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Russellosaurus
''Russellosaurus'' is an extinct genus of tethysaurine mosasauroid from the Late Cretaceous of North America. The genus was described from a skull discovered in an exposure of the Arcadia Park Shale (lower Middle Turonian) at Cedar Hill, Dallas County in south-central Texas, United States. The skull (SMU 73056, Shuler Museum of Paleontology, Southern Methodist University) was found in 1992 by a member of the Dallas Paleontological Society, who then donated to the museum. Other fragmentary specimens of ''Russelosaurus'' have been recovered from the slightly older Kamp Ranch Limestone at two other localities in the Dallas area. Etymology The type species, ''R. coheni'', was named for the amateur fossil collector who discovered SMU 73056, and the genus name honours paleontologist Dale A. Russell for his extensive work on mosasaurs ("Russell's lizard"). This is the second species of mosasaur to have been named for Russell, the first being ''Selmasaurus russelli'' (Wright and Shannon, ...
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Tethysaurine
The Tethysaurinae are a subfamily of mosasaurs, a diverse group of Late Cretaceous marine squamates. Members of the subfamily are informally and collectively known as "tethysaurines" and have been recovered from North America and Africa. Only two tethysaurine genera are known, ''Pannoniasaurus'' and ''Tethysaurus''. The genera ''Yaguarasaurus'' and ''Russellosaurus'' were previously considered tethysaurines until they were grouped with '' Romeosaurus'' in the new subfamily Yaguarasaurinae. A possible member of this clade (subfamily) is a mosasaur specimen known from a maxilla fragment, found in 1960 in the Czech Republic (then Czechoslovakia), in Dolní Újezd near Litomyšl. Like the closely related yaguarasaurines, all tethysaurines were plesiopedal (meaning primitive and not as well adapted to marine life as later mosasaurs). They generally retained relatively small sizes compared to later giant mosasaurs. The tethysaurines appeared during the Turonian and went extinct in the S ...
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Mosasauroid
Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Ancient Greek, Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. They belong to the order Squamata, which includes lizards and snakes. Mosasaurs probably evolved from an extinct group of aquatic lizards known as Aigialosauridae, aigialosaurs in the Late Cretaceous, Earliest Late Cretaceous with 42 described genera. During the last 20 million years of the Cretaceous period (Turonian–Maastrichtian ages), with the extinction of the ichthyosaurs and Pliosauridae, pliosaurs, mosasaurs became the dominant marine predators. They themselves became extinct as a result of the Cretaceous–Paleogene extinction event, K-Pg event at the end of the Cretaceous period, about 66 million years ago. Description Mosasaurs breathed air, were powerful swimmers, and were well-adapted to livi ...
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Selmasaurus
''Selmasaurus'' is an extinct genus of marine lizard belonging to the mosasaur family. It is classified as part of the Plioplatecarpinae subfamily alongside genera like '' Angolasaurus'' and ''Platecarpus''. Two species are known, ''S. russelli'' and ''S. johnsoni''; both are exclusively known from Santonian deposits in the United States. ''Selmasaurus'' is unique among the mosasaurs in that its skull is unusually akinetic, meaning that it is incapable of widening to swallow larger prey. Most mosasaurs have skulls which possess "coupled kinesis" (mesokinesis and streptostyly), that is, parts of the jaw can open widely to accommodate large prey. Description ''Selmasaurus'' was a small predatory mosasaur at approximately 3–5 meters in length. It possesses a relatively low number of teeth for a mosasaur, the lowest of any known species at the time of its discovery. Originally classified as a plioplatecarpine mosasaur, it differs from all other plioplatecarpine mosasaurs in seve ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. However, only about 14% of these had been described by 2011. All species (except viruses) are given a two-part name, a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes i ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Cartilaginous
Cartilage is a resilient and smooth type of connective tissue. In tetrapods, it covers and protects the ends of long bones at the joints as articular cartilage, and is a structural component of many body parts including the rib cage, the neck and the bronchial tubes, and the intervertebral discs. In other taxa, such as chondrichthyans, but also in cyclostomes, it may constitute a much greater proportion of the skeleton. It is not as hard and rigid as bone, but it is much stiffer and much less flexible than muscle. The matrix of cartilage is made up of glycosaminoglycans, proteoglycans, collagen fibers and, sometimes, elastin. Because of its rigidity, cartilage often serves the purpose of holding tubes open in the body. Examples include the rings of the trachea, such as the cricoid cartilage and carina. Cartilage is composed of specialized cells called chondrocytes that produce a large amount of collagenous extracellular matrix, abundant ground substance that is rich in proteoglyc ...
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Septum
In biology, a septum (Latin for ''something that encloses''; plural septa) is a wall, dividing a cavity or structure into smaller ones. A cavity or structure divided in this way may be referred to as septate. Examples Human anatomy * Interatrial septum, the wall of tissue that is a sectional part of the left and right atria of the heart * Interventricular septum, the wall separating the left and right ventricles of the heart * Lingual septum, a vertical layer of fibrous tissue that separates the halves of the tongue. *Nasal septum: the cartilage wall separating the nostrils of the nose * Alveolar septum: the thin wall which separates the alveoli from each other in the lungs * Orbital septum, a palpebral ligament in the upper and lower eyelids * Septum pellucidum or septum lucidum, a thin structure separating two fluid pockets in the brain * Uterine septum, a malformation of the uterus * Vaginal septum, a lateral or transverse partition inside the vagina * Intermuscular sep ...
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Foramen
In anatomy and osteology, a foramen (;Entry "foramen"
in
Merriam-Webster Online Dictionary
'. plural foramina, or foramens ) is an open hole that is present in extant or extinct s. Foramina inside the of typically allow ,

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Jugal
The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anatomy The jugal bone is located on either side of the skull in the circumorbital region. It is the origin of several masticatory muscles in the skull. The jugal and lacrimal bones are the only two remaining from the ancestral circumorbital series: the prefrontal, postfrontal, postorbital, jugal, and lacrimal bones. During development, the jugal bone originates from dermal bone. In dinosaurs This bone is considered key in the determination of general traits in cases in which the entire skull has not been found intact (for instance, as with dinosaurs in paleontology). In some dinosaur genera the jugal also forms part of the lower margin of either the antorbital fenestra or the infratemporal fenestra, or both. Most commonly, this bone articu ...
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Pterygoid Process
The pterygoid processes of the sphenoid (from Greek ''pteryx'', ''pterygos'', "wing"), one on either side, descend perpendicularly from the regions where the body and the greater wings of the sphenoid bone unite. Each process consists of a medial pterygoid plate and a lateral pterygoid plate, the latter of which serve as the origins of the medial and lateral pterygoid muscles. The medial pterygoid, along with the masseter allows the jaw to move in a vertical direction as it contracts and relaxes. The lateral pterygoid allows the jaw to move in a horizontal direction during mastication (chewing). Fracture of either plate are used in clinical medicine to distinguish the Le Fort fracture classification for high impact injuries to the sphenoid and maxillary bones. The superior portion of the pterygoid processes are fused anteriorly; a vertical groove, the pterygopalatine fossa, descends on the front of the line of fusion. The plates are separated below by an angular cleft, the pt ...
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