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Uktenadactylus
''Uktenadactylus'' is a genus of anhanguerid pterodactyloid pterosaurs from the Lower Cretaceous Paw Paw Formation of Texas, United States and the Wessex Formation on the Isle of Wight, England. Fossil remains of ''Uktenadactylus'' dated back to the Early Cretaceous period (Barremian to Cenomanian stages), from about 125 to 100 million years ago. Discovery and naming In 1994, Yuong Nam-Lee named a new species within the genus ''Coloborhynchus'': ''Coloborhynchus wadleighi'', based on a partial snout found in 1992 in Albian layers in Tarrant County, holotype SMU 73058 ( Shuler Museum of Paleontology, Southern Methodist University at Dallas). The specific name honors the collector of the fossil, Chris Wadleigh. The reference of the species to the genus ''Coloborhynchus'' was based on the fact that both ''C. wadleighi'' and the type species of ''Coloborhynchus'', ''Coloborhynchus clavirostris'', share the trait of having three pairs of teeth laterally placed within a broad snout ti ...
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Coloborhynchus
''Coloborhynchus'' is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. ''Coloborhynchus'' is known from the Lower Cretaceous of England (Valanginian age, 140 to 136 million years ago), and depending on which species are included, possibly the Albian and Cenomanian ages (113 to 93.9 million years ago) as well. ''Coloborhynchus'' was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related ''Tropeognathus'' is now thought to have had a larger wingspan. History and classification Like many ornithocheiroid pterosaurs named during the 19th century, ''Coloborhynchus'' has a highly convoluted history of classification. Over the years numerous species have been assigned to it, and often, species have been shuffled between ''Coloborhynchus'' and related genera by various researchers. In 1874 Richard Owen, rejecting the creation by H ...
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Wessex Formation
The Wessex Formation is a fossil-rich English geological formation that dates from the Berriasian to Barremian stages (about 145–125 million years ago) of the Early Cretaceous. It forms part of the Wealden Group and underlies the younger Vectis Formation and overlies the Durlston Formation. The dominant lithology of this unit is mudstone with some interbedded sandstones. It is part of the strata of the Wessex Basin, exposed in both the Isle of Purbeck and the Isle of Wight. While the Purbeck sections are largely barren of vertebrate remains, the Isle of Wight sections are well known for producing the richest and most diverse fauna in Early Cretaceous Europe. Nomenclatural History The Wessex Formation has historically alternately been called the "Variegated Marls And Sandstones", a name used by W. J. Arkell in his 1947 map of the Isle of Purbeck as well as the "Wealden Marls" It was given its current formal name by Daley and Stewart in 1979 Stratigraphy and Lithology In ...
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Anhanguerid
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested ''Tropeognathus''. Relationships There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set. The cladogram below follows Pêgas ''et al.'' (2019), who recovered Anhangueridae as a much more inclusive group. The analysis found most of the ornithocheirids falling into this family, while ''Ornithocheirus'' itself was recovered as a basal member of Ornithocheirae Ornithocheirae is an extinct clad ...
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Paw Paw Formation
The Paw Paw Formation is a geological formation in Texas whose strata date back to the late Albian stage of the Early Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution (Early Cretaceous, North America)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 553-556. . Fossil content * '' Pawpawsaurus campbelli'' - "Skull.""Table 17.1," in Weishampel, et al. (2004). Page 365. * '' Texasetes pleurohalio'' - "Partial skeleton.""Table 17.1," in Weishampel, et al. (2004). Page 367.Coombs, 1995 * '' Turrilites'' * ''Scaphites'' * '' Pachyamia'' * '' Feldmannia'' * '' Cretacoranina'' * ''Coloborhynchus'' * ''Uktenadactylus'' * '' Engonoceras sp.'' * '' Pseudohypolophus sp.'' * '' Squalicorax sp.'' * '' Leptostyrax macrorhiza'' * '' Paraisurus compressus'' * '' Cretolamna appendiculata'' * Testudin ...
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Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java-Manihiki-Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kerguelen P ...
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Type Species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen(s). Article 67.1 A similar concept is used for suprageneric groups and called a type genus. In botanical nomenclature, these terms have no formal standing under the code of nomenclature, but are sometimes borrowed from zoological nomenclature. In botany, the type of a genus name is a specimen (or, rarely, an illustration) which is also the type of a species name. The species name that has that type can also be referred to as the type of the genus name. Names of genus and family ranks, the various subdivisions of those ranks, and some higher-rank names based on genus names, have such types.
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Southern Methodist University
, mottoeng = "The truth will make you free" , established = , type = Private research university , accreditation = SACS , academic_affiliations = , religious_affiliation = United Methodist Church , president = R. Gerald Turner , provost = Elizabeth G. Loboa , coor = , students = 12,373 (fall 2020) , undergrad = 6,827 (fall 2020) , postgrad = 5,546 (fall 2020) , faculty = 1,151; 754 full time (Fall 2019) , endowment = $2.0 billion (2021)As of June 30, 2020. , city = Dallas , state = Texas , country = United States , campus = Large City , campus_size= (main) , colors =  SMU Red SMU Blue , sports_nickname = Mustangs , athletics_affiliations = NCAA Division I FBS – AAC , mascot = Peruna , website = , logo = Southern Methodist University logo.svg , logo_upright = .8 , free_label2 = Newspaper , free2 = ''The Daily Campus'' , free_label = Other campuses , free = Taos Southern Methodist University (SMU) is a private research university in Univ ...
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Dallas
Dallas () is the List of municipalities in Texas, third largest city in Texas and the largest city in the Dallas–Fort Worth metroplex, the List of metropolitan statistical areas, fourth-largest metropolitan area in the United States at 7.5 million people. It is the largest city in and County seat, seat of Dallas County, Texas, Dallas County with portions extending into Collin County, Texas, Collin, Denton County, Texas, Denton, Kaufman County, Texas, Kaufman and Rockwall County, Texas, Rockwall counties. With a 2020 United States census, 2020 census population of 1,304,379, it is the List of United States cities by population, ninth most-populous city in the U.S. and the List of cities in Texas by population, third-largest in Texas after Houston and San Antonio. Located in the North Texas region, the city of Dallas is the main core of the largest metropolitan area in the Southern United States and the largest inland metropolitan area in the U.S. that lacks any navigable link ...
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Specific Name (zoology)
In zoological nomenclature, the specific name (also specific epithet or species epithet) is the second part (the second name) within the scientific name of a species (a binomen). The first part of the name of a species is the name of the genus or the generic name. The rules and regulations governing the giving of a new species name are explained in the article species description. For example, the scientific name for humans is ''Homo sapiens'', which is the species name, consisting of two names: ''Homo'' is the " generic name" (the name of the genus) and ''sapiens'' is the "specific name". Historically, ''specific name'' referred to the combination of what are now called the generic and specific names. Carl Linnaeus, who formalized binomial nomenclature, made explicit distinctions between specific, generic, and trivial names. The generic name was that of the genus, the first in the binomial, the trivial name was the second name in the binomial, and the specific the proper term for ...
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Reginald Walter Hooley
Reginald Walter Hooley (5 September 1865 – 5 May 1923) was a businessman and amateur paleontologist, collecting on the Isle of Wight. He is probably best remembered for describing the dinosaur ''Iguanodon atherfieldensis'', now ''Mantellisaurus.'' Biography Reginald Hooley was born on 5 September 1865 in Southampton, the son of William Hooley, a wealthy gentleman. In 1889 R.W. Hooley began to work for ''Godrich & Petman'', wine merchants, and later in life became managing director of that firm. Living in Portswood, in 1912 he married E.E. Holden and moved to Winchester. In 1913 he was elected a member of the Winchester city council. Hooley was a member of the Hampshire Field Club & Archaeological Society at Winchester from 1890. He was one of the founders of the Isle of Wight Natural History and Archaeological Society. He was an honorary curator of the Winchester Museum between 1918 and 1923. Isle of Wight Hooley made regular visits to the Isle of Wight and found hundreds of ...
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Criorhynchus
''Ornithocheirus'' (from Ancient Greek "ὄρνις", meaning ''bird'', and "χεῖρ", meaning ''hand'') is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the UK and possibly Morocco. Several species have been referred to the genus, most of which are now considered as dubious species, or members of different genera, and the genus is now often considered to include only the type species, ''Ornithocheirus simus''. Species have been referred to ''Ornithocheirus'' from the mid-Cretaceous period of both Europe and South America, but ''O. simus'' is known only from the UK, though a specimen referred to ''O.'' cf. ''simus'' is also known from Morocco. Because ''O. simus'' was originally named based on poorly preserved fossil material, the genus ''Ornithocheirus'' has suffered enduring problems of zoological nomenclature. Fossil remains of ''Ornithocheirus'' have been recovered mainly from the Cambridge Greensand of England, dating to the beginni ...
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Tarrant County
Tarrant County is located in the U.S. state of Texas. As of 2020, it had a population of 2,110,640. It is Texas' third-most populous county and the 15th-most populous in the United States. Its county seat is Fort Worth. Tarrant County, one of 26 counties created out of the Peters Colony, was established in 1849 and organized the next year. It was named in honor of General Edward H. Tarrant of the Republic of Texas militia. Geography According to the U.S. Census Bureau, the county has an area of , of which is land and (4.3%) is water. Adjacent counties * Denton County (north) * Dallas County (east) * Ellis County (southeast) * Johnson County (south) * Parker County (west) * Wise County (northwest) Communities Cities (multiple counties) * Azle (partly in Parker County) * Burleson (mostly in Johnson County) * Crowley (small part in Johnson County) * ''Fort Worth'' (small parts in Denton, Johnson, Parker and Wise counties) * Grand Prairie (partly in Dallas Coun ...
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