Proterotheriids
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Proterotheriids
Proterotheriidae is an extinct family of fossil ungulates from the Cenozoic era that displays toe reduction. Despite resembling primitive, small horses, they were only distantly related to them, and instead belonged to the native South American ungulate order Litopterna. Description Two subfamilies and 18 genera of Proterotheriidae are known. All forms were small or medium-sized. Typical is a reduction of the number of toes and brachydont or mesodont teeth. The family is recorded since the late Palaeocene. Various fossils are known from many parts of the South American continent. The diversity decreased in the Miocene to Pliocene and it has been assumed for a long time that they entirely disappeared in the late Pliocene. However, fossils found in Argentina, Brazil and Uruguay show that one member of the group, '' Neolicaphrium recens'' survived into the Late Pleistocene. Better known genera of the family include '' Diadiaphorus'' and '' Thoatherium'' from the Miocene. Ta ...
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Paramacrauchenia
''Paramacrauchenia'' is an extinct genus of proterotheriid litopterns from the Early Miocene of what is now Argentina and Chile. Its fossils have been found in the Sarmiento and Santa Cruz Formations of Argentina and Chile. Taxonomy ''Paramacrauchenia'' was originally assigned to the family Macraucheniidae, however, recent studies now consider it a member of the family Proterotheriidae, where it is found to be a derived member, closely related to ''Lambdaconus ''Lambdaconus'' is a genus of proterotheriid from the Late Oligocene to Early Miocene of Argentina. The type species is ''L. suinus'', named in 1897 by Ameghino, with referred species including ''L. lacerum'', named as ''Proterotherium ''Prot ...''. Below is a phylogenetic tree of the Proterotheriidae, based on the work of McGrath ''et al''. 2020. References Proterotheriids Miocene mammals of South America Colhuehuapian Santacrucian Neogene Argentina Fossils of Argentina Neogene Chile Fossils of Chi ...
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Neolicaphrium
''Neolicaphrium'' is an extinct genus of ungulate mammal belonging to the extinct order Litopterna. This animal lived from the Late Pliocene (Chapadmalalan) to the Late Pleistocene (Lujanian) in southern South America, being the last survivor of the family Proterotheriidae. Species The genus includes two species, the type species ''N. recens'' and ''N. major''. The fossil found of ''N. major'', one jaw, come from the Miramar Formation in Chapadmalal, Argentina and correspond to the Chapadmalalan mammal age of South America (4.0 to 3.0 million years ago, in the Pliocene). ''N. recens'' appeared in the Ensenadan age (1.2-0.8 million years ago) and the species survived until the Lujanian age (800,000 and 11,000 years ago). Fossils of this species have been found in the Argentine provinces of the northeast, in Córdoba Province, Argentina, Córdoba, Corrientes Province, Corrientes, Tezzanos Pinto Formation, Santa Fe Province, Santa Fe and Santiago del Estero Province, Santiago del E ...
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Neodolodus
''Neodolodus'' is a genus of extinct litoptern from the late middle Miocene of southern Colombia. It was named in 1986 by Hoffstetter and Soria, for the species ''Neodolodus colombianus'' from the Castilletes Formation and the La Victoria and Villavieja Formations of the Honda Group. The type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ... ''N. colombianus'' is known from a right mandibular ramus, teeth, and part of the fore- and hindlimbs. ''Neodolodus'' has been classified as a species of '' Prothoatherium'' or '' Lambdaconus'', but was recognized as a distinct genus by McGrath and colleagues in 2020. The taxon was recovered in a phylogenetic analysis as the sister taxon of '' Protheosodon'', between other proterotheriid genera like '' Picturotherium'' and '' Anisolop ...
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Neobrachytherium
''Neobrachytherium'' is an extinct genus of proterotheriid mammal from the Late Miocene of Argentina and Uruguay. It is represented by multiple species, including the type ''N. intermedium'', originally named in 1891 by Moreno and Mercerat as a species of '' Licaphrium'', ''N. morenoi'', originally named in 1914 by Rovereto as a species of ''Brachytherium'', and ''N. ameghinoi'' and ''N. ullumense'', named in 2001 by Soria, who reclassified all the species in the new genus ''Neobrachytherium''. The various species are known from cranial and dental material from the Corral Quemado, Loma de las Tapias and Ituzaingó Formations. ''Neobrachytherium'' may be closely related to ''Thoatherium ''Thoatherium'' (meaning "active swift-beast") is an extinct genus of litoptern mammals from the Early Miocene of Argentina. Fossils of the genus have been found in the Santa Cruz Formation in Argentina.
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Lambdaconus
''Lambdaconus'' is a genus of proterotheriid from the Late Oligocene to Early Miocene of Argentina. The type species is ''L. suinus'', named in 1897 by Ameghino, with referred species including ''L. lacerum'', named as ''Proterotherium ''Proterotherium'' (meaning "first beast") is an extinct genus of litoptern mammal of the family Proterotheriidae that lived during the Late Miocene of Argentina and Chile. Fossils of this genus have been found in the Ituzaingó Formation of Arge ... lacerum'' in 1902 by Ameghino, and ''L. inaqeuifacies''. References {{Taxonbar, from=Q111838995 Proterotheriids Oligocene mammals of South America Miocene mammals of South America Paleogene Argentina Neogene Argentina Fossils of Argentina Colhuehuapian Deseadan Fossil taxa described in 1897 Taxa named by Florentino Ameghino Prehistoric placental genera Golfo San Jorge Basin Sarmiento Formation ...
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Guilielmofloweria
''Guilielmofloweria'' is an extinct genus of proterotheriid litoptern that lived from the Middle to Late Eocene of what is now Argentina. Fossils have been found in the Sarmiento Formation of Argentina. Taxonomy ''Guilielmofloweria'' was first named in 1901 by Florentino Ameghino based on fragmentary remains found in the Sarmiento Formation of Argentina, in rocks dating back to the Middle Eocene. He originally considered it a member of the pantodont family Pantolambdidae, but recent studies have shown this to be incorrect, as Richard Cifelli considered it as a member of the litoptern family Proterotheriidae, more specifically within the subfamily Anisolambdinae. Anisolambdinae (also called Anisolambdidae in some studies) was proposed to unite the primitive and earlier forms '' Anisolambda'', '' Eolicaphrium'', '' Heteroglyphis'', '' Lambdaconops'', '' Paranisolambda'', '' Protheosodon'', '' Wainka'' and '' Xesmodon''. However, the phylogenetic analysis of McGrath and colleagu ...
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Epitherium
''Epitherium'' is an extinct genus of Litopterna, who belonged to the family Proterotheriidae. It lived during the Pliocene in South America. The fossils of this herbivorous ungulate were found in Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ....The Paleobiology Database


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Proterotheriids
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Eoauchenia
''Eoauchenia'' is a genus of extinct proterotheriidae, proterotheriid from the Late Miocene and Early Pliocene of Argentina. The genus was named by Ameghino in 1887 for the type species ''E. primitiva'', which was originally known from the early Pliocene Monte Hermoso Formation of the Montehermosan age of Buenos Aires Province, but has since been found in the Cerro Azul Formation of the late Huayquerian of La Pampa Province. The fossils originally known included postcranial remains like foot material, but teeth, a skull and skeleton, and mandibles and further postcrania have since been referred to ''Eoauchenia'', which can be distinguished from ''Epitherium'' found alongside it by more gracile bones. References

{{Taxonbar, from=Q112077467 Proterotheriids Miocene mammals of South America Neogene Argentina Fossils of Argentina Fossil taxa described in 1887 Taxa named by Florentino Ameghino Cerro Azul Formation ...
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Diplasiotherium
''Diplasiotherium'' is an extinct genus of litoptern belonging to the family Proterotheriidae, that lived between the late Miocene and the early Pliocene (in the SALMAs Huayquerian and Montehermosan).Villafañé et al., 2006, p.161 The fossils of this animal have been found in Argentina, in the Monte Hermoso Formation.Deschamps & Tomassini, 2016, p.224 ''Diplasiotherium'' is distinguished from other proterotheriids by the crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ... of its molars, which were higher than in other related genera (protohypsodont), and by its larger body size; the species ''D. robustum'' reached approximately in weight, making it the largest known proterotheriid.Villafañe, A. L. (2005). ''Paleoecología de los Proterotheriidae (Mammalia, Litopterna): U ...
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Brachytherium
''Brachytherium'' is an extinct genus of proterotheriid mammal from the Late Miocene to Late Pliocene of Argentina. It is represented by the type and only species ''B. cuspidatum'', a taxon named in 1883 by Ameghino for a partial mandible with teeth. Though it was considered a dubious taxon at times, ''Brachytherium'' was revised as valid by Schmidt in 2015, who also synonymized the species ''Proterotherium gradatum'' and ''Lophogonodon paranensis'', expanding the material known, all of which is from the Ituzaingó Formation The Ituzaingó Formation ( es, Formación Ituzaingó), in older literature also described as Entre Ríos or Entrerriana Formation, is an extensive geological formation of Late Miocene (Tortonian, or Huayquerian in the SALMA classification) age in .... Some material previously referred to ''Brachytherium'' has been given the new name '' Neobrachytherium''. References {{Taxonbar, from=Q111828098 Proterotheriids Miocene mammals of South America Pliocen ...
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Anisolambda
''Anisolambda'' is an extinct genus of litoptern. It lived from the Late Paleocene to the Middle Eocene in what is now Argentina. Description This animal is mostly known from fossils of its maxilla, mandible and teeth, and it is therefore difficult to speculate on its appearance. Its molars were primitive in shape, and closely resembled those of the enigmatic '' Didolodus''. They were distinguished from the latter by the presence of a strong paraconid, in an internal position, almost identical in size to the metaconid, and separated from the latter by a narrow indentation. ''Anisolambda'' may have been similar to more recent genera of Proterotheriidae, such as '' Diadiaphorus'' or ''Proterotherium'', but without the characteristic limb specializations of the latter genera. Classification The genus ''Anisolambda'' was first described in 1901 by Florentino Ameghino, based on a mandible with teeth from the Eocene of Argentina. Ameghino latter described fossils of the maxilla, tha ...
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Diadiaphorus
''Diadiaphorus'' is an extinct genus of litoptern mammal from the Miocene of Argentina ( Ituzaingó, Pinturas, Chiquimil and Santa Cruz Formations) and Bolivia ( Nazareno Formation), South America. Description ''Diadiaphorus'' closely resembled a horse, but was only around in body length with a weight , similar to a modern sheep.D. Patterson, Bruce (2012) ''Bones, Clones, and Biomes: The History and Geography of Recent Neotropical Mammals'' p.92 It had three toes, only one of which touched the ground. This toe had a large hoof; the two outer toes were rudimentary, much like those of early horses such as ''Merychippus''. Unlike horses, however, ''Diadiaphorus'' lacked fused limb bones. Its skull was short and had a relatively large brain cavity. Judging from its low molars The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from La ...
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