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Thalattosaur
Thalattosauria (Greek for "sea lizards") is an extinct order of prehistoric marine reptiles that lived in the middle to late Triassic period. Thalattosaurs were diverse in size and shape, and are divided into two superfamilies: Askeptosauroidea and Thalattosauroidea. Askeptosauroids were endemic to the Tethys Ocean, their fossils have been found in Europe and China, and they were likely semiaquatic fish eaters with straight snouts and decent terrestrial abilities. Thalattosauroids were more specialized for aquatic life and most had unusual downturned snouts and crushing dentition. Thalattosauroids lived along the coasts of both Panthalassa and the Tethys Ocean, and were most diverse in China and western North America. The largest species of thalattosaurs grew to over 4 meters (13 feet) in length, including a long, flattened tail utilized in underwater propulsion. Although thalattosaurs bore a superficial resemblance to lizards, their exact relationships are unresolved. They ar ...
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Thalattosaurus Alexandrae
''Thalattosaurus'' (pronounced: , "tha-la-to-SORE-us") meaning "sea lizard," from the Attic Greek ' (), "sea," and ' (), "lizard," is an extinct genus of marine reptile in the family Thalattosauroidea. They were aquatic diapsids that are known exclusively from the Triassic period. It was a long shellfish-eating reptile with paddle-like limbs and a down-turned rostrum occurring in the Lower and Middle Triassic Sulphur Mountain Formation of British Columbia as well as the Upper Triassic Hosselkus Limestone of California. It has gained notoriety as a result of studies on general diapsid phylogeny. Although originally described as four distinct species by Merriam in 1905, one was proven to be ''T. alexandrae'' upon further inspection and another has a missing type specimen. Currently it is believed to include two known species; ''Thalattosaurus alexandrae'' and ''T. borealis''. Discovery and naming In the summer of 1903 Annie Alexander led an expedition with Miss Edna Wemple ...
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Thalattosauroidea
Thalattosauroidea is a superfamily of thalattosaurs, a Triassic group of marine reptiles. It was named in 1904 by paleontologist John Campbell Merriam to include the genus '' Thalattosaurus'' from California. Thalattosauroids are one of two groups of Thalattosauria, the other being Askeptosauroidea. Thalattosauroids make up the "traditional" thalattosaurs with large downturned snouts, short necks, and long, paddle-like tails. Classification Thalattosauria includes North American genera such as ''Thalattosaurus'' and ''Nectosaurus'' as well as recently described Chinese forms such as '' Xinpusaurus''. A 1999 study of thalattosaurs, which established much of the currently accepted phylogeny of the group, referred to Thalattosauroidea as Thalattosauria, while calling the larger group Thalattosauriformes. More recent phylogenetic studies have come upon the same conclusions, but refer to the group as Thalattosauroidea in order to contrast it with another superfamily of thalattosaurs ...
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Gunakadeit Joseeae
''Gunakadeit'' is an extinct genus of thalattosaur. It is known from a single species, ''Gunakadeit joseeae'', which is based on an articulated and mostly complete skeleton from the late Triassic (middle Norian) Hound Island Volcanics of Alaska. ''Gunakadeit'' possessed a variety of features from the two major suborders of thalattosaurs, Askeptosauroidea and Thalattosauroidea, and it is considered the most basal member of the latter group. Despite this, it is also the youngest known thalattosaur genus, with the group going extinct at the end of the Triassic. ''Gunakadeits basal position and relatively recent occurrence implies a 20-million-year ghost lineage connecting it to the rest of Thalattosauria. The skull ends in a sharply pointed and toothless tip like the askeptosauroid ''Endennasaurus'', but unlike ''Endennasaurus'', ''Gunakadeit'' had poorly developed joints and was likely exclusively aquatic in behavior. Discovery The holotype, UAMES 23258, was discovered by r ...
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Askeptosauroidea
Askeptosauroidea is a superfamily of thalattosaurs, a Triassic group of marine reptiles. Askeptosauroidea is one of two major subgroups of Thalattosauria, the other being Thalattosauroidea. It includes the family Askeptosauridae and a more basal form called ''Endennasaurus''. Phylogeny Below is a cladogram from Wu ''et al.'' (2009) showing the phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... relationships of Askeptosauroidea: References Thalattosaurs Triassic first appearances Triassic extinctions {{triassic-reptile-stub ...
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Askeptosaurus Italicus
''Askeptosaurus'' is an extinct genus of askeptosauroid, a marine reptile from the extinct order Thalattosauria. ''Askeptosaurus'' is known from several well-preserved fossils found in Middle Triassic marine strata in what is now Italy and Switzerland. History of discovery ''Askeptosaurus,'' and its only known species ''Askeptosaurus italicus,'' were first named and described in 1925 by Hungarian paleontologist Franz Nopcsa von Felső-Szilvás. It was most recently redescribed by Dr. Johannes Müller in 2005. ''Askeptosaurus'' is known from several disarticulated and articulated skeletons preserved at the MSNM (Museo Civico di Storia Naturale di Milano) in Milan, Italy, and the PIMUZ ( Paläontologisches Institut und Museum der Universität Zürich) in Zürich, Switzerland. These specimens were discovered in the Grenzbitumenzone of Monte San Giorgio, a UNESCO World Heritage Site on the Swiss-Italian border. Also known as the Besano Formation in Italy, the Grenzbitumenzone has ...
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Diapsid
Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The group first appeared about three hundred million years ago during the late Carboniferous period. All diapsids other than the most primitive ones in the clade Araeoscelidia are sometimes placed into the clade Neodiapsida. The diapsids are extremely diverse, and include birds and all modern reptile groups, including turtles, which were historically thought to lie outside the group. Although some diapsids have lost either one hole (lizards), or both holes (snakes and turtles), or have a heavily restructured skull (modern birds), they are still classified as diapsids based on their ancestry. At least 17,084 species of diapsid animals are extant: 9,159 birds, and 7,925 snakes, lizards, tuatara, turtles, and crocodiles. Characteristics The name Diapsida means "two arches", and diapsids are tradition ...
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Triassic
The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic. The Triassic began in the wake of the Permian–Triassic extinction event, which left the Earth's biosphere impoverished; it was well into the middle of the Triassic before life recovered its former diversity. Three categories of organisms can be distinguished in the Triassic record: survivors from the extinction event, new groups that flourished briefly, and other new groups that went on to dominate the Mesozoic Era. Reptiles, especially archosaurs, were the chief terrestrial vertebrates during this time. A specialized subgroup of archo ...
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Endennasaurus Acutirostris
''Endennasaurus'' is an extinct genus of thalattosaurian from the Upper Triassic of Italy. It was found in and named after the Endenna cave, composed of Zorzino Limestone in Lombardia.''Endennasaurus''
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Fossilworks Fossilworks is a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals ...
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Thalattosaurs
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Blezingeria
''Blezingeria'' is an extinct genus of marine reptile from the Middle Triassic of Germany. The type and only species ''Blezingeria ichthyospondyla'' was named by German paleontologist Eberhard Fraas in 1896. It is known from many isolated bones that come from a deposit in southwestern Germany called the Upper Muschelkalk, which dates back to the Ladinian stage. The relationships of ''Blezingeria'' are uncertain. Fraas identified it as a nothosaur, but it has also been classified as a cymbospondylid ichthyosaur, and most recently a thalattosaur. Many thalattosaur fossils have been found in a slightly olderLate Ladinian-age rock unit in Monte San Giorgio, Switzerland, so if ''Blezingeria'' is a thalattosaur, it may represent an early stage in an evolutionary radiation An evolutionary radiation is an increase in taxonomic diversity that is caused by elevated rates of speciation, that may or may not be associated with an increase in morphological disparity. Radiations may affect ...
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Marine Reptile
Marine reptiles are reptiles which have become secondarily adapted for an aquatic or semiaquatic life in a marine environment. The earliest marine reptile mesosaurus (not to be confused with mosasaurus), arose in the Permian period during the Paleozoic era. During the Mesozoic era, many groups of reptiles became adapted to life in the seas, including such familiar clades as the ichthyosaurs, plesiosaurs (these two orders were once thought united in the group "Enaliosauria", a classification now cladistically obsolete), mosasaurs, nothosaurs, placodonts, sea turtles, thalattosaurs and thalattosuchians. Most marine reptile groups became extinct at the end of the Cretaceous period, but some still existed during the Cenozoic, most importantly the sea turtles. Other Cenozoic marine reptiles included the bothremydids, palaeophiid snakes, a few choristoderes such as ''Simoedosaurus'' and dyrosaurid crocodylomorphs. Various types of marine gavialid crocodilians remained widespread as ...
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Archosauromorpha
Archosauromorpha (Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) rather than lepidosaurs (such as tuataras, lizards, and snakes). Archosauromorphs first appeared during the late Middle Permian or Late Permian, though they became much more common and diverse during the Triassic period. Although Archosauromorpha was first named in 1946, its membership did not become well-established until the 1980s. Currently Archosauromorpha encompasses four main groups of reptiles: the stocky, herbivorous allokotosaurs and rhynchosaurs, the hugely diverse Archosauriformes, and a polyphyletic grouping of various long-necked reptiles including ''Protorosaurus'', tanystropheids, and ''Prolacerta''. Other groups including pantestudines (turtles and their extinct relatives) and the semiaquatic choristoderes have also been placed in Archosauromorpha by some authors. A ...
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Archosaur
Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian dinosaurs, pterosaurs, and extinct relatives of crocodilians. Modern paleontologists define Archosauria as a crown group that includes the most recent common ancestor of living birds and crocodilians, and all of its descendants. The base of Archosauria splits into two clades: Pseudosuchia, which includes crocodilians and their extinct relatives, and Avemetatarsalia, which includes birds and their extinct relatives (such as non-avian dinosaurs and pterosaurs). Older definitions of the group Archosauria rely on shared morphological characteristics, such as an antorbital fenestra in the skull, serrated teeth, and an upright stance. Some extinct reptiles, such as proterosuchids and euparkeriids, possessed these features yet originated pri ...
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