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Ragadidae
Ragadidae is a family of true flies in the superfamily Empidoidea. It was formerly considered a lower taxon, but was published as a new subfamily within Empididae in 2016.Sinclair (2016) Since then, it has been classified as the sister group to Empididae, and has been elevated to family level based on the genetic differences which separate it from Empididae.Wahlberg & Johanson (2018) Description Ragadidae are similar to Empididae and Atelestidae in the sense that their genitalia have symmetrical and straight terminalia. In their wings, the point of origin of the Rs vein is located at a distance from the humeral crossvein (h) as long as, or longer than, h. There is also a circumambient costa in the wing, which distinguishes Ragadidae from Atelestidae. Furthermore, the prosternum being separated from the proepisternum sets Ragadidae aside from Empididae—this is true for all except one genera (''Hydropeza'' ''spp''.), which can instead be characterized through its recurved labr ...
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Atelestidae
Atelestidae is a family of flies in the superfamily Empidoidea. The four genera were placed in a separate family in 1983; they were formerly either in Platypezidae (which are not even particularly closely related) or considered ''incertae sedis''. While they are doubtless the most basal of the living Empidoidea, the monophyly of the family is not fully proven. The genus '' Nemedina'' seems to represent a most ancient lineage among the entire superfamily, while '' Meghyperus'' is probably not monophyletic in its present delimitation, and it is liable to be split up eventually, with some species being placed elsewhere. In 2010, the genus '' Alavesia'', previously only known from Cretaceous fossils, was found alive in Namibia, subsequent species were also described from Brazil. Atelestidae has been shown to form the sister group to the remaining members of the Empidoidea superfamily. Subfamilies include Atelestinae and Nemedininae.Wahlberg & Johanson (2018) Description Atelestid ...
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Empididae
__NOTOC__ Empididae is a family of flies with over 3,000 described species occurring worldwide in all the biogeographic realms but the majority are found in the Holarctic. They are mainly predatory flies like most of their relatives in the Empidoidea, and exhibit a wide range of forms but are generally small to medium-sized, non-metallic and rather bristly. Common names for members of this family are dagger flies (referring to the sharp piercing mouthparts of some species) and balloon flies. The term "dance flies" is sometimes used for this family too, but the dance flies proper, formerly included herein, are now considered a separate family Hybotidae. Description For terms see Morphology of Diptera. Empididae are small to medium-sized flies, rarely large (1.0 to 15.0mm.). The body is slender, or elongated and rarely thickset. The colour ranges from yellow to black, and they may be pollinose or lustrous, but never have a metallic gloss. The head is often small and rounded with ...
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Empidoidea
The Empidoidea are a large monophyletic superfamily of true flies, the sister taxon to the Muscomorpha (Cyclorrhapha). These two groups are sometimes united in the unranked taxon Eremoneura. There are some 10,000 known species within Empidoidea,Moulton & Wiegmann (2007) which are represented on all continents except Antarctica. They are known to have existed since the Jurassic period.Grimaldi & Engel (2005) Empidoidea has been subject to much debate regarding its phylogeny. Based on morphology alone, three major hypotheses had been proposed until 2007 and seemed to be consensus for some time—however, in 2018, Wahlberg & Johanson published the most current phylogeny supported by extensive genetic data, changing the relationship between quite a few families and tribes.Wahlberg & Johanson (2018) Description The majority of these insects are predatory, often with large compound eyes (sometimes covering almost the entire surface of the head), and tend to be associated with moist, t ...
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Hybotidae
Hybotidae, the typical dance flies, are a family of true flies. They belong to the superfamily Empidoidea and were formerly included in the Empididae as a subfamily. Some, such as '' Tachydromia'', are predators that run around on the bark of trees in complex patterns, hence the common name. ''Tachydromia'' species are only about three millimeters long. Description Hybotidae share some similarities with the family Dolichopodidae, when looking at rotation of genitalia and wing characteristics. Male terminalia are rotated dextrally between 45° and 90°, excluding segment 7. Hybotidae wings always have a simple R4+5 vein, where the costa either ends near or at M1/M1+2, or near or at R4+5/R5. Furthermore, it can be distinguished from Dolichopodidae by the point of vein Rs, which it at a distance from the humeral crossvein (h) equal to or longer than the length of h. Systematics The Hybotidae clearly form a lineage quite distinct from the Empididae. Among the Empidoidea, they re ...
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Dolichopodidae
Dolichopodidae, the long-legged flies, are a large, cosmopolitan family of true flies with more than 7,000 described species in about 230 genera. The genus ''Dolichopus'' is the most speciose, with some 600 species. Dolichopodidae generally are small flies with large, prominent eyes and a metallic cast to their appearance, though there is considerable variation among the species. Most have long legs, though some do not. In many species, the males have unusually large genitalia which are taxonomically useful in identifying species. Most adults are predatory on other small animals, though some may scavenge or act as kleptoparasites of spiders or other predators. An expanded concept of the family (Dolichopodidae ''sensu lato'') includes the subfamilies Parathalassiinae and Microphorinae. The latter of these was formerly placed in the Empididae, and was at one time considered a separate family (Microphoridae). However, some authors propose instead that Dolichopodidae ''s.l.'' shou ...
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Tachydromiinae
Tachydromiinae is a subfamily of hybotid flies widespread in the world. Tribes and Genera * Tribe Symballophthalmini Bradley, Sinclair & Cumming, 2006 ** '' Symballophthalmus'' Becker'', 1889'' * Tribe Drapetini Collin, 1961 ** '' Allodromia'' Smith, 1962 ** '' Atodrapetis'' Plant, 1997 ** '' Austrodrapetis'' Smith, 1964 ** '' Austrodromia'' Collin, 1961 ** '' Chaetodromia'' Chillcott & Teskey, 1983 ** '' Chersodromia'' Haliday in Walker, 1851 ** '' Crossopalpus'' Bigot, 1857 ** '' Drapetis'' Meigen, 1822 ** '' Dusmetina'' Gil Collado, 1930 ** '' Elaphropeza'' Macquart, 1827 ** '' Isodrapetis'' Collin, 1961 ** '' Megagrapha'' Melander, 1928 ** '' Micrempis'' Melander, 1928 ** '' Nanodromia'' Grootaert, 1994 ** '' Ngaheremyia'' Plant & Didham, 2006 ** '' Pontodromia'' Grootaert, 1994 ** '' Sinodrapetis'' Yang, Gaimari & Grootaert, 2004 ** '' Stilpon'' Loew, 1859 *Tribe Tachydromiini ** '' Charadrodromia'' Melander, 1928 ** '' Dysaletria'' Loew, 1864 ** '' Pieltaini ...
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Ocydromiinae
Ocydromiinae is a subfamily of hybotid flies. Tribes * Tribe Bicellariini Bradley, Sinclair & Cumming, 2006 ** Genus '' Bicellaria'' Macquart, 1823 ** Genus '' Hoplocyrtoma'' ** Genus '' Leptocyrtoma'' * Tribe Ocydromiini ** Genus '' Apterodromia'' Oldroyd, 1949 ** Genus '' Leptodromiella'' Tuomikoski, 1936 ** Genus '' Leptodromia'' Sinclair & Cumming, 2000 ** Genus '' Leptopeza'' Macquart, 1834 ** Genus '' Neotrichina'' Sinclair & Cumming, 2000 ** Genus '' Ocydromia'' Meigen, 1820 ** Genus '' Oropezella'' Collin, 1926 * Tribe Oedaleini Chvála, 1983 ** Genus '' Allanthalia'' Melander, 1927 ** Genus '' Anthalia'' Zetterstedt, 1838 ** Genus ''Euthyneura Euthyneura is a taxonomic infraclass of snails and slugs, which includes species exclusively from marine, aquatic and terrestrial gastropod mollusks in the clade Heterobranchia. Euthyneura are characterised by several autapomorphies, but ar ...'' Macquart, 1836 ** Genus '' Oedalea'' Meigen, 1820 References ...
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