Ovoo Gurvel

''Ovoo gurvel'' is an extinct varanid lizard from the Late Cretaceous of Mongolia. It is one of the smallest and earliest monitor lizards. It was described in 2008. ''Ovoo'' possesses a pair of two small bones in its skull that are not seen in any other lizard.

Description and history

''Ovoo'' is only known from a fossilized skull cataloged as IGM 3/767 and designated the holotype. The skull was discovered in 2001 near the rich Ukhaa Tolgod fossil site in a locality known as Little Ukhaa. The deposits at Little Uhhaa date back to the Campanian stage of the Late Cretaceous. The generic name is derived from the name of a type of cairn called ovoos that are found along roads near Little Ukhaa. The specific name, ''gurvel'', comes from the Mongolian word for lizard.

''Ovoo'' was very small compared to living monitor lizards, with the exception of the Short-tailed monitor. Nevertheless, the structures of its skull are very similar to those of living monitors. ''Ovoo'' shares many similarities with the extinct monitors ''Aiolosaurus'' and ''Cherminotus'', which are also known from Little Ukhaa and Ukhaa Tolgod. Differences between these genera are seen in the shape of the bones in the skull. The distinguishing features of ''Ovoo'' include:
* Nasals, located behind the nostril openings, that are divided into two bones (in living monitors they are fused into one bone).
*The large size of a pair of holes called premaxillary fenestrae in front of the nostril openings.
*A bone called the septomaxilla that separates the premaxilla (the bone at the very tip of the snout) from the maxilla (the bone that makes up the upper jaw).
*The small size of a hole in the septomaxilla bone called the septomaxillary foramen.

The most unusual feature of ''Ovoo'' is the presence of two small bones that are not present in any other lizard. When it was first described, these structures were called "mystery bones". The two bones are located between the eye sockets. They are positioned behind the nasal bones and in front of the frontal bones. There are no homologous bones in any other animal, making their presence a mystery.

Classification

''Ovoo'' is one of many Late Cretaceous lizards belonging to a group called Varanoidea, which includes the living monitor and helodermatid lizards and the extinct mosasaurs. Features linking it with these lizards include a rounded snout and a lack of contact between the maxillae and frontal bones. A phylogenetic analysis conducted with its first description placed it within the monitor subfamily Varaninae. ''Ovoo'' is the oldest known member of Varaninae. Of the other Mongolian varanoids, ''Aiolosaurus'' was placed as a basal member of Varanidae (the family to which Varaninae belongs) and ''Cherminotus'' was placed in Lanthanotinae (another subfamily within Varanidae). Because of their age, ''Ovoo'', ''Aiolosaurus'', and ''Cherminotus'' may be representatives of the first radiation of monitor lizards. The Late Cretaceous varanoids '' Saniwides'' and ''Telmasaurus'' are just as old as these lizards and have traditionally been viewed as the oldest monitors, but they were placed outside the monitor family in the 2008 analysis. Below is a cladogram from the analysis:

References

{{Taxonbar, from1=Q4046102, from2=Q7114202

Late Cretaceous lepidosaurs of Asia
Monitor lizards
Fossil taxa described in 2008