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Tyrannosaurid
Tyrannosauridae (or tyrannosaurids, meaning "tyrant lizards") is a family of coelurosaurian theropod dinosaurs that comprises two subfamilies containing up to thirteen genera, including the eponymous ''Tyrannosaurus''. The exact number of genera is controversial, with some experts recognizing as few as three. All of these animals lived near the end of the Cretaceous Period and their fossils have been found only in North America and Asia. Although descended from smaller ancestors, tyrannosaurids were almost always the largest predators in their respective ecosystems, putting them at the apex of the food chain. The largest species was ''Tyrannosaurus rex'', one of the largest and most massive known land predators, which measured over in length and according to most modern estimates to in weight. Tyrannosaurids were bipedal carnivores with massive skulls filled with large teeth. Despite their large size, their legs were long and proportioned for fast movement. In contrast, their ...
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Tyrannosaurus
''Tyrannosaurus'' is a genus of large theropoda, theropod dinosaur. The species ''Tyrannosaurus rex'' (''rex'' meaning "king" in Latin), often called ''T. rex'' or colloquially ''T-Rex'', is one of the best represented theropods. ''Tyrannosaurus'' lived throughout what is now western North America, on what was then an island continent known as Laramidia. ''Tyrannosaurus'' had a much wider range than other Tyrannosauridae, tyrannosaurids. Fossils are found in a variety of geologic formation, rock formations dating to the Maastrichtian Age (geology), age of the Upper Cretaceous Period (geology), period, 68 to 66 mya (unit), million years ago. It was the last known member of the tyrannosaurids and among the last non-aves, avian dinosaurs to exist before the Cretaceous–Paleogene extinction event. Like other tyrannosaurids, ''Tyrannosaurus'' was a bipedal carnivore with a massive skull balanced by a long, heavy tail. Relative to its large and powerful hind limbs, the foreli ...
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Tyrannosaurus Rex
''Tyrannosaurus'' is a genus of large theropod dinosaur. The species ''Tyrannosaurus rex'' (''rex'' meaning "king" in Latin), often called ''T. rex'' or colloquially ''T-Rex'', is one of the best represented theropods. ''Tyrannosaurus'' lived throughout what is now western North America, on what was then an island continent known as Laramidia. ''Tyrannosaurus'' had a much wider range than other tyrannosaurids. Fossils are found in a variety of rock formations dating to the Maastrichtian age of the Upper Cretaceous period, 68 to 66 million years ago. It was the last known member of the tyrannosaurids and among the last non-avian dinosaurs to exist before the Cretaceous–Paleogene extinction event. Like other tyrannosaurids, ''Tyrannosaurus'' was a bipedal carnivore with a massive skull balanced by a long, heavy tail. Relative to its large and powerful hind limbs, the forelimbs of ''Tyrannosaurus'' were short but unusually powerful for their size, and they had two clawed ...
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Daspletosaurus
''Daspletosaurus'' ( ; meaning "frightful lizard") is a genus of tyrannosaurid dinosaur that lived in Laramidia between about 79.5 and 74 million years ago, during the Late Cretaceous Period. The genus ''Daspletosaurus'' contains three species. Fossils of the earlier type species, ''D. torosus'', have been found in Alberta, and fossils of a later second species, ''D. wilsoni,'' and third species, ''D. horneri'', have been found only in Montana. A possible fourth species, also from Alberta, awaits formal identification. The taxon ''Thanatotheristes'' has been suggested to represent a species of ''Daspletosaurus'', ''D. degrootorum'', but this has not been widely supported. ''Daspletosaurus'' is closely related to the much larger and more recent tyrannosaurid ''Tyrannosaurus rex''. Like most tyrannosaurids, ''Daspletosaurus'' was a multi-tonne bipedal predator equipped with dozens of large, sharp teeth. ''Daspletosaurus'' had the small forelimbs typical of tyrannosaurids, although ...
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Gorgosaurus
''Gorgosaurus'' ( ; ) is a genus of tyrannosaurid theropod dinosaur that lived in western North America during the Late Cretaceous Period (Campanian), between about 76.6 and 75.1 million years ago. Fossil remains have been found in the Canadian province of Alberta and the U.S. state of Montana. Paleontologists recognize only the type species, ''G. libratus'', although other species have been erroneously referred to the genus. Like most known tyrannosaurids, ''Gorgosaurus'' was a large bipedal predator, measuring in length and in body mass. Dozens of large, sharp teeth lined its jaws, while its two-fingered forelimbs were comparatively small. ''Gorgosaurus'' was most closely related to ''Albertosaurus'', and more distantly related to the larger ''Tyrannosaurus''. ''Gorgosaurus'' and ''Albertosaurus'' are extremely similar, distinguished mainly by subtle differences in the teeth and skull bones. Some experts consider ''G. libratus'' to be a species of ''Albertosaurus''; thi ...
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Tarbosaurus
''Tarbosaurus'' ( ; meaning "alarming lizard") is a genus of tyrannosaurid dinosaur that flourished in Asia about 70 million years ago, at the end of the Late Cretaceous Period, considered to contain a single known species, ''Tarbosaurus bataar''. Fossils have been recovered from the Nemegt Formation of Mongolia, with more fragmentary remains found further afield in the Subashi Formation of China. Although many species have been named, modern paleontologists recognize only one, ''T. bataar'', as valid. Some experts see this species as an Asian representative of the North American genus ''Tyrannosaurus''; this would make the genus ''Tarbosaurus'' redundant. ''Tarbosaurus'' and ''Tyrannosaurus'', if not synonymous, are considered to be at least closely related genera. ''Alioramus'', also from Mongolia, has previously been thought by some authorities to be the closest relative of ''Tarbosaurus'', though this has since been disproven with the discovery of '' Qianzhousaurus'' and ...
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Albertosaurinae
Albertosaurines, or dinosaurs of the subfamily Albertosaurinae, lived in the Late Cretaceous of United States and Canada. The subfamily was first used by Philip J. Currie, Jørn H. Hurum and Karol Sabath as a group of tyrannosaurid dinosaurs. It was originally defined as "(''Albertosaurus'' + ''Gorgosaurus'')", including only the two genera. The group is sister clade to Tyrannosaurinae. In 2007, it was found that the group also contained '' Maleevosaurus'', often synonymized with ''Tarbosaurus''. However, this classification has not been accepted, and '' Maleevosaurus'' is still considered a juvenile ''Tarbosaurus'' or ''Tyrannosaurus''. Description Albertosaurines are large, lightly built tyrannosaurids. Compared to tyrannosaurines, they are lightly built, have shorter, flatter skulls, had shorter ilia, and had proportionally longer tibiae. Albertosaurines and tyrannosaurines share arms or about equal length, with the exception of ''Tarbosaurus'', which had sh ...
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Alioramus
''Alioramus'' (; meaning 'different branch') is a genus of tyrannosaurid theropod dinosaurs from the Late Cretaceous period of Asia. It currently contains two species. The type species, ''A. remotus'' is known from a partial skull and three foot bones recovered from the Mongolian Nemegt Formation, which was deposited in a humid floodplain about 70 million years ago. These remains were named and described by Soviet paleontologist Sergei Kurzanov in 1976. A second species, ''A. altai'', known from a much more complete skeleton also from the Nemegt Formation, was named and described by Stephen L. Brusatte and colleagues in 2009. Its relationships to other tyrannosaurid genera were at first unclear, with some evidence supporting a hypothesis that ''Alioramus'' was closely related to the contemporary species ''Tarbosaurus bataar''. However, the discovery of ''Qianzhousaurus'' indicates that it belongs to a distinct branch of tyrannosaurs, namely the tribe Alioramini. ''Alioramu ...
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Tyrannosauroidea
Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent beginning in the Jurassic Period. By the end of the Cretaceous Period, tyrannosauroids were the dominant large predators in the Northern Hemisphere, culminating in the gigantic ''Tyrannosaurus''. Fossils of tyrannosauroids have been recovered on what are now the continents of North America, Europe and Asia, with fragmentary remains possibly attributable to tyrannosaurs also known from South America and Australia. Tyrannosauroids were bipedal carnivores, as were most theropods, and were characterized by numerous synapomorphy, skeletal features, especially of the skull and pelvis. Early in their existence, tyrannosauroids were small predators with long, three-fingered forelimbs. Late Cretaceous genera became much larger, including some of the ...
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Tyrannosaurinae
The tyrannosaurines (Tyrannosaurinae) is an extinct subfamily of Tyrannosauridae a family of coelurosaurian theropod dinosaurs that consists of at least three tribes and several genera. All fossils of these genera have been found in the Late Cretaceous deposits of western North America and east Asia. Compare to the related subfamily Albertosaurinae, tyrannosaurines overall are more robust and larger though the alioramins were gracile by comparison. This subfamily also includes the oldest known tyrannosaurid genus ''Lythronax'' as well as the youngest and most famous member of the group, ''Tyrannosaurus rex''. History of discovery The first remains of tyrannosaurids were uncovered during expeditions led by the Geological Survey of Canada, which located numerous scattered teeth. These distinctive dinosaur teeth were given the name ''Deinodon'' ("terrible tooth") by Joseph Leidy in 1856. In 1892 Edward Drinker Cope described more tyrannosaur material in the form of isolated vertebra ...
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Raptorex
''Raptorex'' is a dubious genus of tyrannosaurid dinosaur. Its fossil remains consist of a single juvenile specimen probably uncovered in Mongolia, or possibly northeastern China. The type species is ''R. kriegsteini'', described in 2009 by Sereno and colleagues. The genus name is derived from Latin ', "robber", and ', "king". The specific name honours Roman Kriegstein, a survivor of the Holocaust, whose son Henry Kriegstein donated the specimen to the University of Chicago for scientific study. While initially considered to have come from the Yixian Formation of China, dated to approximately 125 million years ago during the early Cretaceous period, later studies showed that such an early date for the fossil are unlikely, and given its extremely close similarity to juvenile tyrannosaurids of the late Cretaceous, it probably came from the Nemegt or similar formation. Because the specimen is a juvenile, and the changes undergone by tyrannosaurids during growth are not yet well u ...
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Zhuchengtyrannus
''Zhuchengtyrannus'' (meaning "Zhucheng tyrant") is a genus of tyrannosaurid theropod dinosaur known from the Campanian stage of the Late Cretaceous of Shandong Province, China. It belongs to the subfamily Tyrannosaurinae, and contains a single species, ''Zhuchengtyrannus magnus''. Discovery and naming ''Zhuchengtyrannus'' was first described and named by David W. E. Hone, Kebai Wang, Corwin Sullivan, Xijin Zhao, Shuqing Chen, Dunjin Li, Shuan Ji, Qiang Ji and Xing Xu in 2011 and the type species is ''Zhuchengtyrannus magnus''. The generic name is derived from the word ''Zhucheng'', which refers to the type locality, and ''tyrant'' in reference to its phylogenetic position as a tyrannosaurid. The specific name ''magnus'' meaning "great" in Latin refers to the relatively large size of ''Zhuchengtyrannus''. ''Zhuchengtyrannus'' is known solely from the holotype ZCDM V0031, a nearly complete right maxilla and associated left dentary (lower jaw, both with teeth) discovered around 2 ...
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Aublysodon
''Aublysodon'' (“backwards-flowing tooth") is a genus of carnivorous dinosaurs known only from the Judith River Formation in Montana, which has been dated to the late Campanian age of the late Cretaceous period (about 75 million years ago). The only currently recognized species, ''Aublysodon mirandus'', was named by paleontologist Joseph Leidy in 1868. It is now considered dubious, because the type specimen consists only of an isolated premaxillary (front) tooth. Although this specimen is now lost, similar teeth have been found in many US states, western Canada, and Asia. These teeth almost certainly belong to juvenile tyrannosaurine tyrannosaurids, but most have not been identified to species level. However, it is likely that the type tooth (and therefore the name ''Aublysodon mirandus'' itself) belongs to one of the species in the genus ''Daspletosaurus'', which was present in contemporary formations, and which matches specific details of the original tooth. The synapomorphies ...
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