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Nicorhynchus Fluviferox
''Nicorhynchus'' (meaning "knucker snout", in reference to its likely ecology) is a genus of anhanguerid pterosaur from the Cretaceous period. It contains two species, the type species, ''N. capito'', from the Cambridge Greensand of England, and ''N. fluviferox'' from the Kem Kem Group of Morocco. These species were previously assigned to ''Coloborhynchus''. History The genus ''Coloborhynchus'' has had a convoluted history, with many species having been assigned to the genus. In 2013, Rodrigues and Kellner considered ''Coloborhynchus'' to be monotypic, containing only ''C. clavirostris'', and placed most other species in other genera, or declared them ''nomina dubia''. One of these species was ''Coloborhynchus capito'', which was originally named by Harry Seeley in 1870 as a species of ''Ornithocheirus''. Its holotype is CAMSM B 54625, a snout. In 2001, this species was moved to ''Coloborhynchus'' by David Unwin, who also synonymized ''Ornithocheirus reedi'' (known from a lost spe ...
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Albian
The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 Ma (million years ago). The Albian is preceded by the Aptian and followed by the Cenomanian. Stratigraphic definitions The Albian Stage was first proposed in 1842 by Alcide d'Orbigny. It was named after Alba, the Latin name for River Aube in France. A Global Boundary Stratotype Section and Point (GSSP), ratified by the IUGS in 2016, defines the base of the Albian as the first occurrence of the planktonic foraminiferan '' Microhedbergella renilaevis'' at the Col de Pré-Guittard section, Arnayon, Drôme, France. The top of the Albian Stage (the base of the Cenomanian Stage and Upper Cretaceous Series) is defined as the place where the foram species '' Rotalipora globotruncanoides'' first appears in the stratigraphic column. The Albia ...
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Nicorhynchus Fluviferox
''Nicorhynchus'' (meaning "knucker snout", in reference to its likely ecology) is a genus of anhanguerid pterosaur from the Cretaceous period. It contains two species, the type species, ''N. capito'', from the Cambridge Greensand of England, and ''N. fluviferox'' from the Kem Kem Group of Morocco. These species were previously assigned to ''Coloborhynchus''. History The genus ''Coloborhynchus'' has had a convoluted history, with many species having been assigned to the genus. In 2013, Rodrigues and Kellner considered ''Coloborhynchus'' to be monotypic, containing only ''C. clavirostris'', and placed most other species in other genera, or declared them ''nomina dubia''. One of these species was ''Coloborhynchus capito'', which was originally named by Harry Seeley in 1870 as a species of ''Ornithocheirus''. Its holotype is CAMSM B 54625, a snout. In 2001, this species was moved to ''Coloborhynchus'' by David Unwin, who also synonymized ''Ornithocheirus reedi'' (known from a lost spe ...
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Aerodraco
''Aerodraco'' (meaning "air dragon") is a genus of anhanguerid pterosaur from the Albian–Cenomanian-age Cambridge Greensand of England. It contains only one species, ''Aerodraco sedgwickii''. It was originally assigned to the genus ''Pterodactylus''. Discovery and naming In 1859, Sir Richard Owen named pterosaur material from the Cambridge Greensand of England as ''Pterodactylus sedgwickii''.R. Owen. 1859. Supplement (No. I) to the Monograph on the Fossil Reptilia of the Cretaceous Formations. Palaeontographical Society Monographs 1–19 At the time, ''Pterodactylus'' was a wastebasket taxon; all sorts of unrelated pterosaurs were assigned to that genus. In 1870, Harry Seeley reassigned it to ''Ornithocheirus'', another wastebasket taxon. Its specific name honors Adam Sedgwick. It was in 1869 renamed by Seeley into a ''Ptenodactylus sedgwickii'',Seeley, H.G., 1869, ''Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arrange ...
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Ferrodraco
''Ferrodraco'' ("Iron Dragon" after the ironstone the fossil was found in) is an extinct genus of anhanguerid pterosaur known from the Late Cretaceous Winton Formation of Queensland, Australia, containing the single species ''F. lentoni''. The species was named after the former mayor of Winton, Graham Thomas ‘Butch’ Lenton. It is the most complete pterosaur fossil from Australia, being known from the holotype specimen AODF 876, consisting primarily of the anterior portion of the skull and dentary, cervical vertebral centra and a partial wing. Its wingspan was estimated to be about . ''Ferrodraco'' was found to have been within the subfamily Ornithocheirinae, as sister taxon to '' Mythunga''. A recent study also recovered ''Ferrodraco'' as sister taxon to ''Mythunga'', but both placed within the family Anhangueridae, more specifically within the subfamily Tropeognathinae. ''Ferrodraco'' is also the latest surviving member of Anhangueria. Discovery and naming The holotyp ...
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Mythunga
''Mythunga'' is a genus of anhanguerid pterosaur from the late Early Cretaceous of Australia. Fossil remains of ''Mythunga'' dated back to the Albian stage of the Early Cretaceous, and the animal itself was found to be a close relative of another Australian anhanguerid called ''Ferrodraco''. Discovery and naming ''Mythunga'' is known from a partial skull, holotype QM F18896 found in April 1991 by Philip Gilmore in marine rocks of the late Albian-age Toolebuc Formation at Dunluce Station west of Hughenden, Queensland. Only the middle snout and corresponding parts of the lower jaws are known, including the rear of a left premaxilla, the lower parts of both maxillae, the rear dentaries and a right splenial. They were three-dimensionally preserved, associated in a chalk nodule. It represents a subadult individual. The fossil was prepared by Angela Hatch of the Queensland Museum, both by mechanical means and by an acid bath. The type species ''Mythunga camara'' was named and descri ...
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Siroccopteryx
''Siroccopteryx'' is an extinct genus of anhanguerid pterodactyloid pterosaur, known from middle Cretaceous (between the Albian and Cenomanian stages, about 105 million years ago) sediments in modern-day Morocco. Some researchers, such as David M. Unwin, consider the genus a junior synonym of ''Coloborhynchus''. Description The genus was named and described in 1999 by Bryn Mader and Alexander Kellner. The name ''Siroccopteryx'' means "wing of the Sirocco", referring to the warm wind that originates in the North Africa and then goes through the Mediterranean, and the Greek word ''pteryx'', a standard suffix for pterosaur genera that means "wing". The epithet of the type species ''S. moroccensis'' refers to its country of origin. This pterosaur is known only from the front part of the jaw with teeth. The holotype fossil, LINHM 016 (Long Island Natural History Museum), was found near Ksar es Souk, in the province of Errachidia in the region of Meknes-Tafilalet at 30.4 ° N, 4 ...
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Tropeognathinae
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a Cladistics, cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested ''Tropeognathus''. Relationships There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set. The cladogram below follows Pêgas ''et al.'' (2019), who recovered Anhangueridae as a much more inclusive group. The analysis found most of the ornithocheirids falling into this family, while ''Ornithocheirus'' itself was recovered as a basal member of Ornithocheirae. References

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Anhangueridae
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested ''Tropeognathus''. Relationships There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set. The cladogram below follows Pêgas ''et al.'' (2019), who recovered Anhangueridae as a much more inclusive group. The analysis found most of the ornithocheirids falling into this family, while ''Ornithocheirus'' itself was recovered as a basal member of Ornithocheirae Ornithocheirae is an extinct clad ...
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Uktenadactylus
''Uktenadactylus'' is a genus of anhanguerid pterodactyloid pterosaurs from the Lower Cretaceous Paw Paw Formation of Texas, United States and the Wessex Formation on the Isle of Wight, England. Fossil remains of ''Uktenadactylus'' dated back to the Early Cretaceous period (Barremian to Cenomanian stages), from about 125 to 100 million years ago. Discovery and naming In 1994, Yuong Nam-Lee named a new species within the genus ''Coloborhynchus'': ''Coloborhynchus wadleighi'', based on a partial snout found in 1992 in Albian layers in Tarrant County, holotype SMU 73058 ( Shuler Museum of Paleontology, Southern Methodist University at Dallas). The specific name honors the collector of the fossil, Chris Wadleigh. The reference of the species to the genus ''Coloborhynchus'' was based on the fact that both ''C. wadleighi'' and the type species of ''Coloborhynchus'', ''Coloborhynchus clavirostris'', share the trait of having three pairs of teeth laterally placed within a broad snout ti ...
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England
England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe by the North Sea to the east and the English Channel to the south. The country covers five-eighths of the island of Great Britain, which lies in the North Atlantic, and includes over 100 smaller islands, such as the Isles of Scilly and the Isle of Wight. The area now called England was first inhabited by modern humans during the Upper Paleolithic period, but takes its name from the Angles, a Germanic tribe deriving its name from the Anglia peninsula, who settled during the 5th and 6th centuries. England became a unified state in the 10th century and has had a significant cultural and legal impact on the wider world since the Age of Discovery, which began during the 15th century. The English language, the Anglican Church, and Engli ...
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Tropeognathus
''Tropeognathus'' (meaning "keel jaw") is a genus of large pterosaurs from the late Early Cretaceous of South America. This genus is considered to be a member of the family Anhangueridae, however, several studies have also recovered it within another family called Ornithocheiridae. Both of these families are diverse groups of pterosaurs known for their keel-tipped snouts and large size. ''Tropeognathus'' is regarded as the largest pterosaur found in the Southern Hemisphere, only rivaled by the huge azhdarchids. The type and only species is ''Tropeognathus mesembrinus''. Fossil remains of ''Tropeognathus'' have been recovered from the Romualdo Formation, which is a Lagerstätte located in the Santana Group of the Araripe Basin in northeastern Brazil. Discovery and naming In the 1980s the German paleontology museum '' Bayerische Staatssammlung für Paläontologie und historische Geologie'' in Munich acquired a pterosaur skull from Brazilian fossil dealers that had probably bee ...
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Coloborhynchus Fluviferox By Nick Longrich
''Coloborhynchus'' is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. ''Coloborhynchus'' is known from the Lower Cretaceous of England (Valanginian age, 140 to 136 million years ago), and depending on which species are included, possibly the Albian and Cenomanian ages (113 to 93.9 million years ago) as well. ''Coloborhynchus'' was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related ''Tropeognathus'' is now thought to have had a larger wingspan. History and classification Like many ornithocheiroid pterosaurs named during the 19th century, ''Coloborhynchus'' has a highly convoluted history of classification. Over the years numerous species have been assigned to it, and often, species have been shuffled between ''Coloborhynchus'' and related genera by various researchers. In 1874 Richard Owen, rejecting the creation by Ha ...
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