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Coloborhynchus
''Coloborhynchus'' is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. ''Coloborhynchus'' is known from the Lower Cretaceous of England (Valanginian age, 140 to 136 million years ago), and depending on which species are included, possibly the Albian and Cenomanian ages (113 to 93.9 million years ago) as well. ''Coloborhynchus'' was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related ''Tropeognathus'' is now thought to have had a larger wingspan. History and classification Like many ornithocheiroid pterosaurs named during the 19th century, ''Coloborhynchus'' has a highly convoluted history of classification. Over the years numerous species have been assigned to it, and often, species have been shuffled between ''Coloborhynchus'' and related genera by various researchers. In 1874 Richard Owen, rejecting the creation by H ...
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Coloborhynchus
''Coloborhynchus'' is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. ''Coloborhynchus'' is known from the Lower Cretaceous of England (Valanginian age, 140 to 136 million years ago), and depending on which species are included, possibly the Albian and Cenomanian ages (113 to 93.9 million years ago) as well. ''Coloborhynchus'' was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related ''Tropeognathus'' is now thought to have had a larger wingspan. History and classification Like many ornithocheiroid pterosaurs named during the 19th century, ''Coloborhynchus'' has a highly convoluted history of classification. Over the years numerous species have been assigned to it, and often, species have been shuffled between ''Coloborhynchus'' and related genera by various researchers. In 1874 Richard Owen, rejecting the creation by H ...
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Ornithocheiridae
Ornithocheiridae (or ornithocheirids, meaning "bird hands") is a group of pterosaurs within the suborder Pterodactyloidea. These pterosaurs were among the last to possess teeth. Members that belong to this group lived from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 90 million years ago. Ornithocheirids are generally infamous for having an enormously controversial and very confusing taxonomy. Although agreements that these animals were related, and therefore similar to istiodactylids and pteranodontians, there is still no virtual consensus over the exact content and interrelationships of this group. Ornithocheirids were the most successful pterosaurs during their reign, they were also the largest pterosaurs before the appearance of the azhdarchids such as ''Quetzalcoatlus''. Ornithocheirids were excellent fish hunters, they used various flight techniques to catch their prey, and they are also capable of flying great distances without fl ...
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Anhanguera Piscator
''Anhanguera'' () is a genus of pterodactyloid pterosaur known from the Early Cretaceous (Albian age, 125 to 112 million years ago) Romualdo Formation of Brazil and the Late Cretaceous ( Cenomanian age, 98 to 92.5 million years ago) Kem Kem Group of Morocco. This pterosaur is closely related to ''Ornithocheirus'', but belongs in the family Anhangueridae.Campos, D. de A., and Kellner, A. W. (1985). "Um novo exemplar de ''Anhanguera blittersdorffi'' (Reptilia, Pterosauria) da formação Santana, Cretaceo Inferior do Nordeste do Brasil." In Congresso Brasileiro de Paleontologia, Rio de Janeiro, Resumos, p. 13. The generic name comes from the Tupi words ''añanga'', meaning "spirit protector of the animals" + ''wera'' "bygone". Description ''Anhanguera'' was a fish-eating animal with a wingspan of about .Aureliano, T., Ghilardi, A. M., Duque, R. R., & Barreto, A. M. (2014). ON THE OCCURRENCE OF PTEROSAURIA IN EXU, PERNAMBUCO (LOWER CRETACEOUS ROMUALDO FORMATION, ARARIPE BASIN), N ...
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Anhanguera Robustus
''Anhanguera'' () is a genus of pterodactyloid pterosaur known from the Early Cretaceous (Albian age, 125 to 112 million years ago) Romualdo Formation of Brazil and the Late Cretaceous ( Cenomanian age, 98 to 92.5 million years ago) Kem Kem Group of Morocco. This pterosaur is closely related to ''Ornithocheirus'', but belongs in the family Anhangueridae.Campos, D. de A., and Kellner, A. W. (1985). "Um novo exemplar de ''Anhanguera blittersdorffi'' (Reptilia, Pterosauria) da formação Santana, Cretaceo Inferior do Nordeste do Brasil." In Congresso Brasileiro de Paleontologia, Rio de Janeiro, Resumos, p. 13. The generic name comes from the Tupi words ''añanga'', meaning "spirit protector of the animals" + ''wera'' "bygone". Description ''Anhanguera'' was a fish-eating animal with a wingspan of about .Aureliano, T., Ghilardi, A. M., Duque, R. R., & Barreto, A. M. (2014). ON THE OCCURRENCE OF PTEROSAURIA IN EXU, PERNAMBUCO (LOWER CRETACEOUS ROMUALDO FORMATION, ARARIPE BASIN), N ...
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Anhanguera (pterosaur)
''Anhanguera'' () is a genus of pterodactyloid pterosaur known from the Early Cretaceous (Albian age, 125 to 112 million years ago) Romualdo Formation of Brazil and the Late Cretaceous ( Cenomanian age, 98 to 92.5 million years ago) Kem Kem Group of Morocco. This pterosaur is closely related to ''Ornithocheirus'', but belongs in the family Anhangueridae.Campos, D. de A., and Kellner, A. W. (1985). "Um novo exemplar de ''Anhanguera blittersdorffi'' (Reptilia, Pterosauria) da formação Santana, Cretaceo Inferior do Nordeste do Brasil." In Congresso Brasileiro de Paleontologia, Rio de Janeiro, Resumos, p. 13. The generic name comes from the Tupi words ''añanga'', meaning "spirit protector of the animals" + ''wera'' "bygone". Description ''Anhanguera'' was a fish-eating animal with a wingspan of about .Aureliano, T., Ghilardi, A. M., Duque, R. R., & Barreto, A. M. (2014). ON THE OCCURRENCE OF PTEROSAURIA IN EXU, PERNAMBUCO (LOWER CRETACEOUS ROMUALDO FORMATION, ARARIPE BASIN), N ...
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Nicorhynchus
''Nicorhynchus'' (meaning "knucker snout", in reference to its likely ecology) is a genus of anhanguerid pterosaur from the Cretaceous period. It contains two species, the type species, ''N. capito'', from the Cambridge Greensand of England, and ''N. fluviferox'' from the Kem Kem Group of Morocco. These species were previously assigned to ''Coloborhynchus''. History The genus ''Coloborhynchus'' has had a convoluted history, with many species having been assigned to the genus. In 2013, Rodrigues and Kellner considered ''Coloborhynchus'' to be monotypic, containing only ''C. clavirostris'', and placed most other species in other genera, or declared them '' nomina dubia''. One of these species was ''Coloborhynchus capito'', which was originally named by Harry Seeley in 1870 as a species of ''Ornithocheirus''. Its holotype is CAMSM B 54625, a snout. In 2001, this species was moved to ''Coloborhynchus'' by David Unwin, who also synonymized ''Ornithocheirus reedi'' (known from a lost sp ...
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Nicorhynchus Capito
''Nicorhynchus'' (meaning "knucker snout", in reference to its likely ecology) is a genus of anhanguerid pterosaur from the Cretaceous period. It contains two species, the type species, ''N. capito'', from the Cambridge Greensand of England, and ''N. fluviferox'' from the Kem Kem Group of Morocco. These species were previously assigned to ''Coloborhynchus''. History The genus ''Coloborhynchus'' has had a convoluted history, with many species having been assigned to the genus. In 2013, Rodrigues and Kellner considered ''Coloborhynchus'' to be monotypic, containing only ''C. clavirostris'', and placed most other species in other genera, or declared them ''nomina dubia''. One of these species was ''Coloborhynchus capito'', which was originally named by Harry Seeley in 1870 as a species of ''Ornithocheirus''. Its holotype is CAMSM B 54625, a snout. In 2001, this species was moved to ''Coloborhynchus'' by David Unwin, who also synonymized ''Ornithocheirus reedi'' (known from a lost spe ...
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Nicorhynchus Fluviferox
''Nicorhynchus'' (meaning "knucker snout", in reference to its likely ecology) is a genus of anhanguerid pterosaur from the Cretaceous period. It contains two species, the type species, ''N. capito'', from the Cambridge Greensand of England, and ''N. fluviferox'' from the Kem Kem Group of Morocco. These species were previously assigned to ''Coloborhynchus''. History The genus ''Coloborhynchus'' has had a convoluted history, with many species having been assigned to the genus. In 2013, Rodrigues and Kellner considered ''Coloborhynchus'' to be monotypic, containing only ''C. clavirostris'', and placed most other species in other genera, or declared them ''nomina dubia''. One of these species was ''Coloborhynchus capito'', which was originally named by Harry Seeley in 1870 as a species of ''Ornithocheirus''. Its holotype is CAMSM B 54625, a snout. In 2001, this species was moved to ''Coloborhynchus'' by David Unwin, who also synonymized ''Ornithocheirus reedi'' (known from a lost spe ...
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Ornithocheirus
''Ornithocheirus'' (from Ancient Greek "ὄρνις", meaning ''bird'', and "χεῖρ", meaning ''hand'') is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the UK and possibly Morocco. Several species have been referred to the genus, most of which are now considered as dubious species, or members of different genera, and the genus is now often considered to include only the type species, ''Ornithocheirus simus''. Species have been referred to ''Ornithocheirus'' from the mid-Cretaceous period of both Europe and South America, but ''O. simus'' is known only from the UK, though a specimen referred to ''O.'' cf. ''simus'' is also known from Morocco. Because ''O. simus'' was originally named based on poorly preserved fossil material, the genus ''Ornithocheirus'' has suffered enduring problems of zoological nomenclature. Fossil remains of ''Ornithocheirus'' have been recovered mainly from the Cambridge Greensand of England, dating to the beginni ...
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Maaradactylus
''Maaradactylus'' is a genus of anhanguerid pterodactyloid pterosaur known from the Lower Cretaceous period (Aptian to Albian stages) of the Romualdo Formation of northeastern Brazil. Discovery ''Maaradactylus'' is based on the Museu Paleontologico de Santana do Cariri specimen MPSC R 2357, a skull, atlas, and axis discovered in 2010 in the Aptian—Albian-age Romualdo Formation of Sítio São Gonçalo, Santana do Cariri, Ceará, in the Araripe Basin of Brazil. ''Maaradactylus'' was described by Renan Bantim and colleagues in 2014. The type species is ''Maaradactylus kellneri''. The generic name refers to Maara, in the legends of the Cariri the daughter of a chief, by sorcery changed into a river monster with long teeth, devouring fishermen. The suffix ~''dactylus'' is common in the names of pterosaurs and is derived from Greek δάκτυλος, ''daktylos'', "finger", referring to the long (fourth) wing finger. The specific name honors Alexander Kellner, Brazil's f ...
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Tropeognathus
''Tropeognathus'' (meaning "keel jaw") is a genus of large pterosaurs from the late Early Cretaceous of South America. This genus is considered to be a member of the family Anhangueridae, however, several studies have also recovered it within another family called Ornithocheiridae. Both of these families are diverse groups of pterosaurs known for their keel-tipped snouts and large size. ''Tropeognathus'' is regarded as the largest pterosaur found in the Southern Hemisphere, only rivaled by the huge azhdarchids. The type and only species is ''Tropeognathus mesembrinus''. Fossil remains of ''Tropeognathus'' have been recovered from the Romualdo Formation, which is a Lagerstätte located in the Santana Group of the Araripe Basin in northeastern Brazil. Discovery and naming In the 1980s the German paleontology museum '' Bayerische Staatssammlung für Paläontologie und historische Geologie'' in Munich acquired a pterosaur skull from Brazilian fossil dealers that had probably bee ...
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Maaradactylus Spielbergi
''Maaradactylus'' is a genus of anhanguerid pterodactyloid pterosaur known from the Lower Cretaceous period (Aptian to Albian stages) of the Romualdo Formation of northeastern Brazil. Discovery ''Maaradactylus'' is based on the Museu Paleontologico de Santana do Cariri specimen MPSC R 2357, a skull, atlas, and axis discovered in 2010 in the Aptian—Albian-age Romualdo Formation of Sítio São Gonçalo, Santana do Cariri, Ceará, in the Araripe Basin of Brazil. ''Maaradactylus'' was described by Renan Bantim and colleagues in 2014. The type species is ''Maaradactylus kellneri''. The generic name refers to Maara, in the legends of the Cariri the daughter of a chief, by sorcery changed into a river monster with long teeth, devouring fishermen. The suffix ~''dactylus'' is common in the names of pterosaurs and is derived from Greek δάκτυλος, ''daktylos'', "finger", referring to the long (fourth) wing finger. The specific name honors Alexander Kellner, Brazil's fo ...
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