Scaphoideus Titanus
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Scaphoideus Titanus
''Scaphoideus titanus'', the American grapevine leafhopper, is an insect of the leafhopper family (Cicadellidae) which feeds on various plants of the family Vitaceae. Native to North America, it was introduced by accident to Europe where it has become a pest by acting as a vector (epidemiology), vector of the Vitis, grapevine phytoplasma disease flavescence dorée. Mating requires species-specific vibrational patterns that males emit to the females, which are often victim to reproductive interference, including vibrational mating disruption caused by humans for pest control purposes. Nymphs do not engage in vibrational communication. References

Insect vectors of plant pathogens Insects described in 1932 Hemiptera of North America Scaphoideini Taxa named by Elmer Darwin Ball {{Deltocephalinae-stub ...
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Elmer D
Elmer is a name of Germanic British origin. The given name originated as a surname, a medieval variant of the given name Aylmer, derived from Old English ''æþel'' (noble) and ''mær'' (famous). It was adopted as a given name in the United States, "in honor of the popularity of the brothers Ebenezer and Jonathan Elmer, leading supporters of the American Revolution." The name has declined in popularity since the first decades of the 20th century and fell out of the top 1,000 names used for American boys in 2009. However, it continues in use for newborn boys in the United States, where 154 boys born there in 2021 received the name. The name is common in the United States and Canada. Notable people with the name include: Mononym * Eilmer of Malmesbury (or Elmer), 11th-century English Benedictine monk * In the amateur radio subculture, an '' Elmer'' is a mentor to a newcoming amateur radio operatorThe term first appeared in the March, 1971 issue of ''QST'' magazine's "How's DX" ...
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Leafhopper
A leafhopper is the common name for any species from the family Cicadellidae. These minute insects, colloquially known as hoppers, are plant feeders that suck plant sap from grass, shrubs, or trees. Their hind legs are modified for jumping, and are covered with hairs that facilitate the spreading of a secretion over their bodies that acts as a water repellent and carrier of pheromones. They undergo a partial metamorphosis, and have various host associations, varying from very generalized to very specific. Some species have a cosmopolitan distribution, or occur throughout the temperate and tropical regions. Some are pests or vectors of plant viruses and phytoplasmas. The family is distributed all over the world, and constitutes the second-largest hemipteran family, with at least 20,000 described species. They belong to a lineage traditionally treated as infraorder Cicadomorpha in the suborder Auchenorrhyncha, but as the latter taxon is probably not monophyletic, many modern au ...
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Vitaceae
The Vitaceae are a family of flowering plants, with 14 genera and around 910 known species, including common plants such as grapevines (''Vitis'' spp.) and Virginia creeper (''Parthenocissus quinquefolia''). The family name is derived from the genus ''Vitis''. Most ''Vitis'' species have 38 chromosomes (n=19), but 40 (n=20) in subgenus ''Muscadinia'', while ''Ampelocissus'', ''Parthenocissus'', and '' Ampelopsis'' also have 40 chromosomes (n=20) and ''Cissus'' has 24 chromosomes (n=12). The family is economically important as the berries of ''Vitis'' species, commonly known as grapes, are an important fruit crop and, when fermented, produce wine. Species of the genus ''Tetrastigma'' serve as hosts to parasitic plants in the family Rafflesiaceae. Taxonomy The name sometimes appears as Vitidaceae, but Vitaceae is a conserved name and therefore has priority over both Vitidaceae and another name sometimes found in the older literature, Ampelidaceae. In the APG III system (2009) onw ...
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North America
North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Caribbean Sea, and to the west and south by the Pacific Ocean. Because it is on the North American Plate, North American Tectonic Plate, Greenland is included as a part of North America geographically. North America covers an area of about , about 16.5% of Earth's land area and about 4.8% of its total surface. North America is the third-largest continent by area, following Asia and Africa, and the list of continents and continental subregions by population, fourth by population after Asia, Africa, and Europe. In 2013, its population was estimated at nearly 579 million people in List of sovereign states and dependent territories in North America, 23 independent states, or about 7.5% of the world's population. In Americas (terminology)#Human ge ...
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Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia and it is located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. Comprising the westernmost peninsulas of Eurasia, it shares the continental landmass of Afro-Eurasia with both Africa and Asia. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south and Asia to the east. Europe is commonly considered to be Boundaries between the continents of Earth#Asia and Europe, separated from Asia by the drainage divide, watershed of the Ural Mountains, the Ural (river), Ural River, the Caspian Sea, the Greater Caucasus, the Black Sea and the waterways of the Turkish Straits. "Europe" (pp. 68–69); "Asia" (pp. 90–91): "A commonly accepted division between Asia and E ...
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Vector (epidemiology)
In epidemiology, a disease vector is any living agent that carries and transmits an infectious pathogen to another living organism; agents regarded as vectors are organisms, such as parasites or microbes. The first major discovery of a disease vector came from Ronald Ross in 1897, who discovered the malaria pathogen when he dissected a mosquito. Arthropods Arthropods form a major group of pathogen vectors with mosquitoes, flies, sand flies, lice, fleas, ticks, and mites transmitting a huge number of pathogens. Many such vectors are haematophagous, which feed on blood at some or all stages of their lives. When the insects feed on blood, the pathogen enters the blood stream of the host. This can happen in different ways. The ''Anopheles'' mosquito, a vector for malaria, filariasis, and various arthropod-borne-viruses (arboviruses), inserts its delicate mouthpart under the skin and feeds on its host's blood. The parasites the mosquito carries are usually located in its salivary gla ...
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Vitis
''Vitis'' (grapevine) is a genus of 79 accepted species of vining plants in the flowering plant family Vitaceae. The genus is made up of species predominantly from the Northern Hemisphere. It is economically important as the source of grapes, both for direct consumption of the fruit and for fermentation to produce wine. The study and cultivation of grapevines is called viticulture. Most cultivated ''Vitis'' varieties are wind-pollinated with hermaphroditic flowers containing both male and female reproductive structures, while wild species are dieceous. These flowers are grouped in bunches called inflorescences. In many species, such as ''Vitis vinifera'', each successfully pollinated flower becomes a grape berry with the inflorescence turning into a cluster of grapes. While the flowers of the grapevines are usually very small, the berries are often large and brightly colored with sweet flavors that attract birds and other animals to disperse the seeds contained within the berrie ...
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Phytoplasma
Phytoplasmas are obligate intracellular parasites of plant phloem tissue and of the insect vectors that are involved in their plant-to-plant transmission. Phytoplasmas were discovered in 1967 by Japanese scientists who termed them mycoplasma-like organisms. Since their discovery, phytoplasmas have resisted all attempts at ''in vitro'' culture in any cell-free medium; routine cultivation in an artificial medium thus remains a major challenge. Phytoplasmas are characterized by the lack of a cell wall, a pleiomorphic or filamentous shape, a diameter normally less than 1 μm, and a very small genome. Phytoplasmas are pathogens of agriculturally important plants, including coconut, sugarcane, and sandalwood, in which they cause a wide variety of symptoms ranging from mild yellowing to death. Phytoplasmas are most prevalent in tropical and subtropical regions. They are transmitted from plant to plant by vectors (normally sap-sucking insects such as leafhoppers) in which they both sur ...
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Flavescence Dorée
Flavescence dorée (from French "Flavescence" : yellowing and "dorée" : golden) is one of the most important and damaging phytoplasma diseases of the vine with the potential to threaten vineyards. The bacterial agent has recently been named '' Candidatus Phytoplasma vitis'', and its vector is the leafhopper, '' Scaphoideus titanus''. Infection may kill young vines and greatly reduce the productivity of old vines. It is classified as a phytoplasma disease belonging to the group generically termed grapevine yellows.winepros.com.au. Occurrences are in sporadic epidemics, and varieties vary in their sensitivity to it. There is no cure at the moment and the way to manage its spread is by uprooting infected plants and by using selective insecticides in an area-wide pest management approach in order to reduce the hemipteran vectors. Symptoms The incubation period between infection and obvious symptoms is usually one year or more, depending on the response of the plant to both pathogen ...
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Reproductive Interference
Reproductive interference is the interaction between individuals of different species during mate acquisition that leads to a reduction of Fitness (biology), fitness in one or more of the individuals involved. The interactions occur when individuals make mistakes or are unable to recognise their own species, labelled as ‘incomplete Intra-species recognition, species recognition'. Reproductive interference has been found within a variety of taxa, including insects, mammals, birds, amphibians, Marine life, marine organisms, and plants. There are seven causes of reproductive interference, namely signal jamming, heterospecific rivalry, misdirected courtship, heterospecific mating attempts, erroneous Mate choice, female choice, heterospecific mating, and Hybridisation (biology), hybridisation. All types have fitness costs on the participating individuals, generally from a reduction in reproductive success, a waste of gametes, and the expenditure of energy and nutrients. These costs are ...
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