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Punatitan
''Punatitan'' (meaning " puna giant") is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Ciénaga del Río Huaco Formation of La Rioja, Argentina. It contains one species, ''Punatitan coughlini''. Etymology The generic name ''Punatitan'' refers to the oxygen-depleted atmosphere characteristic of the Andes. The specific name refers to geologist Tim Coughlin, who first reported dinosaur fossils in the discovery locality. Description Known from the holotype CRILAR-Pv 614 (Paleovertebrate Collection of Centro Regional de Investigaciones Científicas y Transferencia Tecnológica de La Rioja, Argentina), a partial skeleton composed of the anterior portion of posterior cervical vertebra (likely C12), two middle dorsal vertebrae (likely D6–D7), a partial sacrum, 13 articulated caudal vertebrae (some with articulated haemal arches), the right pubis, the left ischium, and several dorsal ribs, ''Punatitan'' was roughly long. Classification The describers' phyl ...
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Bravasaurus
''Bravasaurus'' (meaning Laguna Brava lizard) is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Ciénaga del Río Huaco Formation of La Rioja, Argentina. It contains one species, ''Bravasaurus arreirosorum''. Etymology The generic name ''Bravasaurus'' is derived from the Laguna Brava National Park in Argentina. The specific name refers to the people, the ''arreiros'' or drivers in Spanish, who carried cattle through the Andes in the 19th century. Description ''Bravasaurus'' was roughly long. It is known from the holotype CRILAR-Pv 612, which consists of the right quadrate and quadratojugal, four cervical, five dorsal, and three caudal vertebrae, few dorsal ribs, three haemal arches, the left humerus, a fragmentary ulna, the metacarpal IV, a partial left ilium with sacral ribs, the right pubis, a partial ischium, the left femur, and both fibulae, and the paratype CRILAR-Pv 613, which consists of an isolated tooth, the right ilium, the right femur, an ...
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Arrudatitan
''Arrudatitan'' (meaning " Arruda's giant") is an extinct genus of titanosaur sauropod dinosaur known from the Late Cretaceous (Campanian-Maastrichtian)-aged Adamantina Formation of Brazil. The type species, ''A. maximus'', was named and described in 2011 as a species of ''Aeolosaurus'', but was separated into its own genus in 2021. It was relatively gracile for a titanosaur. Discovery and naming The holotype, MPMA 12-0001-97, which includes two partial posterior cervicals, fragments of several dorsals, parts of nine caudals, seven partial cervical ribs, twelve partial dorsal ribs, eight chevrons, a fragmentary scapula and arm bones, the left and partial right femur, the left ischium, and fragments; with other specimens known, such as the isolated middle caudal vertebra MPP 248, was discovered in 1997 by Ademir Frare and his 12-year-old nephew Luiz Augusto dos Santos Frare in a field in Cândido Rodrigues. They notified palaeontologist Antônio Celso de Arruda Campos and the holo ...
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Aeolosaurini
Aeolosaurini is an extinct clade of titanosaurian dinosaurs known from the Cretaceous period of Argentina and Brazil. Rodrigo M. Santucci and Antonio C. de Arruda-Campos (2011) in their cladistic analysis found ''Aeolosaurus'', ''Gondwanatitan'', '' Maxakalisaurus'', '' Panamericansaurus'' and '' Rinconsaurus'' to be aeolosaurids. Aeolosaurini is characterized by several synapomorphies of the caudal vertebrae, such as angled centra, elongate prezygapophyses, and neural arches shifted anteriorly relative to the centra. In life, their tails may have been strongly curved downward as a result of these traits, which may have increased the force exerted by the caudofemoralis longus muscle in retracting the hindlimb. Some aeolosaurins, such as ''Shingopana'' and ''Overosaurus'', were relatively small compared to other titanosaurs, whereas others, such as ''Aeolosaurus maximus'', were large. Phylogeny Aeolosaurini was defined by Franco-Rosas, Salgado, Rosas and Carvalho (2004) as the ste ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Gondwanatitan
''Gondwanatitan'' (meaning "giant from Gondwana") was a titanosaurian sauropod dinosaur. ''Gondwanatitan'' was found in Brazil, at the time part of the southern supercontinent Gondwana, in the late Cretaceous Period (70 mya). Like some other sauropods, ''Gondwanatitan'' was tall and ate tough shoots and leaves off of the tops of trees. ''G. faustois closest relative was ''Aeolosaurus''. The type species is ''Gondwanatitan faustoi'', formally described by Kellner and de Azevedo in 1999. Etymology ''Gondwanatitan'' means "Gondwana Titan", and is named after Gondwana, the supercontinent that the genus' South American range was once part of, and the Titans of classical Greek mythology. The type and only named species, ''G. faustoi'', is a patronym honoring Dr. Fausto L. de Souza Cunha, a former curator at the Museu Nacional/UFRJ who led the excavation of the type specimen. Description ''Gondwanatitan'' was a fairly small sauropod, only 7 meters (23 ft) long and weighing about 1 ...
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Trigonosaurus
''Trigonosaurus'' (meaning "triangle lizard" after Triangulo Mineiro, where it was found), is a genus of saltasaurid dinosaurs from the Maastrichtian Serra da Galga Formation of Brazil. The type species, ''Trigonosaurus pricei'', was first described by Campos, Kellner, Bertini, and Santucci in 2005. It was based on two specimens, both consisting mainly of vertebrae. The two specimens were believed to have come from the same individual. However, one specimen was described as the holotype of ''Caieiria'' in 2022. Before its description, it was known as the "Peirópolis titanosaur", after the place it was found. History From the 1940s to 1960s, Brazilian paleontologist Llewellyn Ivor Price excavated several series of titanosaur fossils in the "Caieira" locality of the Serra da Galga Formation in Minas Gerais, Brazil. Two of these, "Series B" or MCT 1488-R and "Series C" or MCT 1490-R, were named the holotypes of two titanosaur genera in 2005: ''Trigonosaurus'' and ''Baurutitan ...
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Overosaurus
''Overosaurus'' (meaning "Overo lizard", after the Cerro Overo locality) is an extinct genus of sauropod dinosaurs, containing only a single species, ''Overosaurus paradasorum''. This species lived approximately 86 to 84 million years ago during the latter part of the Cretaceous Period in what is now Patagonia (in southern Argentina). ''Overosaurus paradasorum'' was relatively small compared to other sauropods from Patagonia, like the saltasaurids and other aeolosaurines, estimated as approximately . It was a ground-dwelling herbivore. Discovery and naming The only known specimen of ''Overosaurus paradasorum'' was recovered at the Cerro Overo locality in Patagonia, Argentina. The specimen was collected in 2002 by researchers from the Museo Carmen Funes, the Museo Argentino Urquiza, and the Natural History Museum of Los Angeles County, in terrestrial sediments deposited during the Campanian stage of the Late Cretaceous period, approximately 84 to 78 million years ago. This speci ...
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Muyelensaurus
''Muyelensaurus'' (meaning "Muyelen lizard", after an indigenous name for the Colorado River in Argentina) is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous of Argentina. It was more slender than other titanosaurs. Fossils have been recovered from the Plottier Formation in the Neuquén province of Patagonia. The type species is ''M. pecheni''.''Muyelensaurus''
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The name ''Muyelensaurus'' first appeared in a 2007 paper by

Rinconsaurus
''Rinconsaurus'' is a genus of titanosaur sauropod dinosaur from the Late Cretaceous in what is now Argentina. The type species, ''Rinconsaurus caudamirus'', was described by Calvo and Riga in 2003, and is based on three partial skeletons. Description Like all sauropods, ''Rinconsaurus'' was a large long-necked quadrupedal animal, with a long, whip-like tail and four pillar-like legs. ''Rinconsaurus'' was an unusually slender sauropod. Although fossil discoveries are incomplete, and no complete necks or heads have been found, fully grown ''Rinconsaurus'' are estimated to have been 11 meters (36 ft) long and approximately 2.5 meters (8 ft) high at the shoulder. The body mass of ''Rinconsaurus'' has been estimated to be between 3.21 and 5.39 tonnes. Discovery and species Fossils of ''Rinconsaurus'' were discovered in 1997 by Gabriel Benítezin strata belonging to the Bajo de la Carpa Formation near Rincón de los Sauces, in the Neuquen province of Argentina. ...
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Lognkosauria
Lognkosauria is a clade of giant long-necked sauropod dinosaurs within the clade Titanosauria. It includes some of the largest and heaviest dinosaurs known. Description Lognkosaurians can be distinguished from other titanosaurs by the wide and unusually thick cervical rib loops on their neck vertebrae, their extremely robust neck neural spines, the relatively narrow neural canal, and their huge vaulted neural arches. They also had very wide dorsal vertebrae with wing-like side processes, and extremely wide rib cages. Their dorsal side processes are also fairly in-line with the level of the neural canal, instead of being attached further up the neural arch as in lithostrotians. Skull material from ''Malawisaurus'', the sister taxon to Lognkosauria, indicates that lognkosaurians at least began with the big-nosed, rounded head shape of earlier titanosaurs and more basal macronarians. Classification Lognkosauria was defined as the clade encompassing the most recent common ancestor o ...
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Notocolossus
''Notocolossus'' is a genus of titanosaurian sauropod dinosaur from late Cretaceous strata of Mendoza Province, Argentina. Discovery and naming A fossil of a large sauropod was discovered by the Argentine paleontologist Dr. Bernardo Javier González Riga in Mendoza province In 2016, the type species ''Notocolossus gonzalezparejasi'' was named and described by Bernardo Javier González Riga, Matthew Carl Lamanna, Leonardo Daniel Ortiz David, Jorge Orlando Calvo and Juan P. Coria. The generic name combines the Greek words νότος, ''notos'', "south wind", and κολοσσός, ''kolossos'', "giant statue", in reference to the provenance from the Southern Hemisphere and the gigantic size of the animal. The specific name honours Jorge González Parejas, for having studied the dinosaur fossils of Mendoza province for two decades. The holotype, UNCUYO-LD 301, was found in a layer of the Plottier Formation dating from the Coniacian-Santonian, about eighty-six million y ...
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Bonitasaura
''Bonitasaura'' is a titanosaurian dinosaur hailing from uppermost layers of the Late Cretaceous (Santonian) Bajo de la Carpa Formation, Neuquén Group of the eastern Neuquén Basin, located in Río Negro Province, Northwestern Patagonia, Argentina. The remains, consisting of a partial sub-adult skeleton jumbled in a small area of fluvial sandstone, including lower jaw with teeth, partial vertebrae series and limb bones, were described by Sebastian Apesteguía in 2004. The genus name ''Bonitasaura'' refers to the fossil quarry's name, "La Bonita", while the name of the type species, ''B. salgadoi'', pays homage to Leonardo Salgado, a renowned Argentine paleontologist.Gallina, P. A. (2011)Notes on the axial skeleton of the titanosaur Bonitasaura salgadoi (Dinosauria-Sauropoda).''Anais da Academia Brasileira de Ciências'', ''83''(1), 235-246. Description ''Bonitasaura'' measured in length, and had a skull similar to another group of sauropods, the diplodocids. The lower jaw ha ...
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