassic temnospondyls of Europe Olenekian life Fossils of Russia Fossil taxa described in 1930 {{triassic-animal-stub ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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