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Tetraconata
Pancrustacea is the clade that comprises all crustaceans and hexapods. This grouping is contrary to the Atelocerata hypothesis, in which Myriapoda and Hexapoda are sister taxa, and Crustacea are only more distantly related. As of 2010, the Pancrustacea taxon is considered well accepted, with most studies recovering Hexapoda within Crustacea. The clade has also been called Tetraconata, referring to having four cone cells in the ommatidia. This name is preferred by some scientists as a means of avoiding confusion with the use of "pan-" to indicate a clade that includes a crown group and all of its stem group representatives. Molecular studies A monophyletic Pancrustacea has been supported by several molecular studies, in most of which the subphylum Crustacea is paraphyletic with regard to hexapods (that is, that hexapods, including insects, are derived from crustacean ancestors). The evidence for this clade derives from molecular data and morphological characteristics. The molecu ...
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Paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of Synapomorphy and apomorphy, synapomorphies and symplesiomorphy, symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia (reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles a ...
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Grapsus Grapsus
''Grapsus grapsus'' is one of the most common crabs along the western coast of the Americas. It is known as the red rock crab, or, along with crabs such as ''Percnon gibbesi'', as the Sally Lightfoot crab. Distribution ''Grapsus grapsus'' is found along the Pacific coast of Mexico, Central America, and South America (as far south as northern Peru), and on nearby islands, including the Galápagos Islands. It is also found along the Atlantic coast of South America, but is replaced in the eastern Atlantic Ocean ( Ascension Island and West Africa) by its congener ''Grapsus adscensionis''. Description ''Grapsus grapsus'' is a typically shaped crab, with five pairs of legs, the front two bearing small, blocky, symmetrical chelae (claws). The other legs are broad and flat, with only the tips touching the substrate. The crab's round, flat carapace is slightly longer than . Young ''G. grapsus'' are black or dark brown in colour and are camouflaged well on the black lava coasts of vo ...
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Monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have tak ...
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Remipedia
Remipedia is a class of blind crustaceans found in coastal aquifers which contain saline groundwater, with populations identified in almost every ocean basin so far explored, including in Australia, the Caribbean Sea, and the Atlantic Ocean. The first described remipede was the fossil ''Tesnusocaris goldichi'' (Early Pennsylvanian, Lower Pennsylvanian). Since 1979, at least seventeen living species have been identified in subtropical regions around the world. Description Remipedes are long and comprise a head and an elongate trunk of up to thirty-two similar body segments. Pigmentation and eyes are absent. Biramous swimming appendages are laterally present on each segment. The animals swim on their backs and are generally slow-moving. They are the only known venomous crustaceans, and have fangs connected to secretory glands, which inject a combination of digestive enzymes and venom into their prey, but they also feed through Filter feeder, filter feeding. Being hermaphrodites, th ...
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Cephalocarida
The Cephalocarida are a class in the subphylum Crustacea comprising only 12 benthic species. They were discovered in 1955 by Howard L. Sanders, and are commonly referred to as horseshoe shrimp. They have been grouped together with the Remipedia in the Xenocarida. Although a second family, Lightiellidae, is sometimes used, all cephalocaridans are generally considered to belong in just one family: Hutchinsoniellidae. Though no fossil record of cephalocaridans has been found, most specialists believe them to be primitive among crustaceans. Taxonomy * Class Cephalocarida Sanders 1955 ** Order Brachypoda Birshteyn 1960 *** Family Hutchinsoniellidae Sanders 1955 **** Genus '' Chiltoniella'' Knox & Fenwick 1977 ***** ''Chiltoniella elongata'' Knox & Fenwick 1977 **** Genus '' Hampsonellus'' Hessler & Wakabara 2000 ***** ''Hampsonellus brasiliensis'' Hessler & Wakabara 2000 **** Genus '' Hutchinsoniella'' Sanders 1955 ***** ''Hutchinsoniella macracantha'' Sanders 1955 **** Genus '' Li ...
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Branchiopoda
Branchiopoda is a class of crustaceans. It comprises fairy shrimp, clam shrimp, Diplostraca (or Cladocera), Notostraca and the Devonian ''Lepidocaris''. They are mostly small, freshwater animals that feed on plankton and detritus. Description Members of the Branchiopoda are unified by the presence of gills on many of the animals' appendages, including some of the mouthparts. This is also responsible for the name of the group (from the grc, βράγχια, gills, akin to , windpipe; el, πούς, foot). They generally possess compound eyes and a carapace, which may be a shell of two valves enclosing the trunk (as in most Cladocera), broad and shallow (as in the Notostraca), or entirely absent (as in the Anostraca). In the groups where the carapace prevents the use of the trunk limbs for swimming (Cladocera, clam shrimp and the extinct Lipostraca), the antennae are used for locomotion, as they are in the nauplius. Male fairy shrimp have an enlarged pair of antennae with which ...
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Neuron
A neuron, neurone, or nerve cell is an electrically excitable cell that communicates with other cells via specialized connections called synapses. The neuron is the main component of nervous tissue in all animals except sponges and placozoa. Non-animals like plants and fungi do not have nerve cells. Neurons are typically classified into three types based on their function. Sensory neurons respond to stimuli such as touch, sound, or light that affect the cells of the sensory organs, and they send signals to the spinal cord or brain. Motor neurons receive signals from the brain and spinal cord to control everything from muscle contractions to glandular output. Interneurons connect neurons to other neurons within the same region of the brain or spinal cord. When multiple neurons are connected together, they form what is called a neural circuit. A typical neuron consists of a cell body (soma), dendrites, and a single axon. The soma is a compact structure, and the axon and dend ...
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Axonogenesis
Axon guidance (also called axon pathfinding) is a subfield of neural development concerning the process by which neurons send out axons to reach their correct targets. Axons often follow very precise paths in the nervous system, and how they manage to find their way so accurately is an area of ongoing research. Axon growth takes place from a region called the growth cone and reaching the axon target is accomplished with relatively few guidance molecules. Growth cone receptors respond to the guidance cues. Mechanisms Growing axons have a highly motile structure at the growing tip called the growth cone, which responds to signals in the extracellular environment that instruct the axon in which direction to grow. These signals, called guidance cues, can be fixed in place or diffusible; they can attract or repel axons. Growth cones contain receptors that recognize these guidance cues and interpret the signal into a chemotropic response. The general theoretical framework is that whe ...
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Neuroblasts
In vertebrates, a neuroblast or primitive nerve cell is a postmitotic cell that does not divide further, and which will develop into a neuron after a migration phase. In invertebrates such as ''Drosophila,'' neuroblasts are neural progenitor cells which divide asymmetrically to produce a neuroblast, and a daughter cell of varying potency depending on the type of neuroblast. Vertebrate neuroblasts differentiate from radial glial cells and are committed to becoming neurons. Neural stem cells, which only divide symmetrically to produce more neural stem cells, transition gradually into radial glial cells. Radial glial cells, also called radial glial progenitor cells, divide asymmetrically to produce a neuroblast and another radial glial cell that will re-enter the cell cycle. This mitosis occurs in the germinal neuroepithelium (or germinal zone), when a radial glial cell divides to produce the neuroblast. The neuroblast detaches from the epithelium and migrates while the radial glia ...
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Arthropod Eye
Apposition eyes are the most common form of eye, and are presumably the ancestral form of compound eye. They are found in all arthropod groups, although they may have evolved more than once within this phylum. Some annelids and bivalves also have apposition eyes. They are also possessed by ''Limulus'', the horseshoe crab, and there are suggestions that other chelicerates developed their simple eyes by reduction from a compound starting point. Some caterpillars appear to have evolved compound eyes from simple eyes in the opposite fashion. The arthropods ancestrally possessed compound eyes, but the type and origin of this eye varies between groups, and some taxa have secondarily developed simple eyes. The organ's development through the lineage can be estimated by comparing groups that branched early, such as the velvet worm and horseshoe crab to the advanced eye condition found in insects and other derived arthropods. Eyes and functions Most arthropods have at least one of ...
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Protein
Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, responding to stimuli, providing structure to cells and organisms, and transporting molecules from one location to another. Proteins differ from one another primarily in their sequence of amino acids, which is dictated by the nucleotide sequence of their genes, and which usually results in protein folding into a specific 3D structure that determines its activity. A linear chain of amino acid residues is called a polypeptide. A protein contains at least one long polypeptide. Short polypeptides, containing less than 20–30 residues, are rarely considered to be proteins and are commonly called peptides. The individual amino acid residues are bonded together by peptide bonds and adjacent amino acid residues. The sequence of amino acid residue ...
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Mitochondria
A mitochondrion (; ) is an organelle found in the Cell (biology), cells of most Eukaryotes, such as animals, plants and Fungus, fungi. Mitochondria have a double lipid bilayer, membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is used throughout the cell as a source of chemical energy. They were discovered by Albert von Kölliker in 1857 in the voluntary muscles of insects. The term ''mitochondrion'' was coined by Carl Benda in 1898. The mitochondrion is popularly nicknamed the "powerhouse of the cell", a phrase coined by Philip Siekevitz in a 1957 article of the same name. Some cells in some multicellular organisms lack mitochondria (for example, mature mammalian red blood cells). A large number of unicellular organisms, such as microsporidia, parabasalids and diplomonads, have reduced or transformed their mitochondria into mitosome, other structures. One eukaryote, ''Monocercomonoides'', is known to have completely lost its mitocho ...
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