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Tapejaromorph
Tapejaromorpha is a group of pterosaurs within the clade Azhdarchoidea. The fossil remains of tapejaromorphs dated back to the Cretaceous period. The Tapejaromorpha was defined in 2014 by Andres and colleagues. They made Tapejaromorpha the most inclusive clade containing ''Tapejara'' but not ''Quetzalcoatlus''. Martill and Naish had previously proposed this same definition for Tapejaridae Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primiti ..., but did not formalize it. Andres and colleagues instead defined Tapejaridae more narrowly, as the clade ''Tapejara'' + ''Sinopterus''. References {{pterosaur-stub ...
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Huaxiapterus Jii
''Huaxiapterus'' (meaning " Hua Xia hinawing") is a genus of tapejarid pterodactyloid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. It is the second genus of tapejarid from this formation, after ''Sinopterus''. Three species are known (''H. jii'', "''H. corollatus''", and "''H. benxiensis''"), though they may not actually form a natural group with each other. It was first named by Lü Junchang and Yuan Chongxi. Discovery The type species of ''Huaxiapterus'' is ''H. jii'', which is based on GMN-03-11-001, a nearly complete skeleton and skull. While it was initially separated from the closely related ''Sinopterus'' by several characters including its larger size and larger crest, later studies showed that it was in fact more closely related to ''Sinopterus dongi'' than to either of the other two species of ''Huaxiapterus''. Some researchers have considered ''H. jii'' to be a species of ''Sinopterus'' for this reason, thoug ...
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Sinopterus Dongi
''Sinopterus'' (meaning "Chinese wing") was a genus of tapejarid pterodactyloid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. It was first described and named by Wang Xiaolin and Zhou Zhonghe. Three species have been classified in this genus, though only two are generally considered to be valid. ''Sinopterus'' is known for its proportionally large skull, which has a birdlike pointed beak, a long bony crest that starts with a tall premaxilla and goes back along the middle of the skull to form a point overhanging the rear of the skull, and its lack of teeth. Description The type species, ''S. dongi'', is based on IVPP V13363, an articulated, nearly complete skeleton. The skull of this individual was 17 centimeters (6.7 inches) long, and the wingspan was estimated to be 1.2 meters (3.9 feet). The authors suggested that it was an omnivore, and noted that it was the first record of a tapejarid outside of Brazil, and the earlies ...
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Keresdrakon
''Keresdrakon'' is an extinct genus of tapejaromorph pterosaur from the Goio-Erê Formation of Brazil, which dates back to the Late Cretaceous period (Turonian-Campanian stages), 94 to 71 million years ago. ''Keresdrakon'' contains a single species, ''Keresdrakon vilsoni''. Discovery In 1971, Alexandre Dobruski and his son João Gustavo Dobruski, discovered a fossil site near Cruzeiro do Oeste in Paraná. Only in 2011, paleontologists Paulo César Manzig and Luiz C. Weinschütz visited the location. A bone bed proved to be present with hundreds of specimens of a pterosaur that in 2014 was named ''Caiuajara''. Among them were some bones belonging to a second pterosaur species. These remains, both of ''Caiuajara'' and the new taxon, were prepared by volunteer Vilson Greinert. In 2019, the type species ''Keresdrakon vilsoni'' was named and described by Alexander Wilhelm Armin Kellner, Luiz Carlos Weinschütz, Borja Holgado, Renan Alfredo Machado Bantim and Juliana Manso Sayão. ...
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Azhdarchoidea
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early Cretaceous, Early and Late Cretaceous periods, with one tentative member, ''Tendaguripterus'', that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are ''Quetzalcoatlus'', ''Hatzegopteryx'', and ''Arambourgiania''. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea. Classification Azhdarchoidea was given a phylogenetic definition by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of ''Quetzalcoatlus'' and ''Tapejara (pterosaur), Tapejara'', and all its descendants.Unwin, D. M., (20 ...
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Neoazhdarchia
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, ''Tendaguripterus'', that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are ''Quetzalcoatlus'', ''Hatzegopteryx'', and ''Arambourgiania''. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea. Classification Azhdarchoidea was given a phylogenetic definition by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of ''Quetzalcoatlus'' and '' Tapejara'', and all its descendants.Unwin, D. M., (2003). "On the phylogeny and evolutionar ...
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Sinopterus
''Sinopterus'' (meaning "Chinese wing") was a genus of tapejarid pterodactyloid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. It was first described and named by Wang Xiaolin and Zhou Zhonghe. Three species have been classified in this genus, though only two are generally considered to be valid. ''Sinopterus'' is known for its proportionally large skull, which has a birdlike pointed beak, a long bony crest that starts with a tall premaxilla and goes back along the middle of the skull to form a point overhanging the rear of the skull, and its lack of teeth. Description The type species, ''S. dongi'', is based on IVPP V13363, an articulated, nearly complete skeleton. The skull of this individual was 17 centimeters (6.7 inches) long, and the wingspan was estimated to be 1.2 meters (3.9 feet). The authors suggested that it was an omnivore, and noted that it was the first record of a tapejarid outside of Brazil, and the earlies ...
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Eopteranodon Lii
''Eopteranodon'' (meaning "dawn ''Pteranodon'' (toothless wing)") is a genus of tapejarid pterosaur from the Aptian-age Lower Cretaceous Yixian Formation of Beipiao City, Liaoning, China. The genus was named in 2005 by paleontologists Lü Junchang and Zhang Xingliao. The type and only species is ''Eopteranodon lii''. Description and classification ''Eopteranodon'' is based on the type specimen or holotype BPV-078, an incomplete skeleton and skull. Its skull, including a large crest, was toothless and similar to that of ''Pteranodon''. The skull lacks the point of the snout but it was in life less than 200 millimeters long (7.9 inches), and the animal had a wingspan of about 1.1 meters (3.6 feet). A second specimen, D2526, described in 2006, had a larger wingspan. Despite its similarities to ''Pteranodon'', ''Eopteranodon'' was not placed into a family by its describers, who put it into the clade Pteranodontia as ''incertae sedis'' (uncertain position). Shortly ...
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Bennettazhia
''Bennettazhia'' is a genus of tapejaromorph pterosaur from the Early Cretaceous period of what is now the Hudspeth Formation of the state of Oregon in the United States. Although originally identified as a species of the pteranodontoid pterosaur ''Pteranodon'', ''Bennettazhia'' is now thought to have been a different animal. The type and only species is ''B. oregonensis''. Discovery and history In 1928, Charles Gilmore named a new species of ''Pteranodon'': ''P. oregonensis''. A humerus (holotype MPUC V.126713), two fused dorsal vertebrae and the broken-off end of some joint bone had been unearthed from the Lower Cretaceous (Albian stage) beds of the Hudspeth Formation in Wheeler County, in the state of Oregon, United States, to which the specific epithet refers. Gilmore noted similarities to ''Nyctosaurus'' though the specimens were larger. In 1989, S. Christopher Bennett concluded that the remains might be those of a member of the Azhdarchidae instead of a pteranodont ...
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Tapejaridae
Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primitive genera were found in China, indicating that the family has an Asian origin. Description Tapejarids were small to medium-sized pterosaurs with several unique, shared characteristics, mainly relating to the skull. Most tapejarids possessed a bony crest arising from the snout (formed mostly by the premaxillary bones of the upper jaw tip). In some species, this bony crest is known to have supported an even larger crest of softer, fibrous tissue that extends back along the skull. Tapejarids are also characterized by their large nasoantorbital fenestra, the main opening in the skull in front of the eyes, which spans at least half the length of the entire skull in this family. Their eye sockets were small and pear-shaped. Studies of tapejarid ...
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Bennettazhia Oregonensis
''Bennettazhia'' is a genus of tapejaromorph pterosaur from the Early Cretaceous period of what is now the Hudspeth Formation of the state of Oregon in the United States. Although originally identified as a species of the pteranodontoid pterosaur ''Pteranodon'', ''Bennettazhia'' is now thought to have been a different animal. The type and only species is ''B. oregonensis''. Discovery and history In 1928, Charles Gilmore named a new species of ''Pteranodon'': ''P. oregonensis''. A humerus (holotype MPUC V.126713), two fused dorsal vertebrae and the broken-off end of some joint bone had been unearthed from the Lower Cretaceous ( Albian stage) beds of the Hudspeth Formation in Wheeler County, in the state of Oregon, United States, to which the specific epithet refers. Gilmore noted similarities to '' Nyctosaurus'' though the specimens were larger. In 1989, S. Christopher Bennett concluded that the remains might be those of a member of the Azhdarchidae instead of a pteranodontid. ...
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Thalassodrominae
Thalassodrominae or Thalassodromidae (meaning "sea runners", due to previous misconceptions of skimming behavior; they are now thought to be terrestrial predators) is a group of azhdarchoid pterosaurs from the Cretaceous period. Its traditional members come from Brazil, however, other possible members also come from other places, including the United States, Morocco, and Argentina. Thalassodrominae is considered either to be a subfamily within the pterosaur family Tapejaridae, or as a distinct family, Thalassodromidae, within the clade Neoazhdarchia, closely related to dsungaripterids or azhdarchids. Classification Thalassodrominae traditionally includes only two genera, ''Thalassodromeus'' and ''Tupuxuara'', and was defined to include them and any other descendants of their most recent common ancestor. The classification of thalassodromines is controversial. Some studies, including one by Lü and colleagues in 2008, have found that the thalassodromines are more closely related t ...
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Dsungaripteridae
Dsungaripteridae is a group of pterosaurs within the suborder Pterodactyloidea. They were robust pterosaurs with good terrestrial abilities and flight honed for inland settings. Classification In 1964 Young created a family to place the recently found Chinese genus ''Dsungaripterus''. Later on, also ''Noripterus'' (then now with the name "Phobetor" which was already occupied, therefore the quotation marks) were assigned to the family. In 2003, Alexander Kellner gave the exact definition as a clade:Kellner, A.W.A., 2003. Pterosaur phylogeny and comments on the evolutionary history of the AN group. In: Buffetaut, E., Mazin, J.M. (Eds.), ''Evolution and Palaeobiology of Pterosaurs. Geological Society, London, Special Publication'' 217, 105–137. the group was composed out of the latest common ancestor of ''Dsungaripterus'', ''Noripterus'' and “Phobetor”, and all its descendants. As synapomorphies he gave the next six characteristics: a relatively small eye-socket, which is pla ...
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