Caypullisaurus
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''Caypullisaurus'' is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of platypterygiine ophthalmosaurid ichthyosaur from the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
to the Early Cretaceous ( Tithonian and
Berriasian In the geological timescale, the Berriasian is an age/stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 145.0 ± 4.0 Ma and 139.8 ± 3.0 Ma (million years ago ...
stages) of
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. Its
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...
was collected from the Vaca Muerta Formation of Cerro Lotena, Neuquen, dating to the early Tithonian stage of the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
, about 150
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago) ...
. ''Caypullisaurus'' was first named by Marta Fernández in
1997 File:1997 Events Collage.png, From left, clockwise: The movie set of ''Titanic'', the highest-grossing movie in history at the time; '' Harry Potter and the Philosopher's Stone'', is published; Comet Hale-Bopp passes by Earth and becomes one of ...
and the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
is ''Caypullisaurus bonapartei''. It was a large ichthyosaur, measuring about long.


Classification

It is a member of the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Ophthalmosauridae Ophthalmosauridae is an extinct family of thunnosaur ichthyosaurs from the Middle Jurassic to the early Late Cretaceous (Bajocian - Cenomanian) worldwide. Almost all ichthyosaurs from the Middle Jurassic onwards belong to the family, until the ex ...
, and closely related to ''
Platypterygius ''Platypterygius'' is a historically paraphyletic genus of platypterygiine ichthyosaur from the Cretaceous period. It was historically used as a wastebasket taxon, and most species within ''Platypterygius'' likely are undiagnostic at the genus ...
'' and '' Brachypterygius''.Fernández M. 2007. Redescription and phylogenetic position of ''Caypullisaurus'' (Ichthyosauria: Ophthalmosauridae). ''Journal of Paleontology'' 81 (2): 368-375. In 2012, ''Caypullisaurus'' was found to be most closely related to '' Athabascasaurus'' and "''Platypterygius''" ''australis'', and to nest within the subfamily
Platypterygiinae Platypterygiinae is an extinct subfamily of ophthalmosaurid thunnosaur ichthyosaurs from the early Late Jurassic to the early Late Cretaceous (Kimmeridgian - Cenomanian) of Asia, Australia, Europe, North America and South America. Currently, the ...
.


Phylogeny

The following cladogram shows a possible phylogenetic position of ''Caypullisaurus'' in Ophthalmosauridae according to the analysis performed by Zverkov and Jacobs (2020).


See also

* List of ichthyosaurs * Timeline of ichthyosaur research


References

Early Cretaceous ichthyosaurs Late Jurassic ichthyosaurs Berriasian life Tithonian life Late Jurassic reptiles of South America Early Cretaceous reptiles of South America Cretaceous Argentina Jurassic Argentina Fossils of Argentina Neuquén Basin Fossil taxa described in 1997 Ophthalmosauridae Ichthyosauromorph genera {{cretaceous-reptile-stub