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Santonian
The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. The Santonian is preceded by the Coniacian and is followed by the Campanian.Gradstein ''et al.'' (2004) Stratigraphic definition The Santonian Stage was established by French geologist Henri Coquand in 1857. It is named after the city of Saintes in the region of Saintonge, where the original type locality is located. The base of the Santonian Stage is defined by the appearance of the inoceramid bivalve ''Cladoceramus undulatoplicatus''. The GSSP (official reference profile) for the base of the Santonian Stage is located near Olazagutia, Spain; it was ratified by the Subcommission on Cretaceous Stratigraphy in 2012. The Santonian's top (the base of the Campanian Stage) is informally marked by the extinction of the crinoid '' Marsu ...
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Coniacian
The Coniacian is an age or stage in the geologic timescale. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series and spans the time between 89.8 ± 1 Ma and 86.3 ± 0.7 Ma (million years ago). The Coniacian is preceded by the Turonian and followed by the Santonian. Stratigraphic definitions The Coniacian is named after the city of Cognac in the French region of Saintonge. It was first defined by French geologist Henri Coquand in 1857. The base of the Coniacian Stage is at the first appearance of the inoceramid bivalve species '' Cremnoceramus deformis erectus''. The official reference profile for the base (a GSSP) is located in Salzgitter-Salder, Lower Saxony, Germany. The top of the Coniacian (the base of the Santonian Stage) is defined by the appearance of the inoceramid bivalve ''Cladoceramus undulatoplicatus''. The Coniacian overlaps the regional Emscherian Stage of Germany, which is roughly coeval with the Coniacian and Santonian Stages ...
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Marsupites
''Marsupites'' is an extinct genus of crinoids from the Santonian stage of the Late Cretaceous.''Marsupites''
at Fossilworks.org


Biostratigraphic significance

The (ICS) has assigned the extinction of ''Marsupites testudinarius'' as the defining biological marker for the start of the

Cladoceramus
''Cladoceramus'' is an extinct genus of fossil marine pteriomorphian bivalves that superficially resembled the related winged pearly oysters of the extant genus '' Pteria''. They lived in the Santonian stage of the Late Cretaceous. Description Adult shells of ''Cladoceramus'' are small to very large size (more than 1 m in length). Many species with fine, discrete, juvenile ornamentation on umbo, consisting of closely and evenly to subevenly spaced raised concentric growth lines without rugae. Posterior auricle well-defined, triangular, separated from disc by auricular sulcus; a post-umbonal sulcus occurs in some species, as do very small anterior auricles ("ears").Lower Turonian Euramerican Inoceramidae: A morphologic, taxonomic, and ...
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Geologic Timescale
The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronology (scientific branch of geology that aims to determine the age of rocks). It is used primarily by Earth scientists (including geologists, paleontologists, geophysicists, geochemists, and paleoclimatologists) to describe the timing and relationships of events in geologic history. The time scale has been developed through the study of rock layers and the observation of their relationships and identifying features such as lithologies, paleomagnetic properties, and fossils. The definition of standardized international units of geologic time is the responsibility of the International Commission on Stratigraphy (ICS), a constituent body of the International Union of Geological Sciences (IUGS), whose primary objective is to precisely de ...
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Epoch (geology)
The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronology (scientific branch of geology that aims to determine the age of rocks). It is used primarily by Earth scientists (including geologists, paleontologists, geophysicists, geochemists, and paleoclimatologists) to describe the timing and relationships of events in geologic history. The time scale has been developed through the study of rock layers and the observation of their relationships and identifying features such as lithologies, paleomagnetic properties, and fossils. The definition of standardized international units of geologic time is the responsibility of the International Commission on Stratigraphy (ICS), a constituent body of the International Union of Geological Sciences (IUGS), whose primary objective is to precisely de ...
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Age (geology)
The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronology (scientific branch of geology that aims to determine the age of rocks). It is used primarily by Earth scientists (including geologists, paleontologists, geophysicists, geochemists, and paleoclimatologists) to describe the timing and relationships of events in geologic history. The time scale has been developed through the study of rock layers and the observation of their relationships and identifying features such as lithologies, paleomagnetic properties, and fossils. The definition of standardized international units of geologic time is the responsibility of the International Commission on Stratigraphy (ICS), a constituent body of the International Union of Geological Sciences (IUGS), whose primary objective is to precisely de ...
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Henri Coquand
Henri Coquand (1813, in Aix-en-Provence – 1881, in Marseille) was a French geologist and paleontologist. In 1841 he obtained his doctorate in sciences in Paris, and later served as a professor of geology at the University of Besançon, Poitiers and Marseille.Prosopo
Sociétés savantes
From his geological studies of southwestern France, he introduced the stages: , and

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Saintes, Charente-Maritime
Saintes (; Poitevin-Saintongeais: ''Sénte'') is a commune and historic town in western France, in the Charente-Maritime department of which it is a sub-prefecture, in Nouvelle-Aquitaine. Its inhabitants are called ''Saintaises'' and ''Saintais''. Saintes is the second-largest city in Charente-Maritime, with inhabitants in 2008. The city's immediate surroundings form the second-most populous metropolitan area in the department, with inhabitants. While a majority of the surrounding landscape consists of fertile, productive fields, a significant minority of the region remains forested, its natural state. In Roman times, Saintes was known as '' Mediolanum Santonum''. During much of its history, the name of the city was spelled Xaintes or Xainctes. Primarily built on the left bank of the Charente, Saintes became the first Roman capital of Aquitaine. Later it was designated as the capital of the province of Saintonge under the Ancien Régime. Following the French Revolution, it b ...
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Campanian
The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian spans the time from 83.6 (± 0.2) to 72.1 (± 0.2) million years ago. It is preceded by the Santonian and it is followed by the Maastrichtian. The Campanian was an age when a worldwide sea level rise covered many coastal areas. The morphology of some of these areas has been preserved: it is an unconformity beneath a cover of marine sedimentary rocks. Etymology The Campanian was introduced in scientific literature by Henri Coquand in 1857. It is named after the French village of Champagne in the department of Charente-Maritime. The original type locality was a series of outcrop near the village of Aubeterre-sur-Dronne in the same region. Definition The base of the Campanian Stage is defined as a place in the stratigraphic colu ...
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Bivalvia
Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bivalves have no head and they lack some usual molluscan organs, like the radula and the odontophore. They include the clams, oysters, cockles, mussels, scallops, and numerous other families that live in saltwater, as well as a number of families that live in freshwater. The majority are filter feeders. The gills have evolved into ctenidia, specialised organs for feeding and breathing. Most bivalves bury themselves in sediment, where they are relatively safe from predation. Others lie on the sea floor or attach themselves to rocks or other hard surfaces. Some bivalves, such as the scallops and file shells, can swim. The shipworms bore into wood, clay, or stone and live inside these substances. The shell of a bivalve is composed o ...
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Inoceramid
The Inoceramidae are an extinct family of bivalves ("clams") in the Class Mollusca. Fossils of inoceramids are found in marine sediments of Permian to latest Cretaceous in age. Inoceramids tended to live in upper bathyal and neritic environments. Many species of inoceramid are found all over the world, demonstrating the wide distribution of their preferred ecosystems, and possibly long-lived planktotrophic larvae.Lower Turonian Euramerican Inoceramidae: A morphologic, taxonomic, and biostratigraphic overview
A report from the first Workshop on Early Turonian Inoceramids (Oct. 5-8, 1992) in Hamburg, Germany; organized by Heinz Hilbrecht and Peter J. Harries
Despite their wide distribution, the pace of evolution of inoceramids ...
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France
France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan area extends from the Rhine to the Atlantic Ocean and from the Mediterranean Sea to the English Channel and the North Sea; overseas territories include French Guiana in South America, Saint Pierre and Miquelon in the North Atlantic, the French West Indies, and many islands in Oceania and the Indian Ocean. Due to its several coastal territories, France has the largest exclusive economic zone in the world. France borders Belgium, Luxembourg, Germany, Switzerland, Monaco, Italy, Andorra, and Spain in continental Europe, as well as the Netherlands, Suriname, and Brazil in the Americas via its overseas territories in French Guiana and Saint Martin. Its eighteen integral regions (five of which are overseas) span a combined area of and contain clos ...
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