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Palaeognath
Palaeognathae (; ) is a infraclass of birds, called paleognaths, within the class Aves of the clade Archosauria. It is one of the two extant infraclasses of birds, the other being Neognathae, both of which form Neornithes. Palaeognathae contains five extant branches of flightless lineages (plus two extinct clades), termed ratites, and one flying lineage, the Neotropic tinamous.Wetmore, A. (1960) A Classification for Birds of the World. Smithsonian Miscellaneous Collection (Washington D.C.: Smithsonian Institution) 139: 1–37. There are 47 species of tinamous, five of kiwis (''Apteryx''), three of cassowaries (''Casuarius''), one of emus (''Dromaius'') (another became extinct in historic times), two of rheas (''Rhea'') and two of ostrich (''Struthio'').Clements, J. C. ''et al''. (2010) Recent research has indicated that paleognaths are monophyletic but the traditional taxonomic split between flightless and flighted forms is incorrect; tinamous are within the ratite radiation, me ...
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Ratite
A ratite () is any of a diverse group of flightless, large, long-necked, and long-legged birds of the infraclass Palaeognathae. Kiwi, the exception, are much smaller and shorter-legged and are the only nocturnal extant ratites. The systematics of and relationships within the paleognath clade have been in flux. Previously, all the flightless members had been assigned to the order Struthioniformes, which is more recently regarded as containing only the ostrich. The modern bird superorder Palaeognathae consists of ratites and the flighted Neotropic tinamous (compare to Neognathae). Unlike other flightless birds, the ratites have no keel on their sternum — hence the name, from the Latin ''ratis'' (raft, a vessel which has no keel - in contradistinction to extant flighted birds with a keel). Without this to anchor their wing muscles, they could not have flown even if they developed suitable wings. Ratites are a paraphyletic group; tinamous fall within them, and are the sister gr ...
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Tinamou
Tinamous () form an order of birds called Tinamiformes (), comprising a single family called Tinamidae (), divided into two distinct subfamilies, containing 46 species found in Mexico, Central America, and South America. The word "tinamou" comes from the Galibi term for these birds, ''tinamu''. Tinamous have traditionally been regarded as the sister group of the flightless ratites, but recent work places them well within the ratite radiation, implying basal ratites could fly. Tinamous first appear in the fossil record in the Miocene epoch. They are generally sedentary, ground-dwelling and, though not flightless, when possible avoid flight in favour of hiding or running away from danger. They are found in a variety of habitats, ranging from semi-arid alpine grasslands to tropical rainforests. The two subfamilies are broadly divided by habitat, with the Nothurinae referred to as steppe or open country tinamous, and the Tinaminae known as forest tinamous. Although some species are ...
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Lithornithiformes
Lithornithidae is an extinct, possibly paraphyletic (but see below) group of early paleognath birds. They are known from fossils dating to the Upper Paleocene through the Middle Eocene of North America and Europe, with possible Late Cretaceous representatives. All are extinct today; the youngest specimen is the currently unnamed SGPIMH MEV1 specimen from the mid-Eocene Messel Pit site. Lithornithids had long, slender, bills for probing. They closely resembled modern tinamous. They possessed a rhynchokinetic skull with relatively unfused cranial bones, a weakly fused pygostyle and a splenial. The unguals were more curved than in tinamous and probably allowed better perching in trees. The order Lithornithiformes was erected by Dr. Peter Houde in 1988. Initially, only three genera (''Lithornis'', ''Paracathartes'', and ''Pseudocrypturus'') and eight named species were included. ''Promusophaga'' (Harrison & Walker, 1977) originally considered a stem-turaco, is considered synonymou ...
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Rheidae
Rheidae is a family of flightless ratite birds which first appeared in the Paleocene. It is today represented by the sole living genus '' Rhea'', but also contains several extinct genera. Taxonomy Order Rheiformes (Forbes, 1884) Furbringer, 1888 heimorphae Bonaparte, 1849; Rheae Forbes 1884ref name="mikko"> * Family † Opisthodactylidae Ameghino 1895 ** Genus ?†''Diogenornis'' de Alvarenga 1983 (Late Paleocene) – possibly a member of Casuariiformes instead. ** Genus †''Opisthodactylus'' Ameghino 1895 (Miocene) – rheid? * Family Rheidae (Bonaparte 1849) Bonaparte, 1853 ** Genus †''Heterorhea'' Rovereto 1914 (Pliocene) ** Genus †''Hinasuri ''Hinasuri'' is an extinct genus of rhea from the Montehermosan. Its fossils have been found in the Monte Hermoso Formation of Buenos Aires Province, Argentina. The type species In zoological nomenclature, a type species (''species typica' ...'' Tambussi 1995 ** Genus '' Rhea'' Brisson 1760 References {{Taxonbar, ...
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Neognathae
Neognathae (; ) is a infraclass of birds, called neognaths, within the class Aves of the clade Archosauria. Neognathae includes the majority of living birds; the exceptions being the tinamous and the flightless ratites, which belong instead to the sister taxon Palaeognathae. There are nearly 10,000 living species of neognaths. The earliest fossils are known from the very end of the Cretaceous but molecular clocks suggest that neognaths originated sometime in the first half of the Late Cretaceous, about 90 million years ago. Since then, they have undergone adaptive radiation, producing the diversity of form, function, and behavior that exists today. Neognathae includes the order Passeriformes (perching birds), one of the largest orders of land vertebrates, containing some 60% of living birds. Passeriformes is twice as species-rich as Rodentia and about five times as species-rich as Chiroptera (bats), which are the two largest orders of mammals. Neognathae also contains some ve ...
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Apterygiformes
Kiwi ( ) are flightless birds endemic to New Zealand of the order Apterygiformes. The five extant species fall into the family Apterygidae () and genus ''Apteryx'' (). Approximately the size of a domestic chicken, kiwi are by far the smallest living ratites (which also include ostriches, emus, rheas and cassowaries). However, the ratite group is polyphyletic, and cladistically also includes tinamous, which can also be of moderate size. Members of this expanded group are known as paleognaths. DNA sequence comparisons have yielded the conclusion that kiwi are much more closely related to the extinct Malagasy elephant birds than to the moa with which they shared New Zealand. There are five recognised species, four of which are currently listed as vulnerable, and one of which is near-threatened. All species have been negatively affected by historic deforestation, but their remaining habitat is well-protected in large forest reserves and national parks. At present, the greatest ...
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Casuariiformes
The Casuariiformes is an order of large flightless birds that has four surviving members: the three species of cassowary, and the only remaining species of emu. They are divided into either a single family, Casuariidae, or more typically two, with the emu splitting off into its own family, Dromaiidae. All four living members are native to Australia-New Guinea,Clements, J (2007) but some possible extinct taxa occurred in other landmasses. Systematics and evolution The emus form a distinct family, characterized by legs adapted for running. The total number of cassowary species described, based on minor differences in casque shape and color variations, formerly reached nine. Now, however, only three species are recognized, and most authorities only acknowledge few subspecies or none at all. The fossil record of casuariforms is interesting, but not very extensive. Regarding fossil species of ''Dromaius'' and ''Casuarius'', see their genus pages. As with all ratites, there are sev ...
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Apterygidae
Kiwi ( ) are flightless birds endemic to New Zealand of the order Apterygiformes. The five extant species fall into the family Apterygidae () and genus ''Apteryx'' (). Approximately the size of a domestic chicken, kiwi are by far the smallest living ratites (which also include ostriches, emus, rheas and cassowaries). However, the ratite group is polyphyletic, and cladistically also includes tinamous, which can also be of moderate size. Members of this expanded group are known as paleognaths. DNA sequence comparisons have yielded the conclusion that kiwi are much more closely related to the extinct Malagasy elephant birds than to the moa with which they shared New Zealand. There are five recognised species, four of which are currently listed as vulnerable, and one of which is near-threatened. All species have been negatively affected by historic deforestation, but their remaining habitat is well-protected in large forest reserves and national parks. At present, the greatest ...
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Neotropic
The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In biogeography, the Neotropic or Neotropical realm is one of the eight terrestrial realms. This realm includes South America, Central America, the Caribbean islands, and southern North America. In Mexico, the Yucatán Peninsula and southern lowlands, and most of the east and west coastlines, including the southern tip of the Baja California Peninsula are Neotropical. In the United States southern Florida and coastal Central Florida are considered Neotropical. The realm also includes temperate southern South America. In contrast, the Neotropical Floristic Kingdom excludes southernmost South America, which instead is placed in the Antarctic kingdom. The Neotropic is delimited by similarities in fauna or flora. Its fauna and flora are distinct ...
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Extant Taxon
Neontology is a part of biology that, in contrast to paleontology, deals with living (or, more generally, '' recent'') organisms. It is the study of extant taxa (singular: extant taxon): taxa (such as species, genera and families) with members still alive, as opposed to (all) being extinct. For example: * The moose (''Alces alces'') is an extant species, and the dodo (''Raphus cucullatus'') is an extinct species. * In the group of molluscs known as the cephalopods, there were approximately 600 extant species and 7,500 extinct species. A taxon can be classified as extinct if it is broadly agreed or certified that no members of the group are still alive. Conversely, an extinct taxon can be reclassified as extant if there are new discoveries of living species ("Lazarus species"), or if previously-known extant species are reclassified as members of the taxon. Most biologists, zoologists, and botanists are in practice neontologists, and the term neontologist is used large ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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