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Thoatherium
''Thoatherium'' (meaning "active swift-beast") is an extinct genus of litoptern mammals from the Early Miocene of Argentina. Fossils of the genus have been found in the Santa Cruz Formation in Argentina.''Thoatherium''
at Fossilworks.org


Description

With a length of , the -like ''Thoatherium'' was a small representative of the order
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Proterotheriids
Proterotheriidae is an extinct family of fossil ungulates from the Cenozoic era that displays toe reduction. Despite resembling primitive, small horses, they were only distantly related to them, and instead belonged to the native South American ungulate order Litopterna. Description Two subfamilies and 18 genera of Proterotheriidae are known. All forms were small or medium-sized. Typical is a reduction of the number of toes and brachydont or mesodont teeth. The family is recorded since the late Palaeocene. Various fossils are known from many parts of the South American continent. The diversity decreased in the Miocene to Pliocene and it has been assumed for a long time that they entirely disappeared in the late Pliocene. However, fossils found in Argentina, Brazil and Uruguay show that one member of the group, '' Neolicaphrium recens'' survived into the Late Pleistocene. Better known genera of the family include '' Diadiaphorus'' and '' Thoatherium'' from the Miocene. Ta ...
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Litopterna
Litopterna (from grc, λῑτή πτέρνα "smooth heel") is an extinct order of fossil hoofed mammals from the Cenozoic era. The order is one of the five great orders of South American ungulates that were endemic to the continent, until the Great American Biotic Interchange brought new ungulate species. Like other endemic South American mammals, their relationship to other mammal groups had long been unclear, but recent genetic and proteomic evidence indicates that their closest living relatives are Perissodactyls (odd-toed ungulates) including horses, rhinoceros, and tapirs, and that litopterns are closely related to notoungulates, another widespread group of South American ungulates. There were two major groups of litopterns: Proterotheriidae and Macraucheniidae. Proterotheriids were medium to large animals that evolved adaptations for fast running, and occupied a variety of niches that elsewhere were filled by animals such as goats and antelopes, mouse deer, and h ...
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Litoptern
Litopterna (from grc, λῑτή πτέρνα "smooth heel") is an extinct order of fossil hoofed mammals from the Cenozoic era. The order is one of the five great orders of South American ungulates that were endemic to the continent, until the Great American Biotic Interchange brought new ungulate species. Like other endemic South American mammals, their relationship to other mammal groups had long been unclear, but recent genetic and proteomic evidence indicates that their closest living relatives are Perissodactyls (odd-toed ungulates) including horses, rhinoceros, and tapirs, and that litopterns are closely related to notoungulates, another widespread group of South American ungulates. There were two major groups of litopterns: Proterotheriidae and Macraucheniidae. Proterotheriids were medium to large animals that evolved adaptations for fast running, and occupied a variety of niches that elsewhere were filled by animals such as goats and antelopes, mouse deer, and hors ...
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Santa Cruz Formation, Argentina
The Santa Cruz Formation is a geological formation in the Magallanes/Austral Basin in southern Patagonia in Argentina and in adjacent areas of Chile. It dates to the late Early Miocene epoch, and is contemporaneous with eponymous Santacrucian SALMA. The formation extends from the Andes to the Atlantic coast. In its coastal section it is divided into two members, the lower, fossil rich Estancia La Costa Member, which has a lithology predominantly consisting of tuffaceous deposits and fine grained sedimentary claystone and mudstone, and the upper fossil-poor Estancia La Angelina Member, which consists of sedimentary rock, primarily claystone, mudstone, and sandstone. The environment of deposition is interpreted to have been mostly fluvial, with the lowermost part of the Estancia La Costa Member being transitional between fluvial and marine conditions. The environment of the Estancia La Costa Member is thought to have been relatively warm and humid, but likely became somewhat coole ...
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Santacrucian
The Santacrucian age is a period of geologic time (17.5 – 16.3 Mya (unit), Ma) within the Early Miocene epoch of the Neogene, used more specifically with South American land mammal age, SALMA classification in South America. It follows the Colhuehuapian and precedes the Friasian age. Etymology The age is named after the Santa Cruz Formation in the Magallanes Basin, Austral/Magallanes Basin of southern Patagonia, Argentina and Chile. Formations Fossils References Bibliography

;Santa Cruz Formation * * * * * * * * * * * * * * * * ;Aisol Formation * * ;Cantaure Formation * ;Castillo Formation * * * * * ;Cerro Boleadores Formation * ;Chaguaramas Formation * ;Chilcatay Formation * * * * ;Cura-Mallín Group * * * ;Gran Bajo del Gualicho Formation * ;Jimol Formation * * * * * * * * ;Mariño Formation * * * * * ;Pinturas Formation * * ;Río Yuca Formation * {{SALMA Santacrucian, Miocene South America ...
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Proterotherium
''Proterotherium'' (meaning "first beast") is an extinct genus of litoptern mammal of the family Proterotheriidae that lived during the Late Miocene of Argentina and Chile. Fossils of this genus have been found in the Ituzaingó Formation of Argentina, and the Galera Formation of Chile. Classification The genus ''Proterotherium'' was first described by Florentino Ameghino in 1883, on the basis of fossil remains found in Upper Miocene deposits in Patagonia. The type species is ''Proterotherium cervioides''. Numerous other species have been ascribed to this genus, many of which come from lower-middle Miocene deposits, such as ''P. americanum'', ''P. australe'', ''P. brachygnathum'', ''P. cavum'', ''P. cingulatum'', ''P. curtidens'', ''P. dichotomum'' , ''P. divortium'', ''P. gradatum'', ''P. intermedium'', ''P. karaikense'', ''P. mixtum'', ''P. nitens'', ''P. perpolitum'', ''P. politum'', ''P. main'', and ''P. pyramidatum''. However, many of these species are now considered to be ...
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Equine Forelimb Anatomy
Good conformation in the limbs leads to improved movement and decreased likelihood of injuries. Large differences in bone structure and size can be found in horses used for different activities, but correct conformation remains relatively similar across the spectrum. Structural defects, as well as other problems such as injuries and infections, can cause lameness, or movement at an abnormal gait. Injuries to and problems with horse legs can be relatively minor, such as stocking up, which causes swelling without lameness, or quite serious. Even leg injuries that are not immediately fatal may still be life-threatening to horses, as their bodies are adapted to bear weight on all four legs and serious problems can result if this is not possible. Limb anatomy Horses are odd-toed ungulates, or members of the order Perissodactyla. This order also includes the extant species of rhinos and tapirs, and many extinct families and species. Members of this order walk on either one toe (lik ...
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Fossil Taxa Described In 1887
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the abso ...
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Fossils Of Argentina
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolu ...
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Neogene Argentina
The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. The Neogene is sub-divided into two epochs, the earlier Miocene and the later Pliocene. Some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary. The term "Neogene" was coined in 1853 by the Austrian palaeontologist Moritz Hörnes (1815–1868). During this period, mammals and birds continued to evolve into modern forms, while other groups of life remained relatively unchanged. The first humans (''Homo habilis'') appeared in Africa near the end of the period. Some continental movements took place, the most significant event being the connection of North and South America at the Isthmus of Panama, late in the Pliocene. This cut off the warm ocean currents from the Pacific to the ...
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Miocene Mammals Of South America
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the late ...
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Burdigalian Life
The Burdigalian is, in the geologic timescale, an age or stage in the early Miocene. It spans the time between 20.43 ± 0.05 Ma and 15.97 ± 0.05 Ma (million years ago). Preceded by the Aquitanian, the Burdigalian was the first and longest warming period of the MioceneEdward Petuch, Ph.D. Florida Atlantic University, Department of Geosciences. and is succeeded by the Langhian. Stratigraphic definition The name Burdigalian comes from ''Burdigala'', the Latin name for the city of Bordeaux, France. The Burdigalian Stage was introduced in scientific literature by Charles Depéret in 1892. The base of the Burdigalian is at the first appearance of foram species ''Globigerinoides altiaperturus'' and the top of magnetic chronozone C6An. , an official GSSP for the Burdigalian had not yet been assigned. The top of the Burdigalian (the base of the Langhian) is defined by the first appearance of foram species ''Praeorbulina glomerosa'' and is also coeval with the top of magnetic chro ...
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