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Procolophoninae
Procolophoninae is an extinct subfamily of procolophonid parareptiles from the late Early Triassic to the early Middle Triassic (Olenekian and Anisian stages) of Africa, Antarctica, Asia, Europe and South America. Currently, the oldest-known procolophonine is ''Procolophon'' from the earliest Olenekian stage. Phylogeny Procolophoninae was named in 1890 by Richard Lydekker. It is a stem-based taxon defined phylogenetically for the first time by Modesto ''et al.'' (2002) as "all taxa more related to ''Procolophon trigoniceps'' Owen, 1876 than to ''Leptopleuron lacertinum'' Owen, 1851". The cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... below follows Ruta ''et al.'' 2011. References Procolophonines Early Triassic reptiles Middle Triassic reptiles Late Tria ...
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Procolophonidae
Procolophonidae is an extinct family of small, lizard-like parareptiles known from the Late Permian to Late Triassic that were distributed across Pangaea, having been reported from Europe, North America, China, South Africa, South America, Antarctica and Australia. The most primitive procolophonids were likely insectiovous or omnivorous, more derived members of the clade developed bicusped molars, and were likely herbivorous feeding on high fiber vegetation or durophagous omnivores. Many members of the group are noted for spines projecting from the quadratojugal bone of the skull, which likely served a defensive purpose as well as possibly also for display. At least some taxa were likely fossorial burrowers. While diverse during the Early and Middle Triassic, they had very low diversity during the Late Triassic, and were extinct by the beginning of the Jurassic. Phylogeny Below is a cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a dia ...
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Kapes (genus)
''Kapes'' is an extinct genus of procolophonid parareptile from the Lower and Middle Triassic of the United Kingdom and Russia. It is a member of the subfamily Procolophoninae. The type species ''K. amaenus'' was named in 1975 from the banks of the Vychegda River in the Komi Republic of Russia. In 1983, a new species was brought into the genus, ''K. majmesculae''. ''K. majmesculae'' was first named in 1968 as a member of the genus ''Tichvinskia''. A third Russian species, ''K. serotinus'', was named in 1991. In 2002, ''Kapes bentoni'' was named from the Middle Triassic Otter Sandstone Formation of Devon, England, extending the geographic range of ''Kapes''. In the same paper, ''K. serotinus'' was synonymized with ''K. majmesculae'' and another Russian species was assigned to ''Kapes'' called ''K. komiensis''. ''K. komiensis'' was first named in 1975 (in the same paper ''K. amaenus'' was named in) as a member of the genus '' Macrophon''. Phylogeny Below is a cladogram A clado ...
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Eumetabolodon
''Eumetabolodon'' is an extinct genus of procolophonine procolophonid parareptile from early and middle Triassic (Induan to earliest Anisian stages) deposits of Nei Mongol, northern China. Two species of ''Eumetabolodon'' were named by J. L. Li in 1983 and the type species is ''Eumetabolodon bathycephalus''. Discovery The type species is known from the holotype IVPP V6064, a nearly complete skull which was collected in the Zhuengeerqi locality, from the upper Heshanggou Formation (Olenekian and earliest Anisian stages), Ordos Basin. A second specimen, IVPP V6070 was collected in the Zhuengeerqi locality, from the Ermaying Formation (upper Olenekian stage), Ordos Basin. Additional 17 specimens, IVPP V6065-69, V6166 (1&2), V6167, V6168 (1&2), V6169-75, were collected in the Gucheng locality, from the upper Heshanggou Formation, Ordos Basin. ''"E." dongshengensis'' is known only from the holotype IVPP V6073, a partial skull including right frontal, prefrontal, ...
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Early Triassic
The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a unit in chronostratigraphy. The Early Triassic is the oldest epoch of the Mesozoic Era. It is preceded by the Lopingian Epoch (late Permian, Paleozoic Era) and followed by the Middle Triassic Epoch. The Early Triassic is divided into the Induan and Olenekian ages. The Induan is subdivided into the Griesbachian and Dienerian subages and the Olenekian is subdivided into the Smithian and Spathian subages. The Lower Triassic series is coeval with the Scythian Stage, which is today not included in the official timescales but can be found in older literature. In Europe, most of the Lower Triassic is composed of Buntsandstein, a lithostratigraphic unit of continental red beds. The Early Triassic and partly also the Middle Triassic span the in ...
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Thelerpeton
''Thelerpeton'' is an extinct genus of procolophonine procolophonid parareptile from middle Triassic (early Anisian stage) deposits of Free State Province, South Africa. It is known from the holotype BP/1/4538, a nearly complete skull. It was collected by the South African palaeontologist, James W. Kitching from Hugoskop in the Rouxville District and referred to subzone B of the Cynognathus Assemblage Zone of the Burgersdorp Formation, Beaufort Group (Karoo Basin). It was first named by Sean P. Modesto and Ross J. Damiani in 2003 and the type species is ''Thelerpeton oppressus''. It was first assigned to a species of '' Thelegnathus'' (now considered to be a ''nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...''), ''Thelegnathus oppressus''. References Procolo ...
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Richard Lydekker
Richard Lydekker (; 25 July 1849 – 16 April 1915) was an English naturalist, geologist and writer of numerous books on natural history. Biography Richard Lydekker was born at Tavistock Square in London. His father was Gerard Wolfe Lydekker, a barrister-at-law with Dutch ancestry. The family moved to Harpenden Lodge soon after Richard's birth. He was educated at Trinity College, Cambridge, where he took a first-class in the Natural Science tripos (1872). In 1874 he joined the Geological Survey of India and made studies of the vertebrate palaeontology of northern India (especially Kashmir). He remained in this post until the death of his father in 1881. His main work in India was on the Siwalik palaeofauna; it was published in ''Palaeontologia Indica''. He was responsible for the cataloguing of the fossil mammals, reptiles, and birds in the Natural History Museum (10 vols., 1891). He named a variety of taxa including the golden-bellied mangabey; as a taxon authority he is nam ...
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South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southern subregion of a single continent called America. South America is bordered on the west by the Pacific Ocean and on the north and east by the Atlantic Ocean; North America and the Caribbean Sea lie to the northwest. The continent generally includes twelve sovereign states: Argentina, Bolivia, Brazil, Chile, Colombia, Ecuador, Guyana, Paraguay, Peru, Suriname, Uruguay, and Venezuela; two dependent territories: the Falkland Islands and South Georgia and the South Sandwich Islands; and one internal territory: French Guiana. In addition, the ABC islands of the Kingdom of the Netherlands, Ascension Island (dependency of Saint Helena, Ascension and Tristan da Cunha, a British Overseas Territory), Bouvet Island ( dependency of Norway), Pa ...
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Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia and it is located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. Comprising the westernmost peninsulas of Eurasia, it shares the continental landmass of Afro-Eurasia with both Africa and Asia. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south and Asia to the east. Europe is commonly considered to be Boundaries between the continents of Earth#Asia and Europe, separated from Asia by the drainage divide, watershed of the Ural Mountains, the Ural (river), Ural River, the Caspian Sea, the Greater Caucasus, the Black Sea and the waterways of the Turkish Straits. "Europe" (pp. 68–69); "Asia" (pp. 90–91): "A commonly accepted division between Asia and E ...
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Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area of , about 30% of Earth's total land area and 8.7% of Earth's total surface area. The continent, which has long been home to the majority of the human population, was the site of many of the first civilizations. Its 4.7 billion people constitute roughly 60% of the world's population. In general terms, Asia is bounded on the east by the Pacific Ocean, on the south by the Indian Ocean, and on the north by the Arctic Ocean. The border of Asia with Europe is a historical and cultural construct, as there is no clear physical and geographical separation between them. It is somewhat arbitrary and has moved since its first conception in classical antiquity. The division of Eurasia into two continents reflects East–West cultural, linguistic, ...
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Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest continent, being about 40% larger than Europe, and has an area of . Most of Antarctica is covered by the Antarctic ice sheet, with an average thickness of . Antarctica is, on average, the coldest, driest, and windiest of the continents, and it has the highest average elevation. It is mainly a polar desert, with annual precipitation of over along the coast and far less inland. About 70% of the world's freshwater reserves are frozen in Antarctica, which, if melted, would raise global sea levels by almost . Antarctica holds the record for the lowest measured temperature on Earth, . The coastal regions can reach temperatures over in summer. Native species of animals include mites, nematodes, penguins, seals and tardigrades. Where vegetation o ...
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Africa
Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area and 20% of its land area.Sayre, April Pulley (1999), ''Africa'', Twenty-First Century Books. . With billion people as of , it accounts for about of the world's human population. Africa's population is the youngest amongst all the continents; the median age in 2012 was 19.7, when the worldwide median age was 30.4. Despite a wide range of natural resources, Africa is the least wealthy continent per capita and second-least wealthy by total wealth, behind Oceania. Scholars have attributed this to different factors including geography, climate, tribalism, colonialism, the Cold War, neocolonialism, lack of democracy, and corruption. Despite this low concentration of wealth, recent economic expansion and the large and young population make Afr ...
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Anisian
In the geologic timescale, the Anisian is the lower stage or earliest age of the Middle Triassic series or epoch and lasted from million years ago until million years ago. The Anisian Age succeeds the Olenekian Age (part of the Lower Triassic Epoch) and precedes the Ladinian Age. Stratigraphic definitions The stage and its name were established by Austrian geologists Wilhelm Heinrich Waagen and Carl Diener in 1895. The name comes from ''Anisus'', the Latin name of the river Enns. The original type locality is at Großreifling in the Austrian state of Styria. The base of the Anisian Stage (also the base of the Middle Triassic series) is sometimes laid at the first appearance of conodont species '' Chiosella timorensis'' in the stratigraphic record. Other stratigraphers prefer to use the base of magnetic chronozone MT1n. There is no accepted global reference profile for the base, but one ( GSSP or golden spike) was proposed at a flank of the mountain Deşli Caira in the Roman ...
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