Pannoniasaurus Skeletal Diagram
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Pannoniasaurus Skeletal Diagram
''Pannoniasaurus'' is an extinct genus of tethysaurine mosasauroid known from the Late Cretaceous Csehbánya Formation (Santonian stage) of Hungary. It contains a single species, ''Pannoniasaurus inexpectatus'', Material was copied from this source, which is available under a Creative Commons License. dubbed "unexpected" because it was discovered in freshwater sediments, unlike other mosasaurs, which were marine predators. It was a medium-sized mosasaur, reaching up to in length. Discovery and naming The holotype (MTM 2011.43.1) and referred specimens have been collected from the alluvial sediments of the Csehbánya Formation from various exposures at the Iharkút open-pit bauxite mine, Bakony Hills, Western Hungary since the discovery of the locality in 2000. A single vertebra has been collected in 1999 from the Ajka Coal Formation at the waste dump of the coal mines next to the town of Ajka, 20 km from Iharkút. Currently more than one hundred bones of ''Pannoniasaurus' ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Budapest
Budapest (, ; ) is the capital and most populous city of Hungary. It is the ninth-largest city in the European Union by population within city limits and the second-largest city on the Danube river; the city has an estimated population of 1,752,286 over a land area of about . Budapest, which is both a city and county, forms the centre of the Budapest metropolitan area, which has an area of and a population of 3,303,786; it is a primate city, constituting 33% of the population of Hungary. The history of Budapest began when an early Celtic settlement transformed into the Roman town of Aquincum, the capital of Lower Pannonia. The Hungarians arrived in the territory in the late 9th century, but the area was pillaged by the Mongols in 1241–42. Re-established Buda became one of the centres of Renaissance humanist culture by the 15th century. The Battle of Mohács, in 1526, was followed by nearly 150 years of Ottoman rule. After the reconquest of Buda in 1686, the ...
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Monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have tak ...
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Haasiasaurus
''Haasiasaurus'' is an extinct genus of early mosasaur, originally named "''Haasia''" by M. J. Polcyn ''et al.'', in honour of the palaeontologist Georg Haas (paleontologist), Georg Haas. (The original name was a junior homonym of ''Haasia'' Bollman, 1893, a genus of millipedes.) ''Haasiasaurus'' was the largest cenomanian mosasaur at . The genus contains the species ''Haasiasaurus gittelmani'', which was found in the Cenomanian 100 million years ago (Upper Cretaceous) rocks near Ein Yabrud, in the Palestinian territories, Palestinian West Bank, approximately north of Jerusalem. References

Mosasaurids Extinct animals of Asia Mosasaurs of Asia Fossil taxa described in 2003 {{paleo-lizard-stub ...
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Komensaurus
''Komensaurus'' is a genus of basal aigialosaurid mosasauroid from the Late Cretaceous period. It was found at Komen in Slovenia in limestone dating from the Cenomanian. It was earlier referred to as the "Trieste aigialosaur". In 2007, the type species ''Komensaurus carrolli'' was named. Its holotype, specimen MCSNT 11430, was discovered in Slovenia and lived alongside the related ''Carsosaurus''. It was a relatively small reptile, reaching in length and in body mass. See also * List of mosasaurs This list of mosasaurs is a comprehensive listing of all Genus, genera that have ever been included in the family Mosasauridae or the parent clade Mosasauroidea, excluding purely vernacular terms. The list includes all commonly accepted genera, bu ... External links Oceans of Kansas Notes Mosasaurids Mosasaurs of Europe {{paleo-lizard-stub zh:莫那龍 ...
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Carsosaurus
''Carsosaurus'' is a genus of extinct amphibious reptiles, in the mosasaur superfamily, containing only the species ''Carsosaurus marchesetti''. It is known from a single individual that lived during the Upper Cretaceous in what is now Slovenia. The specimen is well-preserved, containing many different bones as well as some skin impressions and sternal cartilage. While more remains are needed to be certain, it is generally thought to belong to Aigialosauridae. In life, it was an amphibious creature that spent most of its time on land, although its later relatives would become fully aquatic. Discovery and naming ''Carsosaurus marchesetti'' was described from a single, mostly complete skeleton at the Civico Museo di Storia Naturale di Trieste, uncovered from the Karst Plateau near Komen (modern-day Slovenia) by Andreas Kornhuber in 1893. He compared it to ''Acteosaurus tommasinii'', as both were uncovered from the same area. There were numerous noticeable differences between the tw ...
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Russellosaurus
''Russellosaurus'' is an extinct genus of tethysaurine mosasauroid from the Late Cretaceous of North America. The genus was described from a skull discovered in an exposure of the Arcadia Park Shale (lower Middle Turonian) at Cedar Hill, Dallas County in south-central Texas, United States. The skull (SMU 73056, Shuler Museum of Paleontology, Southern Methodist University) was found in 1992 by a member of the Dallas Paleontological Society, who then donated to the museum. Other fragmentary specimens of ''Russelosaurus'' have been recovered from the slightly older Kamp Ranch Limestone at two other localities in the Dallas area. Etymology The type species, ''R. coheni'', was named for the amateur fossil collector who discovered SMU 73056, and the genus name honours paleontologist Dale A. Russell for his extensive work on mosasaurs ("Russell's lizard"). This is the second species of mosasaur to have been named for Russell, the first being ''Selmasaurus russelli'' (Wright and Shannon, ...
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Yaguarasaurus
''Yaguarasaurus'' is an extinct genus of mosasauroid from the Late Cretaceous (Turonian) period of Colombia, South America. The remains discovered (an articulated skull, some vertebrae and ribs) were defined as a new genus and species of mosasaurid, ''Yaguarasaurus columbianus'', by the Colombian paleontologist María Páramo, former director of the Museo de Geología José Royo y Gómez of INGEOMINAS in Bogotá. The first fossils remains of this animal suggested a cranial length of and a total length of ; an additional skull that measures long implies a larger size. This reptile is a member of the family of marine lizards Mosasauridae characteristic of Middle and Upper Cretaceous, with global distribution, but in South America known only through isolated remains (Price, 1957, Pierce and Welles, 1959 ; Bonaparte, 1978; Ameghino, 1918). This mosasaur discovered in Yaguará, was at the moment of discovery the most complete material known in South America. Etymology The remain ...
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Tethysaurus
''Tethysaurus'' is an extinct genus of tethysaurine mosasauroid from the Early Turonian (Late Cretaceous) period. The only species is ''Tethysaurus nopcsai''. Discovery The name means "Tethys' lizard of Nopcsa", a reference to the Greek goddess of the sea Tethys (also the name of the Tethys Ocean, an ancient sea between southern Europe and northern Africa) and to the Hungarian paleontologist Baron Ferenc Nopcsa, who made pioneering studies on Adriatic aquatic squamates. It was found in the Akrabou Formation, near the villages of Tadirhourst and Asfla in the region of Goulmima, Errachidia Province, in Morocco, with three referred specimens that included a nearly complete articulated skull, mandible, vertebrae and portions of the appendicular skeleton. The diagnosis after Bardet ''et al''. is "(...) prefrontal strongly vaulted in anterior view; parietal exhibits a triangular table ending posteriorly in two pointed pegs overlying the supraoccipital; jugal with a large and wide ...
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Sister Taxon
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomi ...
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Tethysaurinae
The Tethysaurinae are a subfamily of mosasaurs, a diverse group of Late Cretaceous marine squamates. Members of the subfamily are informally and collectively known as "tethysaurines" and have been recovered from North America and Africa. Only two tethysaurine genera are known, ''Pannoniasaurus'' and ''Tethysaurus''. The genera ''Yaguarasaurus'' and ''Russellosaurus'' were previously considered tethysaurines until they were grouped with '' Romeosaurus'' in the new subfamily Yaguarasaurinae. A possible member of this clade (subfamily) is a mosasaur specimen known from a maxilla fragment, found in 1960 in the Czech Republic (then Czechoslovakia), in Dolní Újezd near Litomyšl. Like the closely related yaguarasaurines, all tethysaurines were plesiopedal (meaning primitive and not as well adapted to marine life as later mosasaurs). They generally retained relatively small sizes compared to later giant mosasaurs. The tethysaurines appeared during the Turonian and went extinct in the ...
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Phylogenetic Tree Pannoniasaurus
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does n ...
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