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Platypterygiinae
Platypterygiinae is an extinct subfamily of ophthalmosaurid thunnosaur ichthyosaurs from the early Late Jurassic to the early Late Cretaceous (Kimmeridgian - Cenomanian) of Asia, Australia, Europe, North America and South America. Currently, the oldest known platypterygiine is ''Brachypterygius''. Platypterygiines were characterized by square tooth roots in cross-section, an extremely reduced extracondylar area of the basioccipital, prominent dorsal and ventral trochanters on humerus and ischiopubis lacking an obturator foramen. Phylogeny Platypterygiinae was named in 2001 by Maxim S. Arkhangelsky and dually noted by colleague Aleski Masluk. It is a stem-based taxon defined phylogenetically for the first time by Fischer ''et al.'' (2012) as "all taxa closer to '' Platypterygius hercynicus'' than to ''Ophthalmosaurus icenicus''". The cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among org ...
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Ophthalmosaurid
Ophthalmosauridae is an extinct family of thunnosaur ichthyosaurs from the Middle Jurassic to the early Late Cretaceous (Bajocian - Cenomanian) worldwide. Almost all ichthyosaurs from the Middle Jurassic onwards belong to the family, until the extinction of ichthyosaurs in the early Late Cretaceous. Opthalmosaurids appeared worldwide during early Bajocian, subsequent to the disappearance of most other ichthyosaur lineages after the end of the Toarcian. Currently, the oldest known ophthalmosaurids is ''Mollesaurus'' from the early Bajocian of Argentina, as well as indeterminate remains of the same age from Luxembourg and Canada. Named by George H. Baur, in 1887, the family contains the basal taxa like ''Ophthalmosaurus''. Appleby (1956) named the taxon Ophthalmosauria which was followed by some authors, but these two names are often treated as synonyms; Ophthalmosauridae has the priority over Ophthalmosauria. However, some researchers argue that Ophthalmosauridae should be restricte ...
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Sisteronia
''Sisteronia'' is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur known from the 'middle' Cretaceous of southeastern England and southeastern France. It contains a single species, ''Sisteronia seeleyi''. Discovery ''Sisteronia'' was named by Valentin Fischer, Nathalie Bardet, Myette Guiomar and Pascal Godefroit in 2014 and the type species is ''Sisteronia seeleyi''. The generic name honors Sisteron, a commune in the Alpes-de-Haute-Provence, southeastern France, where relatively complete specimens referable to ''Sisteronia'' were collected, including a partial articulated skeleton and at least three additional articulated specimens held in a private collection. The specific name, ''seeleyi'', honors the renowned British paleontologist Harry Govier Seeley who cataloged thousands fragmentary ichthyosaur specimens from the Cambridge Greensand Member of the Lower Chalk Formation. Now housed in the collections of Sedgwick Museum of Earth Sciences (CAMSM), the Royal ...
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Acuetzpalin
''Acuetzpalin'' (meaning "water lizard" in Classical Nahuatl) is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur found in the Kimmeridgian La Casita Formation in Mexico. It is known from a partial skeleton (missing the limbs), of which the skull is surprisingly well preserved. Its length is estimated to have been more than long, considering the missing parts of the holotype. It was the first ichthyosaur described in 2020 and the first new ichthyosaur genus described since 2017 (both ''Gengasaurus'' and ''Keilhauia'' were described in 2017). Classification A phylogenetic analysis conducted in 2020 reveals that ''Acuetzpalin'' is a member of the subfamily Platypterygiinae Platypterygiinae is an extinct subfamily of ophthalmosaurid thunnosaur ichthyosaurs from the early Late Jurassic to the early Late Cretaceous (Kimmeridgian - Cenomanian) of Asia, Australia, Europe, North America and South America. Currently, the ... and is located among the most-derived platyp ...
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Caypullisaurus
''Caypullisaurus'' is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur from the Late Jurassic to the Early Cretaceous ( Tithonian and Berriasian stages) of Argentina. Its holotype was collected from the Vaca Muerta Formation of Cerro Lotena, Neuquen, dating to the early Tithonian stage of the Late Jurassic, about 150 million years ago. ''Caypullisaurus'' was first named by Marta Fernández in 1997 and the type species is ''Caypullisaurus bonapartei''. It was a large ichthyosaur, measuring about long. Classification It is a member of the family Ophthalmosauridae, and closely related to ''Platypterygius'' and '' Brachypterygius''.Fernández M. 2007. Redescription and phylogenetic position of ''Caypullisaurus'' (Ichthyosauria: Ophthalmosauridae). ''Journal of Paleontology'' 81 (2): 368-375. In 2012, ''Caypullisaurus'' was found to be most closely related to '' Athabascasaurus'' and "''Platypterygius''" ''australis'', and to nest within the subfamily Platypte ...
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Grendelius
''Grendelius'' is a genus of platypterygiine ophthalmosaurid ichthyosaur from the Late Jurassic (Kimmeridgian-Tithonian) of the UK and European Russia. It was a medium-sized ichthyosaur measuring about long. Taxonomy The type species, ''Grendelius mordax'', was described in 1976 on the basis of CAMSM J.68516, a complete skull with associated postcranial elements from the Kimmeridge Clay of England. Later, it was reassigned to ''Brachypterygius'' because differences between the two species were considered insufficient to warrant separate genera, or even species.McGowan, C. 1997. The taxonomic status of Grendelius mordax: a preliminary report. Journal of Vertebrate Paleontology, 17, 428–430. doi:10.1080/02724634.1997.10010986McGowan, C. & Motani, R. Ichthyopterygia. In Sues, H.-D. (ed.) Handbook of Paleoherpetology, vol. 8. Verlag Dr. Friedrich Pfeil, Munich, 175 pp., 19 pls. Efimov, 1998 named a new genus of ichthyosaur, ''Otschevia pseudoscythica'' on the basis of a single sp ...
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Kyhytysuka
''Kyhytysuka'' () is an extinct genus of ophthalmosaurian ichthyosaur from Early Cretaceous Colombia. The animal was previously assigned to the genus ''Platypterygius'', but given its own genus in 2021. ''Kyhytysuka'' was a mid-sized ophthalmosaurian with heterodont dentition and several adaptations suggesting that it was a macropredatory vertebrate hunter living in shallow waters. It contains a single species, ''Kyhytysuka sachicarum''. Discovery and naming The first definitive ichthyosaur remains in Colombia were discovered in the 1970s in the Paja Formation of Villa de Leyva. This fossil, a three dimensionally preserved skull discovered by local Jorge Cárdenas, was described as a new species of ''Platypterygius'', ''P. sachicarum'', by María Páramo-Fonseca in 1997. However the taxon went largely ignored in the following years or was unofficially considered to be synonymous with the Argentinian ''Platypterygius hauthali''. Further research on ichthyosaurs since its discovery ...
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Athabascasaurus
''Athabascasaurus'' is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur known from Alberta, Canada. Discovery and etymology ''Athabascasaurus'' is known from the holotype TMP 2000.29.01, articulated nearly complete postcranial skeleton and nearly complete skull preserved in dorsal view, missing the premaxilla. It was collected in 2000 on the western side of the Syncrude Canada Ltd.'s Base Mine, an open-pit mine near Fort McMurray. The specimen was found in the Wabiskaw Member of the Clearwater Formation, which dates to the earliest Albian stage of the Early Cretaceous epoch, about 112 million years ago. The specimen resides at the Royal Tyrrell Museum of Palaeontology. ''Athabascasaurus'' was first named by Patrick S. Druckenmiller and Erin E. Maxwell in 2010 and the type species is ''Athabascasaurus bitumineus''. The generic name is derived from the name of the Athabasca River, which runs through Athabasca oil sands area where the holotype was co ...
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Brachypterygius
''Brachypterygius'' (meaning ″short wing/paddle″ in Greek) is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur known from the Late Jurassic of England.McGowan, C. & Motani, R. ''Ichthyopterygia''. In Sues, H.-D. (ed.) Handbook of Paleoherpetology, vol. 8. Verlag Dr. Friedrich Pfeil, Munich, 175 pp., 19 pls. The type species was originally described and named as ''Ichthyosaurus extremus'' by Boulenger in 1904.Boulenger, G. A. 1904. Exhibition of, and remarks upon, a paddle of a new species of ichthyosaur. ''Proceedings of the Zoological Society of London'', 1904, 424–426. ''Brachypterygius'' was named by Huene in 1922 for the width and shortness of the forepaddle, and the type species is therefore ''Brachypterygius extremus''.Huene, F. F. von 1922. ''Die Ichthyosaurier des Lias und ihre Zusammenhänge''. Verlag von Gebrüder Borntraeger, Berlin, 114 pp., 22 pls. The holotype of ''B. extremus'' was originally thought to be from the Lias Group of Bath, Unite ...
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Aegirosaurus
''Aegirosaurus'' is an extinct genus of platypterygiine ophthalmosaurid ichthyosaurs known from the late Jurassic and early Cretaceous of Europe. It was originally named as a species of ''Ichthyosaurus''. Discovery and species Originally described by Wagner (1853) as a species of the genus ''Ichthyosaurus'' (''I. leptospondylus''), the species ''Aegirosaurus leptospondylus'' has had an unstable taxonomic history. It has been referred to the species '' Ichthyosaurus trigonus posthumus'' (later reclassified in the dubious genus '' Macropterygius'') in the past, and sometimes identified with ''Brachypterygius extremus''. In 2000, Bardet and Fernández selected a complete skeleton in a private collection as the neotype for the species ''I. leptospondylus'', as the only other described specimen was destroyed in World War II. A second specimen from the Munich collection was referred to the same taxon. Bardet and Fernández concluded that the neotype should be assigned to a new genus, ''A ...
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Maiaspondylus
''Maiaspondylus'' is an extinct genus of platypterygiine ophthalmosaurid ichthyosaurs known from Northwest Territories of Canada, the Cambridge Greensand of England and the Voronezh Region of Russia. Description ''Maiaspondylus'' is known from the holotype UALVP 45635, a disarticulated but nearly complete skeleton preserved in three dimensions and from the referred materials UALVP 45639, two articulated, partially preserved embryos and eight articulated vertebrae of an adult, UALVP 45640, 14 articulated vertebrae of a juvenile, UALVP 45640, 12 articulated vertebrae, UALVP 45642, a partial snout and left dentaries with teeth and UALVP 45643, a fragmentary snout. All specimens were collected at Hay River from the Loon River Formation, dating to the early Albian stage of the Early Cretaceous, about 110 million years ago. All ''Maiaspondylus'' specimens were originally referred to ''Platypterygius''. However, all recent cladistic analyses foun ...
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Platypterygius
''Platypterygius'' is a historically paraphyletic genus of platypterygiine ichthyosaur from the Cretaceous period. It was historically used as a wastebasket taxon, and most species within ''Platypterygius'' likely are undiagnostic at the genus or species level, or represent distinct genera, even being argued as invalid. While fossils referred to ''Platypterygius'' have been found throughout different continents, the holotype specimen was found in Germany. Description As ''Platypterygius'' contains multiple species not especially close to each other, little can be said in terms of shared characteristics. According to an analysis by Fischer (2012), all anatomical features used to unify ''Platypterygius'' species are either not actually present in each species, or much more widespread among unrelated ophthalmosaurs. Generally, species referred to this genus were large bodied macropredators based on their robust dentition. This is also supported by ''P. australis'' having been foun ...
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Simbirskiasaurus
''Simbirskiasaurus'' is an extinct genus of ichthyosaur from the Early Cretaceous (early Barremian) of Ulyanovsk Oblast, Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eig ....Valentin Fischer, Maxim S. Arkhangelsky, Darren Naish, Ilya M. Stenshin, Gleb N. Uspensky and Pascal Godefroit (2014) Simbirskiasaurus and Pervushovisaurus reassessed: implications for the taxonomy and cranial osteology of Cretaceous platypterygiine ichthyosaurs. Zoological Journal of the Linnean Society 171(4): 822–841. http://onlinelibrary.wiley.com/doi/10.1111/zoj.12158/abstract Its type specimen is YKM 65119, a fragmentary skull and vertebral column.Storrs, G. W., M. S. Arkhangel'skii and V. M. Efimov. 2000. Mesozoic marine reptiles of Russia and other former Soviet republics. pages 187-210 In Bento ...
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