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Varanid
The Varanidae are a family of lizards in the superfamily Varanoidea within the Anguimorpha group. The family, a group of carnivorous and frugivorous lizards, includes the living genus '' Varanus'' and a number of extinct genera more closely related to ''Varanus'' than to the earless monitor lizard (''Lanthanotus''). ''Varanus'' includes the Komodo dragon (the largest living lizard), crocodile monitor, savannah monitor, the goannas of Australia and Southeast Asia, and various other species with a similarly distinctive appearance. Their closest living relatives are the earless monitor lizard and chinese crocodile lizard. The oldest members of the family are known from the Late Cretaceous of Mongolia. Taxonomy The Varanidae were defined (using morphological characteristics) by Estes, de Queiroz and Gauthier (1988) as the clade containing the most recent common ancestor of ''Lanthanotus'' and ''Varanus'' and all of its descendants. A similar definition was formulated by Conrad ''et al ...
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Komodo Dragon
The Komodo dragon (''Varanus komodoensis''), also known as the Komodo monitor, is a member of the monitor lizard family Varanidae that is endemic to the Indonesian islands of Komodo, Rinca, Flores, and Gili Motang. It is the largest extant species of lizard, growing to a maximum length of , and weighing up to . As a result of their size, Komodo dragons are apex predators, and dominate the ecosystems in which they live. Komodo dragons hunt and ambush prey including invertebrates, birds, and mammals. It has been claimed that they have a venomous bite; there are two glands in the lower jaw that secrete several toxic proteins. The biological significance of these proteins is disputed, but the glands have been shown to secrete an anticoagulant. Komodo dragons' group behavior in hunting is exceptional in the reptile world. The diet of Komodo dragons mainly consists of Javan rusa (''Rusa timorensis''), though they also eat considerable amounts of carrion. Komodo dragons also occ ...
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Anguimorpha
The Anguimorpha is a suborder of squamates. The group was named by Fürbringer in 1900 to include all autarchoglossans closer to '' Varanus'' and '' Anguis'' than ''Scincus ''Scincus'' is a genus of skinks, lizards in the family Scincidae. The genus contains four or five species, all of which are typical desert inhabitants, living in sandy and dune-like areas with a hot and dry climate. Species in the genus ''Scinc ...''. These lizards, along with iguanians and snakes, constitute the proposed "venom clade" Toxicofera of all venomous reptiles. Evolution The oldest widely accepted member of Anguimorpha is ''Dorsetisaurus'' from the Late Jurassic of Europe and North America. In 2022, the genus ''Cryptovaranoides'' was described from the latest Triassic (Rhaetian) of England. ''Cryptovaranoides'' was recovered in the study as a crown-group Anguimorph, and therefore the oldest crown group-squamate, 35 million years older than any previously known crown-group squamate. Famili ...
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Saniwa
''Saniwa'' is an extinct genus of varanid lizard that lived about 48 million years ago during the Eocene epoch. It is known from well-preserved fossils found in the Bridger and Green River Formations of Wyoming. The type species ''S. ensidens'' was described in 1870 as the first fossil lizard known from North America. Several other species have since been added, but their validity is uncertain. It is a close relative of '' Varanus'', the genus that includes monitor lizards. Description ''Saniwa'' measured . Like other varanid lizards, ''Saniwa'' had a long, pointed snout and nostrils placed farther back in the skull than most lizards and a tail that was almost twice as long as the body. Although similar in appearance to extant monitor lizards, ''Saniwa'' had many primitive traits, including teeth on its palate, a jugal bone beneath the eye that extended farther forward, and a suture between the frontal and parietal bones that was straight rather than curved. A study in 2018 by ...
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Earless Monitor Lizard
The earless monitor lizard (''Lanthanotus borneensis'') is a semiaquatic, brown lizard native to the Southeast Asian island of Borneo. It is the only living species in the family Lanthanotidae and it is related to the true monitor lizards. Taxonomy The earless monitor lizard was described in 1878 by Franz Steindachner. The genus name ''Lanthanotus'' means "hidden ear" and the species name ''borneensis'' refers to its home island of Borneo. The uniqueness of the species was recognized from the start and Steindachner placed it in its own family, Lanthanotidae. In 1899 George Albert Boulenger relegated it to the family Helodermatidae, together with the beaded lizards and gila monster. Further studies were conducted in the 1950s where it was found that although it is related to Helodermatidae, this relationship is relatively distant. The similarity is in part the result of convergent evolution and they should be recognized as separate families. Both are part of a broader Anguimorph ...
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Ovoo Gurvel
''Ovoo gurvel'' is an extinct varanid lizard from the Late Cretaceous of Mongolia. It is one of the smallest and earliest monitor lizards. It was described in 2008. ''Ovoo'' possesses a pair of two small bones in its skull that are not seen in any other lizard. Description and history ''Ovoo'' is only known from a fossilized skull cataloged as IGM 3/767 and designated the holotype. The skull was discovered in 2001 near the rich Ukhaa Tolgod fossil site in a locality known as Little Ukhaa. The deposits at Little Uhhaa date back to the Campanian stage of the Late Cretaceous. The generic name is derived from the name of a type of cairn called ovoos that are found along roads near Little Ukhaa. The specific name, ''gurvel'', comes from the Mongolian word for lizard. ''Ovoo'' was very small compared to living monitor lizards, with the exception of the Short-tailed monitor. Nevertheless, the structures of its skull are very similar to those of living monitors. ''Ovoo'' shares many s ...
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Earless Monitor Lizard
The earless monitor lizard (''Lanthanotus borneensis'') is a semiaquatic, brown lizard native to the Southeast Asian island of Borneo. It is the only living species in the family Lanthanotidae and it is related to the true monitor lizards. Taxonomy The earless monitor lizard was described in 1878 by Franz Steindachner. The genus name ''Lanthanotus'' means "hidden ear" and the species name ''borneensis'' refers to its home island of Borneo. The uniqueness of the species was recognized from the start and Steindachner placed it in its own family, Lanthanotidae. In 1899 George Albert Boulenger relegated it to the family Helodermatidae, together with the beaded lizards and gila monster. Further studies were conducted in the 1950s where it was found that although it is related to Helodermatidae, this relationship is relatively distant. The similarity is in part the result of convergent evolution and they should be recognized as separate families. Both are part of a broader Anguimorph ...
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Telmasaurus
''Telmasaurus'' is an extinct genus of varanoid lizard from the Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ... of Mongolia. Fossils have been found from the Djadokhta Formation, Djadokha and Barun Goyot Formations that date between the early and middle Campanian stage from approximately 80 to 75 million years ago. The type species ''Telmasaurus grangeri'' was named in 1943. Description ''Telmasaurus grangeri'' was named in 1943 and is now known from several fossils. Only two skulls are known, and the snout region is missing from both. Based on the size of these skulls, ''Telmasaurus'' was larger than other Cretaceous varanoid lizards, approaching the size of modern monitor lizards. Its skull is flatter than most other varanoids. The fusion of two bones in t ...
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Lace Monitor
The lace monitor or tree goanna (''Varanus varius'') is a member of the monitor lizard family native to eastern Australia. A large lizard, it can reach in total length and in weight. The lace monitor is considered to be a least-concern species according to the International Union for Conservation of Nature. Taxonomy John White, the surgeon-general of New South Wales, described this species as the variegated lizard (''Lacerta varia'') in 1790. George Shaw reported that several specimens were taken back to England. French naturalist François Marie Daudin gave it the name ''Tupinambis variegatus'' in 1802, and noted two forms. German naturalist Blasius Merrem established the genus '' Varanus'' in 1820, with ''V. varius'' as the first mentioned member set as its type species by John Edward Gray in 1827. French zoologists André Marie Constant Duméril and Gabriel Bibron described two specimens in 1836, one in their possession and one from the collection of English zoologist Th ...
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Lanthanotus
The earless monitor lizard (''Lanthanotus borneensis'') is a semiaquatic, brown lizard native to the Southeast Asian island of Borneo. It is the only living species in the family Lanthanotidae and it is related to the true monitor lizards. Taxonomy The earless monitor lizard was described in 1878 by Franz Steindachner. The genus name ''Lanthanotus'' means "hidden ear" and the species name ''borneensis'' refers to its home island of Borneo. The uniqueness of the species was recognized from the start and Steindachner placed it in its own family, Lanthanotidae. In 1899 George Albert Boulenger relegated it to the family Helodermatidae, together with the beaded lizards and gila monster. Further studies were conducted in the 1950s where it was found that although it is related to Helodermatidae, this relationship is relatively distant. The similarity is in part the result of convergent evolution and they should be recognized as separate families. Both are part of a broader Anguimorph ...
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Archaeovaranus
''Archaeovaranus'' (meaning "ancient '' Varanus''") is genus of varanid lizard from the early Eocene (Ypresian) Yuhuangding Formation of Hubei Province, China. The genus contains a single species, ''Archaeovaranus lii'', known from a nearly complete skeleton. The holotype, which includes an intact skull, is associated but disarticulated. ''Archaeovaranus'' fills a gap in the varanid fossil record, as it represents a stem-varanid from the early Eocene of East Asia, and is the closest known relative of '' Varanus''. Discovery and naming The holotype specimen of ''Archaeovaranus'', IVPP V 22770, was discovered at the Dajian locality of the Yuhuangding Formation near Danjiangkou, Hubei Province, China. The generic name, "''Archaeovaranus''," combines the Greek "archaīos," meaning "ancient," with a reference to the closely related ''Varanus''. "''Varanus''" is derived from the Arabic "waral," meaning "lizard beast." The specific name, "''lii''," honors the paleontologist Chuank ...
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Varanoidea
Varanoidea is a superfamily of lizards, including the well-known family Varanidae (the monitors and goannas). Also included in the Varanoidea are the Lanthanotidae (earless monitor lizards), and the extinct Palaeovaranidae. Throughout their long evolutionary history, varanoids have exhibited great diversity, both in habitat and form. This superfamily includes the largest-known terrestrial lizard, Megalania (5–6 meters), and the largest extant lizard, the Komodo dragon (''Varanus komodoensis'', 3+ meters). Evolution Either synonymous with, or a subgroup of, the group Platynota, the varanoids first appear in the fossil record in the latter part of Early Cretaceous, but possible varanoid ancestors have been traced back as far as Early Jurassic times. Among the earliest known varanoids are the monitor-like necrosaurids '' Palaeosaniwa canadensis'' from the Campanian (roughly 71-82 mya) of North America and '' Estesia mongoliensis'' and '' Telmasaurus grangeri'', both from the Cam ...
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Savannah Monitor
The savannah monitor (''Varanus exanthematicus'') is a medium-sized species of monitor lizard native to Africa. The species is known as Bosc's monitor in Europe, since French scientist Louis Bosc first described the species. It belongs to the subgenus '' Polydaedalus''. Etymology The specific name ''exanthematicus'' is derived from the Greek word ''exanthem'' , meaning an eruption or blister of the skin. French botanist and zoologist Louis Augustin Guillaume Bosc originally described this lizard as ''Lacerta exanthematica'' in reference to the large oval scales on the back of its neck. The species was formerly known as ''Lacerta exanthematicus''. Description Savannah monitors are stoutly built, with relatively short limbs and toes, and skulls and dentition adapted to feed on hard-shelled prey. They are robust creatures, with powerful limbs for digging, powerful jaws and blunt, peglike teeth. Maximum size is rarely more than 100 cm. The skin coloration pattern varies accor ...
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