Papilio Nubilus
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Papilio Nubilus
''Papilio nubilus'' is a rare species of swallowtail butterfly from the genus ''Papilio'' that can be found in Borneo. Previously listed as a distinct species ( D'Abrera 1982, Munroe 1961), but now regarded (by Collins & Morris 1985, Tsukada & Nishiyama 1982: 307) as an interspecific hybrid between ''Papilio nephelus'' and ''Papilio polytes''. Subspecies *''Papilio nubilus nubilus'' (northern Borneo) *''Papilio nubilus musianus'' Rothschild, 1899 (Upper Palembang district) Taxonomy ''Papilio nubilus'' is a member of the ''helenus'' species-group. Other members of this clade are *''Papilio helenus'' Linnaeus, 1758 *''Papilio iswara'' White, 1842 *''Papilio iswaroides'' Fruhstorfer, 1898 *''Papilio nephelus ''Papilio nephelus'' is a species of swallowtail butterfly belonging to the family Papilionidae. Subspecies include ''P. n. chaon'', the yellow Helen, and ''P. n. sunatus'', the black and white Helen. Description ''Papilio nephelus'' has a wi ...'' Boisduval, 1836 *'' ...
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Otto Staudinger
Otto Staudinger (2 May 1830 – 13 October 1900) was a German entomologist and a natural history dealer considered one of the largest in the world specialising in the collection and sale of insects to museums, scientific institutions, and individuals. Life Staudinger was born in Groß Wüstenfelde, Duchy of Mecklenburg-Schwerin, Mecklenburg-Schwerin, from a Bavarian family on his father's side. His grandfather was born near Ansbach and came to Holstein at the end of the 18th century where Staudinger's father was born in Groß Flottbeck in 1799. His mother, a born Schroeder, was from Mecklenburg, born in Putzar at the Count of Schwerin's estate in 1794. At the time of Otto Staudinger's birth in 1830 his father was the tenant of the Rittergut Groß Wüstenfelde. At the age of six or seven Otto was introduced into entomology by his private tutor Wagner who collected beetles. In the summer of 1843 his father purchased the Rittergut Lübsee near Güstrow where Otto – now under the ...
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Species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. However, only about 14% of these had been described by 2011. All species (except viruses) are given a two-part name, a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes i ...
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Swallowtail Butterfly
Swallowtail butterflies are large, colorful butterflies in the family Papilionidae, and include over 550 species. Though the majority are tropical, members of the family inhabit every continent except Antarctica. The family includes the largest butterflies in the world, the birdwing butterflies of the genus '' Ornithoptera''. Swallowtails have a number of distinctive features; for example, the papilionid caterpillar bears a repugnatorial organ called the osmeterium on its prothorax. The osmeterium normally remains hidden, but when threatened, the larva turns it outward through a transverse dorsal groove by inflating it with fluid. The forked appearance in some of the swallowtails' hindwings, which can be seen when the butterfly is resting with its wings spread, gave rise to the common name ''swallowtail''. As for its formal name, Linnaeus chose ''Papilio'' for the type genus, as ''papilio'' is Latin for "butterfly". For the specific epithets of the genus, Linnaeus applied ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Papilio
''Papilio'' is a genus in the swallowtail butterfly family, Papilionidae, as well as the only representative of the tribe Papilionini. The word ''papilio'' is Latin for butterfly. It includes the common yellow swallowtail (''Papilio machaon''), which is widespread in the Northern Hemisphere and the type species of the genus, as well as a number of other well-known North American species such as the western tiger swallowtail ('' Papilio rutulus''). Familiar species elsewhere in the world include the Mormons ('' Papilio polytes'', '' Papilio polymnestor'', '' Papilio memnon'', and '' Papilio deiphobus'') in Asia, the orchard and Ulysses swallowtails in Australia (''Papilio aegeus'', '' Papilio ulysses'', respectively) and the citrus swallowtail of Africa (''Papilio demodocus''). Older classifications of the swallowtails tended to use many rather small genera. More recent classifications have been more conservative, and as a result a number of former genera are now absorbed within ...
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Borneo
Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and east of Sumatra. The island is politically divided among three countries: Malaysia and Brunei in the north, and Indonesia to the south. Approximately 73% of the island is Indonesian territory. In the north, the East Malaysian states of Sabah and Sarawak make up about 26% of the island. The population in Borneo is 23,053,723 (2020 national censuses). Additionally, the Malaysian federal territory of Labuan is situated on a small island just off the coast of Borneo. The sovereign state of Brunei, located on the north coast, comprises about 1% of Borneo's land area. A little more than half of the island is in the Northern Hemisphere, including Brunei and the Malaysian portion, while the Indonesian portion spans the Northern and Southern hemisph ...
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Papilio Nephelus
''Papilio nephelus'' is a species of swallowtail butterfly belonging to the family Papilionidae. Subspecies include ''P. n. chaon'', the yellow Helen, and ''P. n. sunatus'', the black and white Helen. Description ''Papilio nephelus'' has a wingspan reaching about . The basic colour of the wings is black, with a chain of white spots on the forewing and a large white or yellow area on the hindwing. The underside and the upperside of the wings in this species are very similar. The hindwings have wavy margins, with long tails. The thorax and the abdomen are black. ''Papilio nephelus'' superficially closely resembles '' P. helenus''. The differences are, however, both of structure and of colour. Male forewing upperside: entirely without the thick coating of short hairs on the outer half. Upperside: black with a sprinkling of yellowish-brown scales on the forewing, that form four longitudinal streaks on the cell and internervular streaks on the outer half. Hindwing: the upper discal ...
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Papilio Polytes
''Papilio polytes'', the common Mormon, is a common species of swallowtail butterfly widely distributed across Asia. This butterfly is known for the mimicry displayed by the numerous forms of its females which mimic inedible red-bodied swallowtails, such as the common rose and the crimson rose. Names The common name is an allusion to the polygamy formerly practiced by members of the Mormon sect according to Harish Gaonkar, of the Natural History Museum in London: The scientific name is constructed from the Latin word for butterfly, ''papilio'', and the Greek word for many, ''poly''. Range Pakistan, India, Bangladesh, Nepal, Sri Lanka, Maldives, Myanmar, Thailand, Singapore, southern and western China (including Hainan and Guangdong provinces), Taiwan, Hong Kong, Japan (Ryukyu Islands), Vietnam, Laos, Cambodia, Andamans, Nicobars, eastern and Peninsular Malaysia, Brunei, Indonesia (except Moluccas and Irian Jaya), Philippines, and Northern Marianas (Saipan). Status Very ...
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Species Group
In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each other, further blurring any distinctions. Terms that are sometimes used synonymously but have more precise meanings are cryptic species for two or more species hidden under one species name, sibling species for two (or more) species that are each other's closest relative, and species flock for a group of closely related species that live in the same habitat. As informal taxonomic ranks, species group, species aggregate, macrospecies, and superspecies are also in use. Two or more taxa that were once considered conspecific (of the same species) may later be subdivided into infraspecific taxa (taxa within a species, such as bacterial strains or plant varieties), that is complex but it is not a species complex. A species complex is in most cas ...
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Cladistics
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics ( synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. R ...
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Papilio Helenus
''Papilio helenus'', the red Helen, is a large swallowtail butterfly found in forests of southern India and parts of southeast Asia. Range ''Papilio helenus'' is rarely found in Sri Lanka, southern and north-east India, Nepal, Bhutan, Bangladesh, Myanmar, Thailand, Laos, Kampuchea, Vietnam, southern China (including Hainan, Guangdong province), Taiwan, southern Japan, South Korea, Ryukyu Islands, peninsular and eastern Malaysia, Brunei, Philippines, and Indonesia (Sumatra, Java, Bangka, Kalimantan, the Lesser Sunda Islands except Tanimbar). In India it occurs along the Western Ghats from Kerala to Gujarat, also Palnis and Shevaroys, in the north from Mussoorie eastwards, to north-east India and onto Myanmar. Description Status ''Papilio helenus'' is generally uncommon and slightly threatened ( in certain places) . It is very commonly found Maharashtra, but also rarely in Gujarat and parts of Kerala. Its history can be traced back to Assam and Western Ghats, where it is ...
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Papilio Iswara
''Papilio iswara'', the great Helen, is a species of large swallowtail butterfly found in parts of Southeast Asia. Status ''Papilio iswara'' is not uncommon and not Threatened species, threatened. Subspecies *''Papilio iswara iswara'' (southern Burma to Singapore, Sumatra, Bangka) *''Papilio iswara araspes'' C. & R. Felder, 1859 (northern Borneo, Natuna Island) Taxonomy ''Papilio iswara'' is a member of the ''helenus'' species group. The members of this Cladistics, clade are: *''Papilio helenus'' Linnaeus, 1758 *''Papilio iswara'' White, 1842 *''Papilio iswaroides'' Fruhstorfer, 1898 *''Papilio nephelus'' Boisduval, 1836 *''Papilio nubilus'' Staudinger, 1895 *''Papilio sataspes'' C. & R. Felder, 1865 Type material The holotype is conserved in the Natural History Museum, London. References *Adam White (zoologist), White, A., 1842, Notice of two New Species of ''Papilio'' from Penang, presented to the British Museum by Sir. Wm. Norris. ''Entomological Magazine, Entomologist'' ...
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